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<document id="7A2013DF39565A43FECC36D04CFF054C" ID-DOI="10.11646/zootaxa.3895.4.3" ID-GBIF-Dataset="19065cdb-384f-4ee1-996f-c16dc64cebe3" ID-ISSN="1175-5326" ID-Zenodo-Dep="227791" ID-ZooBank="C6ECAF21-4E33-40C7-ADA6-4A013818835B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461001425946" checkinUser="plazi" docAuthor="Komai, Tomoyuki, Maenosono, Tadafumi &amp; Fujita, Yoshihisa" docDate="2014" docId="03B9FD7AFFB51163FF37FE69033F7231" docLanguage="en" docName="zt03895p524.pdf" docOrigin="Zootaxa 3895 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cheramus Spence Bate 1888" docType="treatment" docVersion="8" lastPageNumber="505" masterDocId="FF808502FFB41161FFA0FFC400027124" masterDocTitle="Two new species of ghost shrimp assigned to the genus Cheramus Spence Bate, 1888 (Crustacea: Decapoda: Axiidea: Callianassidae) from the Ryukyu Islands, Japan" masterLastPageNumber="524" masterPageNumber="503" pageNumber="504" updateTime="1698564789849" updateUser="plazi">
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<mods:title id="4EE7014CDBB4798A11D98547CAFCBB7F">Two new species of ghost shrimp assigned to the genus Cheramus Spence Bate, 1888 (Crustacea: Decapoda: Axiidea: Callianassidae) from the Ryukyu Islands, Japan</mods:title>
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<mods:namePart id="5E208CE303660284406BA6E89C544EB8">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="9018F52B5FC0674B6403233647B900A7">Maenosono, Tadafumi</mods:namePart>
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<mods:date id="6554DF4B3B8B50F405C8D2C80A899B33">2014</mods:date>
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<subSubSection id="C30A1FE7FFB51160FF37FE69025670E3" box="[151,596,429,455]" pageId="1" pageNumber="504" type="nomenclature">
<paragraph id="8BAF4C6CFFB51160FF37FE69025670E3" blockId="1.[151,596,429,455]" box="[151,596,429,455]" pageId="1" pageNumber="504">
<heading id="D0E7FB00FFB51160FF37FE69025670E3" bold="true" box="[151,596,429,455]" fontSize="11" level="1" pageId="1" pageNumber="504" reason="1">
<emphasis id="B964907EFFB51160FF37FE69025670E3" bold="true" box="[151,596,429,455]" pageId="1" pageNumber="504">
Genus
<taxonomicName id="4C1037EFFFB51160FF4EFE69025670E3" ID-CoL="8f84ed4e-2899-4c8d-87f2-78cc7a7e5a75" authority="Spence Bate, 1888" authorityName="Spence Bate" authorityYear="1888" box="[238,596,429,455]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FF4EFE69016970E3" bold="true" box="[238,363,429,455]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
<bibRefCitation id="EF81319DFFB51160FED3FE69025670E3" author="Spence" box="[371,596,429,455]" pageId="1" pageNumber="504" refString="Spence Bate, C. (1888) Report on the Crustacea Macrura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. &quot; Challenger &quot; during the years 1873 - 76, Zoology, 24 (i - xc), 1 - 942. [plates. 1 - 157]" type="journal article" year="1888">Spence Bate, 1888</bibRefCitation>
</taxonomicName>
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<subSubSection id="C30A1FE7FFB51160FF37FE300234730C" pageId="1" pageNumber="504" type="reference_group">
<paragraph id="8BAF4C6CFFB51160FF37FE300234730C" blockId="1.[151,1436,500,552]" pageId="1" pageNumber="504">
<treatmentCitationGroup id="AB006B42FFB51160FF37FE300234730C" pageId="1" pageNumber="504">
<taxonomicName id="4C1037EFFFB51160FF37FE3001EF732E" ID-CoL="8f84ed4e-2899-4c8d-87f2-78cc7a7e5a75" authority="Spence Bate, 1888: 30" authorityName="Spence Bate" authorityPageNumber="30" authorityYear="1888" box="[151,493,500,522]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FF37FE3000FD732E" box="[151,255,500,522]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
<treatmentCitation id="0AB16A7DFFB51160FEA6FE3001EF732E" author="Spence" box="[262,493,500,522]" page="30" pageId="1" pageNumber="504" year="1888">
<bibRefCitation id="EF81319DFFB51160FEA6FE3001CB732E" author="Spence" box="[262,457,500,522]" pageId="1" pageNumber="504" refString="Spence Bate, C. (1888) Report on the Crustacea Macrura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. &quot; Challenger &quot; during the years 1873 - 76, Zoology, 24 (i - xc), 1 - 942. [plates. 1 - 157]" type="journal article" year="1888">Spence Bate, 1888</bibRefCitation>
: 30
</treatmentCitation>
</taxonomicName>
.—
<treatmentCitation id="0AB16A7DFFB51160FDAAFE3002E4732E" author="Borradaile" box="[522,742,500,522]" page="545" pageId="1" pageNumber="504" year="1903">
<bibRefCitation id="EF81319DFFB51160FDAAFE3002B7732E" author="Borradaile" box="[522,693,500,522]" pageId="1" pageNumber="504" refString="Borradaile, L. A. (1903) On the classification of the Thalassinidea. Annals and Magazine of Natural History, 12 (7), 534 - 551. http: // dx. doi. org / 10.1080 / 00222930308678891" type="journal article" year="1903">Borradaile 1903</bibRefCitation>
: 545
</treatmentCitation>
(part).—de
<treatmentCitation id="0AB16A7DFFB51160FCC4FE3103F1732E" author="Man" box="[868,1011,500,522]" page="95" pageId="1" pageNumber="504" year="1928">
<bibRefCitation id="EF81319DFFB51160FCC4FE3103D2732E" author="Man" box="[868,976,500,522]" pageId="1" pageNumber="504" refString="Man, J. G. de. (1928) The Thalassinidae and Callianassidae collected by the Siboga-Expedition with some remarks on the Laomediidae. The Decapoda of the Siboga Expedition, Part VII. Siboga Expeditie Monographie, 39 a 6, 1 - 187. [plates. 1 - 20]" type="journal article" year="1928">Man 1928</bibRefCitation>
: 95
</treatmentCitation>
.—
<treatmentCitation id="0AB16A7DFFB51160FBB1FE300543732E" author="Manning" box="[1041,1345,500,522]" page="780" pageId="1" pageNumber="504" year="1991">
<bibRefCitation id="EF81319DFFB51160FBB1FE300512732E" author="Manning" box="[1041,1296,500,522]" pageId="1" pageNumber="504" refString="Manning, R. B. &amp; Felder, D. L. (1991) Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington, 104, 764 - 792." type="journal article" year="1991">Manning &amp; Felder 1991</bibRefCitation>
: 780
</treatmentCitation>
.—
<treatmentCitation id="0AB16A7DFFB51160FAFFFE300129730C" author="Poore" page="101" pageId="1" pageNumber="504" year="1994">
<bibRefCitation id="EF81319DFFB51160FAFFFE3000FF730C" author="Poore" pageId="1" pageNumber="504" refString="Poore, G. C. B. (1994) A phylogeny of the families of Thalassinidea (Crustacea: Decapoda) with keys to families and genera. Memoirs of the Museum of Victoria, 54, 79 - 120." type="journal article" year="1994">Poore 1994</bibRefCitation>
: 101
</treatmentCitation>
.—
<treatmentCitation id="0AB16A7DFFB51160FEE9FDD601F2730C" author="Sakai" box="[329,496,530,552]" page="363" pageId="1" pageNumber="504" year="2011">
<bibRefCitation id="EF81319DFFB51160FEE9FDD601C2730C" author="Sakai" box="[329,448,530,552]" pageId="1" pageNumber="504" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616." type="journal article" year="2011">Sakai 2011</bibRefCitation>
: 363
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30A1FE7FFB51163FF37FD91033F7231" lastPageId="2" lastPageNumber="505" pageId="1" pageNumber="504" type="discussion">
<paragraph id="8BAF4C6CFFB51160FF37FD9102B3771D" blockId="1.[151,1437,597,2026]" pageId="1" pageNumber="504">
<emphasis id="B964907EFFB51160FF37FD910106734A" bold="true" box="[151,260,597,622]" pageId="1" pageNumber="504">Remarks</emphasis>
. The taxonomic composition of
<taxonomicName id="4C1037EFFFB51160FDD0FD9102E1734A" box="[624,739,597,622]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FDD0FD9102E1734A" box="[624,739,597,622]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
</taxonomicName>
is still in a state of flux. The genus was established by
<bibRefCitation id="EF81319DFFB51160FAEBFD92011E73B6" pageId="1" pageNumber="504" refString="Spence Bate, C. (1888) Report on the Crustacea Macrura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. &quot; Challenger &quot; during the years 1873 - 76, Zoology, 24 (i - xc), 1 - 942. [plates. 1 - 157]" type="journal article">Spence Bate (1888)</bibRefCitation>
for
<taxonomicName id="4C1037EFFFB51160FEEBFDBD032B73B6" authority="Spence Bate, 1888" authorityName="Spence Bate" authorityYear="1888" box="[331,809,633,658]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="B964907EFFB51160FEEBFDBD024E73B6" box="[331,588,633,658]" italics="true" pageId="1" pageNumber="504">Cheramus occidentalis</emphasis>
<bibRefCitation id="EF81319DFFB51160FDF4FDBE032B73B6" author="Spence" box="[596,809,634,658]" pageId="1" pageNumber="504" refString="Spence Bate, C. (1888) Report on the Crustacea Macrura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. &quot; Challenger &quot; during the years 1873 - 76, Zoology, 24 (i - xc), 1 - 942. [plates. 1 - 157]" type="journal article" year="1888">Spence Bate, 1888</bibRefCitation>
</taxonomicName>
, described on the basis of a damaged specimen, lacking chelipeds, from the West Indies. In the same Challenger volume,
<bibRefCitation id="EF81319DFFB51160FC04FD5A04887392" box="[932,1162,670,694]" pageId="1" pageNumber="504" refString="Spence Bate, C. (1888) Report on the Crustacea Macrura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. &quot; Challenger &quot; during the years 1873 - 76, Zoology, 24 (i - xc), 1 - 942. [plates. 1 - 157]" type="journal article">Spence Bate (1888)</bibRefCitation>
described another new species,
<taxonomicName id="4C1037EFFFB51160FF58FD05020C73FE" box="[248,526,705,730]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="B964907EFFB51160FF58FD05020C73FE" box="[248,526,705,730]" italics="true" pageId="1" pageNumber="504">Callianassa occidentalis</emphasis>
</taxonomicName>
, based on a single cheliped taken in the same haul with the
<typeStatus id="54ABF2CEFFB51160FB6EFD0604FC73FE" box="[1230,1278,706,730]" pageId="1" pageNumber="504">type</typeStatus>
of
<taxonomicName id="4C1037EFFFB51160FA89FD05011C73DA" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="B964907EFFB51160FA89FD05011C73DA" italics="true" pageId="1" pageNumber="504">Cheramus occidentalis</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF81319DFFB51160FE92FD2101FD73DA" author="Borradaile" box="[306,511,741,766]" pageId="1" pageNumber="504" refString="Borradaile, L. A. (1903) On the classification of the Thalassinidea. Annals and Magazine of Natural History, 12 (7), 534 - 551. http: // dx. doi. org / 10.1080 / 00222930308678891" type="journal article" year="1903">Borradaile (1903)</bibRefCitation>
relegated
<taxonomicName id="4C1037EFFFB51160FD20FD2102F173DA" box="[640,755,741,766]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FD20FD2102F173DA" box="[640,755,741,766]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
</taxonomicName>
to a subgenus of
<taxonomicName id="4C1037EFFFB51160FC75FD2104F973DA" authority="Leach, 1914" authorityName="Leach" authorityYear="1914" box="[981,1275,741,766]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FC75FD21045973DA" box="[981,1115,741,766]" italics="true" pageId="1" pageNumber="504">Callianassa</emphasis>
Leach, 1914
</taxonomicName>
, thus making
<taxonomicName id="4C1037EFFFB51160FF37FCCD019A7206" box="[151,408,777,802]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="B964907EFFB51160FF37FCCD019A7206" box="[151,408,777,802]" italics="true" pageId="1" pageNumber="504">Cheramus occidentalis</emphasis>
</taxonomicName>
a homonym of
<taxonomicName id="4C1037EFFFB51160FDECFCCD04157206" authority="Stimpson, 1856" authorityName="Stimpson" authorityYear="1856" box="[588,1047,777,802]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="B964907EFFB51160FDECFCCD03627206" box="[588,864,777,802]" italics="true" pageId="1" pageNumber="504">Callianassa occidentalis</emphasis>
Stimpson, 1856
</taxonomicName>
[= a junior subjective synonym of
<taxonomicName id="4C1037EFFFB51160FF37FCEA02637262" authority="Dana, 1854" authorityName="Dana" authorityYear="1854" box="[151,609,813,838]" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="californiensis">
<emphasis id="B964907EFFB51160FF37FCEA01B97262" box="[151,443,813,838]" italics="true" pageId="1" pageNumber="504">Neotrypaea californiensis</emphasis>
(Dana, 1854)
</taxonomicName>
]. Hence,
<bibRefCitation id="EF81319DFFB51160FD6CFCE903947262" author="Borradaile" box="[716,918,813,838]" pageId="1" pageNumber="504" refString="Borradaile, L. A. (1903) On the classification of the Thalassinidea. Annals and Magazine of Natural History, 12 (7), 534 - 551. http: // dx. doi. org / 10.1080 / 00222930308678891" type="journal article" year="1903">Borradaile (1903)</bibRefCitation>
proposed
<taxonomicName id="4C1037EFFFB51160FBAFFCE905677262" box="[1039,1381,813,838]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="batei" subGenus="Cheramus">
<emphasis id="B964907EFFB51160FBAFFCE904977262" box="[1039,1173,813,838]" italics="true" pageId="1" pageNumber="504">Callianassa</emphasis>
(
<emphasis id="B964907EFFB51160FB08FCE905197262" box="[1192,1307,813,838]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
)
<emphasis id="B964907EFFB51160FA8DFCE905677262" box="[1325,1381,813,838]" italics="true" pageId="1" pageNumber="504">batei</emphasis>
</taxonomicName>
as a replacement name for
<taxonomicName id="4C1037EFFFB51160FE11FC9502BE724E" box="[433,700,849,874]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="B964907EFFB51160FE11FC9502BE724E" box="[433,700,849,874]" italics="true" pageId="1" pageNumber="504">Cheramus occidentalis</emphasis>
</taxonomicName>
. Later,
<bibRefCitation id="EF81319DFFB51160FC87FC9503C4724E" author="Biffar" box="[807,966,849,874]" pageId="1" pageNumber="504" refString="Biffar, T. A. (1973) The taxonomic status of Callianassa occidentalis Bate, 1888, and C. batei Borradaile, 1903 (Decapoda, Callianassidae). Crustaceana, 24, 224 - 230. http: // dx. doi. org / 10.1163 / 156854073 X 00399" type="journal article" year="1973">Biffar (1973)</bibRefCitation>
concluded that Spence Bate’s (1888)
<taxonomicName id="4C1037EFFFB51160FF37FCB101A972AA" box="[151,427,885,910]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="B964907EFFB51160FF37FCB101A972AA" box="[151,427,885,910]" italics="true" pageId="1" pageNumber="504">Callianassa occidentalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1037EFFFB51160FE42FCB102E172AA" box="[482,739,885,910]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="B964907EFFB51160FE42FCB102E172AA" box="[482,739,885,910]" italics="true" pageId="1" pageNumber="504">Cheramus occidentalis</emphasis>
</taxonomicName>
were actually based on the same individual, and proposed the second replacement name,
<taxonomicName id="4C1037EFFFB51160FE70FC5D02C77296" box="[464,709,921,946]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="profunda">
<emphasis id="B964907EFFB51160FE70FC5D02C77296" box="[464,709,921,946]" italics="true" pageId="1" pageNumber="504">Callianassa profunda</emphasis>
</taxonomicName>
, for Spence Bate’s taxon, because the name
<taxonomicName id="4C1037EFFFB51160FB74FC5D015772F2" authority="Borradaile, 1903" authorityName="Borradaile" authorityYear="1903" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="batei">
<emphasis id="B964907EFFB51160FB74FC5D059E7296" box="[1236,1436,921,946]" italics="true" pageId="1" pageNumber="504">Callianassa batei</emphasis>
<bibRefCitation id="EF81319DFFB51160FF37FC79015772F2" author="Borradaile" box="[151,341,957,982]" pageId="1" pageNumber="504" refString="Borradaile, L. A. (1903) On the classification of the Thalassinidea. Annals and Magazine of Natural History, 12 (7), 534 - 551. http: // dx. doi. org / 10.1080 / 00222930308678891" type="journal article" year="1903">Borradaile, 1903</bibRefCitation>
</taxonomicName>
was also preoccupied by a fossil taxon
<taxonomicName id="4C1037EFFFB51160FC84FC7904BB72F2" authority="Woodward, 1868" authorityName="Woodward" authorityYear="1868" box="[804,1209,957,982]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="batei">
<emphasis id="B964907EFFB51160FC84FC7903E872F2" box="[804,1002,957,982]" italics="true" pageId="1" pageNumber="504">Callianassa batei</emphasis>
Woodward, 1868
</taxonomicName>
.
<bibRefCitation id="EF81319DFFB51160FB67FC7900E372DE" author="Manning" pageId="1" pageNumber="504" refString="Manning, R. B. &amp; Felder, D. L. (1991) Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington, 104, 764 - 792." type="journal article" year="1991">Manning &amp; Felder (1991)</bibRefCitation>
recognized
<taxonomicName id="4C1037EFFFB51160FED0FC2501E172DE" box="[368,483,993,1018]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FED0FC2501E172DE" box="[368,483,993,1018]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
</taxonomicName>
as a valid genus and proposed a new subfamily
<taxonomicName id="4C1037EFFFB51160FBBCFC2504A172DE" box="[1052,1187,993,1018]" class="Malacostraca" family="Callianassidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="subFamily" subFamily="Cheraminae">Cheraminae</taxonomicName>
to accommodate two genera,
<taxonomicName id="4C1037EFFFB51160FF4FFBC10160753A" box="[239,354,1029,1054]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FF4FFBC10160753A" box="[239,354,1029,1054]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1037EFFFB51160FE37FBC102D6753A" authority="Spence Bate, 1888" authorityName="Spence Bate" authorityYear="1888" box="[407,724,1029,1054]" class="Malacostraca" family="Callianassidae" genus="Scallasis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FE37FBC101F8753A" box="[407,506,1029,1054]" italics="true" pageId="1" pageNumber="504">Scallasis</emphasis>
<bibRefCitation id="EF81319DFFB51160FDA1FBC202D6753A" author="Spence" box="[513,724,1030,1054]" pageId="1" pageNumber="504" refString="Spence Bate, C. (1888) Report on the Crustacea Macrura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. &quot; Challenger &quot; during the years 1873 - 76, Zoology, 24 (i - xc), 1 - 942. [plates. 1 - 157]" type="journal article" year="1888">Spence Bate, 1888</bibRefCitation>
</taxonomicName>
. They referred
<taxonomicName id="4C1037EFFFB51160FCDEFBC104FF753A" authority="Biffar, 1973" authorityName="Biffar" authorityYear="1973" box="[894,1277,1029,1054]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="profunda">
<emphasis id="B964907EFFB51160FCDEFBC10472753A" box="[894,1136,1029,1054]" italics="true" pageId="1" pageNumber="504">Callianassa profunda</emphasis>
<bibRefCitation id="EF81319DFFB51160FBD6FBC104FF753A" author="Biffar" box="[1142,1277,1029,1054]" pageId="1" pageNumber="504" refString="Biffar, T. A. (1973) The taxonomic status of Callianassa occidentalis Bate, 1888, and C. batei Borradaile, 1903 (Decapoda, Callianassidae). Crustaceana, 24, 224 - 230. http: // dx. doi. org / 10.1163 / 156854073 X 00399" type="journal article" year="1973">Biffar, 1973</bibRefCitation>
</taxonomicName>
[
<typeStatus id="54ABF2CEFFB51160FAAAFBC2053F753A" box="[1290,1341,1030,1054]" pageId="1" pageNumber="504">type</typeStatus>
species; as
<taxonomicName id="4C1037EFFFB51160FF16FBED02457566" authority="Borradaile, 1903" authorityName="Borradaile" authorityYear="1903" box="[182,583,1065,1090]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="batei">
<emphasis id="B964907EFFB51160FF16FBED01687566" box="[182,362,1065,1090]" italics="true" pageId="1" pageNumber="504">Cheramus batei</emphasis>
(
<bibRefCitation id="EF81319DFFB51160FEDAFBED023C7566" author="Borradaile" box="[378,574,1065,1090]" pageId="1" pageNumber="504" refString="Borradaile, L. A. (1903) On the classification of the Thalassinidea. Annals and Magazine of Natural History, 12 (7), 534 - 551. http: // dx. doi. org / 10.1080 / 00222930308678891" type="journal article" year="1903">Borradaile, 1903</bibRefCitation>
)
</taxonomicName>
],
<taxonomicName id="4C1037EFFFB51160FDFAFBED040E7566" authority="Rathbun, 1901" authorityName="Rathbun" authorityYear="1901" box="[602,1036,1065,1090]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="B964907EFFB51160FDFAFBED035E7565" box="[602,860,1065,1090]" italics="true" pageId="1" pageNumber="504">Callianassa marginata</emphasis>
Rathbun, 1901
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FBBAFBED00D07542" authority="Spence Bate, 1888" authorityName="Spence Bate" authorityYear="1888" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B964907EFFB51160FBBAFBED05037566" box="[1050,1281,1065,1090]" italics="true" pageId="1" pageNumber="504">Cheramus orientalis</emphasis>
<bibRefCitation id="EF81319DFFB51160FAA9FBEE00D07542" author="Spence" pageId="1" pageNumber="504" refString="Spence Bate, C. (1888) Report on the Crustacea Macrura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. &quot; Challenger &quot; during the years 1873 - 76, Zoology, 24 (i - xc), 1 - 942. [plates. 1 - 157]" type="journal article" year="1888">Spence Bate, 1888</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C1037EFFFB51160FEB4FB89032F7542" authority="Le Loeuff &amp; Intes, 1974" authorityName="Le Loeuff &amp; Intes" authorityYear="1974" box="[276,813,1101,1126]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="oblonga">
<emphasis id="B964907EFFB51160FEB4FB8902037542" box="[276,513,1101,1126]" italics="true" pageId="1" pageNumber="504">Callianassa oblonga</emphasis>
Le Loeuff &amp; Intès, 1974
</taxonomicName>
, to
<taxonomicName id="4C1037EFFFB51160FCFDFB8903D27542" box="[861,976,1101,1126]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FCFDFB8903D27542" box="[861,976,1101,1126]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF81319DFFB51160FC42FB8904C47542" box="[994,1222,1101,1126]" pageId="1" pageNumber="504" refString="Tudge, C. C., Poore, G. C. B. &amp; Lemaitre, R. (2000) Preliminary phylogenetic analysis of generic relationships within the Callianassidae and Ctenochelidae (Decapoda: Thalassinidea: Callianassoidea). Journal of Crustacean Biology, 20, 129 - 149." type="journal article">Tudge et al. (2000)</bibRefCitation>
, who performed a phylogenetic analysis estimating generic relationships within the
<taxonomicName id="4C1037EFFFB51160FCDAFBB5041E75AE" box="[890,1052,1137,1162]" class="Malacostraca" family="Callianassidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="family">Callianassidae</taxonomicName>
and
<taxonomicName id="4C1037EFFFB51160FBF4FBB504F575AE" box="[1108,1271,1137,1162]" class="Malacostraca" family="Ctenochelidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="family">Ctenochelidae</taxonomicName>
, supported the recognition of
<taxonomicName id="4C1037EFFFB51160FEEBFB5101BC758A" box="[331,446,1173,1198]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FEEBFB5101BC758A" box="[331,446,1173,1198]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
</taxonomicName>
, and added seven species to that genus:
<taxonomicName id="4C1037EFFFB51160FC1FFB510572758A" authority="Sakai, 1967" authorityName="Sakai" authorityYear="1967" box="[959,1392,1173,1198]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="longicauda">
<emphasis id="B964907EFFB51160FC1FFB5104D0758A" box="[959,1234,1173,1198]" italics="true" pageId="1" pageNumber="504">Callianassa longicauda</emphasis>
Sakai, 1967
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FA22FB52010F75F6" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="praedatrix">
<emphasis id="B964907EFFB51160FA22FB52010F75F6" italics="true" pageId="1" pageNumber="504">C. praedatrix</emphasis>
</taxonomicName>
de Man, 1905,
<taxonomicName id="4C1037EFFFB51160FE1DFB7E025075F5" box="[445,594,1210,1233]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="propinqua">
<emphasis id="B964907EFFB51160FE1DFB7E025075F5" box="[445,594,1210,1233]" italics="true" pageId="1" pageNumber="504">C. propinqua</emphasis>
</taxonomicName>
de Man, 1905,
<taxonomicName id="4C1037EFFFB51160FCA1FB7E03BA75F6" box="[769,952,1209,1234]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="rectangularis">
<emphasis id="B964907EFFB51160FCA1FB7E03BA75F6" box="[769,952,1209,1234]" italics="true" pageId="1" pageNumber="504">C. rectangularis</emphasis>
</taxonomicName>
Ngoc-Ho, 1991,
<taxonomicName id="4C1037EFFFB51160FBDBFB7E04F175F6" box="[1147,1267,1209,1234]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="B964907EFFB51160FBDBFB7E04F175F6" box="[1147,1267,1209,1234]" italics="true" pageId="1" pageNumber="504">C. sibogae</emphasis>
</taxonomicName>
de Man, 1905, and
<taxonomicName id="4C1037EFFFB51160FF67FB1A022275D2" authority="Sakai, 1970" authorityName="Sakai" authorityYear="1970" box="[199,544,1245,1270]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="spinophthalma">
<emphasis id="B964907EFFB51160FF67FB1A018D75D2" box="[199,399,1245,1270]" italics="true" pageId="1" pageNumber="504">C. spinophthalma</emphasis>
Sakai, 1970
</taxonomicName>
.
<bibRefCitation id="EF81319DFFB51160FD8AFB1902B875D2" author="Sakai" box="[554,698,1245,1270]" pageId="1" pageNumber="504" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616." type="journal article" year="2011">Sakai (2011)</bibRefCitation>
rediagnosed
<taxonomicName id="4C1037EFFFB51160FCF1FB1903C675D2" box="[849,964,1245,1270]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FCF1FB1903C675D2" box="[849,964,1245,1270]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
</taxonomicName>
, and assigned the following 16 species to it:
<taxonomicName id="4C1037EFFFB51160FF16FAC5024B743E" authority="Sakai, 2002" authorityName="Sakai" authorityYear="2002" box="[182,585,1281,1306]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="anoploura">
<emphasis id="B964907EFFB51160FF16FAC501BB743E" box="[182,441,1281,1306]" italics="true" pageId="1" pageNumber="504">Callianassa anoploura</emphasis>
<bibRefCitation id="EF81319DFFB51160FE61FAC5024B743E" author="Sakai" box="[449,585,1281,1306]" pageId="1" pageNumber="504" refString="Sakai, K. (2002) Callianassidae (Decapoda, Thalassinidea) in the Andaman Sea, Thailand. In: Bruce, N. L., Berggren, M. &amp; Bussarawit, S. (Eds.), Proceedings of the International Workshop on the Biodiversity of Crustacea of the Andama Sea. Phuket Marine Biological Center Special Publication, 23, 461 - 532." type="journal article" year="2002">Sakai, 2002</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FDF5FAC603C5743E" authority="Dworschak, 2003" authorityName="Dworschak" authorityYear="2003" box="[597,967,1281,1306]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="aqabaensis">
<emphasis id="B964907EFFB51160FDF5FAC602F7743E" box="[597,757,1281,1306]" italics="true" pageId="1" pageNumber="504">C. aqabaensis</emphasis>
<bibRefCitation id="EF81319DFFB51160FD5DFAC503C5743E" author="Dworschak" box="[765,967,1281,1306]" pageId="1" pageNumber="504" refString="Dworschak, P. C. (2003) A new species of ghost shrimp from the Gulf of Aqaba, Red Sea (Crustacea: Decapoda: Callianassidae). Annalen des Naturhistorisches Museums in Wien, 104 (B), 415 - 428." type="journal article" year="2003">Dworschak, 2003</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FC73FAC604E0743E" authority="Sakai, 2002" authorityName="Sakai" authorityYear="2002" box="[979,1250,1281,1306]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="contipes">
<emphasis id="B964907EFFB51160FC73FAC60450743D" box="[979,1106,1282,1305]" italics="true" pageId="1" pageNumber="504">C. contipes</emphasis>
<bibRefCitation id="EF81319DFFB51160FBFAFAC504E0743E" author="Sakai" box="[1114,1250,1281,1306]" pageId="1" pageNumber="504" refString="Sakai, K. (2002) Callianassidae (Decapoda, Thalassinidea) in the Andaman Sea, Thailand. In: Bruce, N. L., Berggren, M. &amp; Bussarawit, S. (Eds.), Proceedings of the International Workshop on the Biodiversity of Crustacea of the Andama Sea. Phuket Marine Biological Center Special Publication, 23, 461 - 532." type="journal article" year="2002">Sakai, 2002</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FB4EFAC6059E743E" box="[1262,1436,1281,1306]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="lobetobensis">
<emphasis id="B964907EFFB51160FB4EFAC6059E743E" box="[1262,1436,1281,1306]" italics="true" pageId="1" pageNumber="504">C. lobetobensis</emphasis>
</taxonomicName>
de Man, 1905,
<taxonomicName id="4C1037EFFFB51160FE9FFAE201DF741A" box="[319,477,1317,1342]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="longicauda">
<emphasis id="B964907EFFB51160FE9FFAE201DF741A" box="[319,477,1317,1342]" italics="true" pageId="1" pageNumber="504">C. longicauda</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FE4BFAE20332741A" authority="Sakai, 2002" authorityName="Sakai" authorityYear="2002" box="[491,816,1317,1342]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="malaccaensis">
<emphasis id="B964907EFFB51160FE4BFAE202A1741A" box="[491,675,1317,1342]" italics="true" pageId="1" pageNumber="504">C. malaccaensis</emphasis>
<bibRefCitation id="EF81319DFFB51160FD09FAE10332741A" author="Sakai" box="[681,816,1317,1342]" pageId="1" pageNumber="504" refString="Sakai, K. (2002) Callianassidae (Decapoda, Thalassinidea) in the Andaman Sea, Thailand. In: Bruce, N. L., Berggren, M. &amp; Bussarawit, S. (Eds.), Proceedings of the International Workshop on the Biodiversity of Crustacea of the Andama Sea. Phuket Marine Biological Center Special Publication, 23, 461 - 532." type="journal article" year="2002">Sakai, 2002</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FC9BFAE203BB741A" box="[827,953,1317,1342]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="B964907EFFB51160FC9BFAE203BB741A" box="[827,953,1317,1342]" italics="true" pageId="1" pageNumber="504">C. modesta</emphasis>
</taxonomicName>
de Man, 1905,
<taxonomicName id="4C1037EFFFB51160FBC8FAE2059A741A" authority="Sakai, 2002" authorityName="Sakai" authorityYear="2002" box="[1128,1432,1317,1342]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="nigroculata">
<emphasis id="B964907EFFB51160FBC8FAE20508741A" box="[1128,1290,1317,1342]" italics="true" pageId="1" pageNumber="504">C. nigroculata</emphasis>
<bibRefCitation id="EF81319DFFB51160FAB1FAE1059A741A" author="Sakai" box="[1297,1432,1317,1342]" pageId="1" pageNumber="504" refString="Sakai, K. (2002) Callianassidae (Decapoda, Thalassinidea) in the Andaman Sea, Thailand. In: Bruce, N. L., Berggren, M. &amp; Bussarawit, S. (Eds.), Proceedings of the International Workshop on the Biodiversity of Crustacea of the Andama Sea. Phuket Marine Biological Center Special Publication, 23, 461 - 532." type="journal article" year="2002">Sakai, 2002</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FF37FA8E011A7446" box="[151,280,1353,1378]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="oblonga">
<emphasis id="B964907EFFB51160FF37FA8E011A7446" box="[151,280,1353,1378]" italics="true" pageId="1" pageNumber="504">C. oblonga</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FE8BFA8E02427446" authority="Sakai, 1988" authorityName="Sakai" authorityYear="1988" box="[299,576,1353,1378]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="B964907EFFB51160FE8BFA8E01AA7446" box="[299,424,1353,1378]" italics="true" pageId="1" pageNumber="504">C. parvula</emphasis>
<bibRefCitation id="EF81319DFFB51160FE14FA8D02427446" author="Sakai" box="[436,576,1353,1378]" pageId="1" pageNumber="504" refString="Sakai, K. (1988) A new genus and five new species of Callianassidae (Crustacea: Decapoda: Thalassinidea) from northern Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences, 5, 51 - 69." type="journal article" year="1988">Sakai, 1988</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FDF0FA8E02E97446" box="[592,747,1353,1378]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="praedatrix">
<emphasis id="B964907EFFB51160FDF0FA8E02E97446" box="[592,747,1353,1378]" italics="true" pageId="1" pageNumber="504">C. praedatrix</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FD5FFA8E03887446" box="[767,906,1353,1378]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="profunda">
<emphasis id="B964907EFFB51160FD5FFA8E03887446" box="[767,906,1353,1378]" italics="true" pageId="1" pageNumber="504">C. profunda</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FC3DFA8E04357445" box="[925,1079,1354,1377]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="propinqua">
<emphasis id="B964907EFFB51160FC3DFA8E04357445" box="[925,1079,1354,1377]" italics="true" pageId="1" pageNumber="504">C. propinqua</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FBEAFA8E05177446" box="[1098,1301,1353,1378]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="spinophthalma">
<emphasis id="B964907EFFB51160FBEAFA8E05177446" box="[1098,1301,1353,1378]" italics="true" pageId="1" pageNumber="504">C. spinophthalma</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1037EFFFB51160FA89FA8D010074A2" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B964907EFFB51160FA89FA8D010074A2" italics="true" pageId="1" pageNumber="504">Cheramus orientalis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1037EFFFB51160FE9FFAAA031374A2" authority="Rodrigues &amp; Manning, 1992" authorityName="Rodrigues &amp; Manning" authorityYear="1992" box="[319,785,1389,1414]" class="Malacostraca" family="Callianassidae" genus="Poti" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="gaucho">
<emphasis id="B964907EFFB51160FE9FFAAA01C574A2" box="[319,455,1389,1414]" italics="true" pageId="1" pageNumber="504">Poti gaucho</emphasis>
Rodrigues &amp; Manning, 1992
</taxonomicName>
.
<bibRefCitation id="EF81319DFFB51160FCBCFAA903A974A2" author="Sakai" box="[796,939,1389,1414]" pageId="1" pageNumber="504" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616." type="journal article" year="2011">Sakai (2011)</bibRefCitation>
also established a new genus,
<taxonomicName id="4C1037EFFFB51160FAA0FAA9059774A2" box="[1280,1429,1389,1414]" class="Malacostraca" family="Callianassidae" genus="Cheramoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FAA0FAA9059774A2" box="[1280,1429,1389,1414]" italics="true" pageId="1" pageNumber="504">Cheramoides</emphasis>
</taxonomicName>
, for
<taxonomicName id="4C1037EFFFB51160FF61FA5501C6748D" box="[193,452,1425,1450]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="B964907EFFB51160FF61FA5501C6748D" box="[193,452,1425,1450]" italics="true" pageId="1" pageNumber="504">Callianassa marginata</emphasis>
</taxonomicName>
, because of the structural difference in the male second pleopod.
<bibRefCitation id="EF81319DFFB51160FB63FA5500E374EA" author="Komai" pageId="1" pageNumber="504" refString="Komai, T. &amp; Fujiwara, Y. (2012) New records of callianassid ghost shrimp (Crustacea: Decapoda: Axiidea) from reducing environments in Kyushu, southwestern Japan. Zootaxa, 3271, 55 - 67." type="journal article" year="2012">Komai &amp; Fujiwara (2012)</bibRefCitation>
described a new species,
<taxonomicName id="4C1037EFFFB51160FDA2FA7102E674EA" box="[514,740,1461,1486]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="cavifrons">
<emphasis id="B964907EFFB51160FDA2FA7102E674EA" box="[514,740,1461,1486]" italics="true" pageId="1" pageNumber="504">Cheramus cavifrons</emphasis>
</taxonomicName>
, from
<collectingCountry id="F3070CFCFFB51160FC8FFA72037074E9" box="[815,882,1462,1485]" name="Japan" pageId="1" pageNumber="504">Japan</collectingCountry>
.
<bibRefCitation id="EF81319DFFB51160FCDCFA71044C74EA" author="Dworschak" box="[892,1102,1461,1486]" pageId="1" pageNumber="504" refString="Dworschak, P. C. (2012) On the identities of Callianassa bouvieri Nobili, 1904, C. maldivensis Borradaile, 1904, and C. gravieri Nobili, 1905 (Crustacea: Decapoda: Callianassidae): a morphometric approach. Zootaxa, 3149, 39 - 56." type="journal article" year="2012">Dworschak (2012)</bibRefCitation>
has clarified that
<taxonomicName id="4C1037EFFFB51160FAB6FA71012F74D6" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="rectangularis">
<emphasis id="B964907EFFB51160FAB6FA71012F74D6" italics="true" pageId="1" pageNumber="504">Callianassa rectangularis</emphasis>
</taxonomicName>
, once synonymized with
<taxonomicName id="4C1037EFFFB51160FDFDFA1D03EB74D6" authority="Nobili, 1904" authorityName="Nobili" authorityYear="1904" box="[605,1001,1497,1522]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="bouvieri">
<emphasis id="B964907EFFB51160FDFDFA1D034974D6" box="[605,843,1497,1522]" italics="true" pageId="1" pageNumber="504">Callianassa bouvieri</emphasis>
Nobili, 1904
</taxonomicName>
by
<bibRefCitation id="EF81319DFFB51160FBBCFA1D04A874D6" author="Sakai" box="[1052,1194,1497,1522]" pageId="1" pageNumber="504" refString="Sakai, K. (1999) Synopsis of the family Callianassidae, with keys to subfamilies, genera and species, and the description of new taxa (Crustacea: Decapoda: Thalassinidea). Zoologische Verhandelingen, 326, 1 - 152." type="journal article" year="1999">Sakai (1999</bibRefCitation>
;
<bibRefCitation id="EF81319DFFB51160FB1AFA1D04F474D6" author="Sakai" box="[1210,1270,1497,1522]" pageId="1" pageNumber="504" refString="Sakai, K. (2005) Callianassoidea of the world (Decapoda, Thalassinidea). Crustaceana Monographs, 4, 1 - 285." type="journal article" year="2005">2005</bibRefCitation>
), is actually a junior synonym of
<taxonomicName id="4C1037EFFFB51160FEDEFA3A03837732" authority="Borradaile, 1904" authorityName="Borradaile" authorityYear="1904" box="[382,897,1533,1558]" class="Malacostraca" family="Callianassidae" genus="Paratrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="maldivensis">
<emphasis id="B964907EFFB51160FEDEFA3A029F7732" box="[382,669,1533,1558]" italics="true" pageId="1" pageNumber="504">Paratrypaea maldivensis</emphasis>
(Borradaile, 1904)
</taxonomicName>
. Sakai &amp; Türkay (2014) transferred
<taxonomicName id="4C1037EFFFB51160FA9AFA3A0211771E" authority="Guzman &amp; Thatje, 2003" authorityName="Guzman &amp; Thatje" authorityYear="2003" class="Malacostraca" family="Callianassidae" genus="Biffarius" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B964907EFFB51160FA9AFA3A00FE771E" italics="true" pageId="1" pageNumber="504">Biffarius pacificus</emphasis>
<bibRefCitation id="EF81319DFFB51160FEA3F9E60211771E" author="Guzman" box="[259,531,1569,1594]" pageId="1" pageNumber="504" refString="Guzman, G. L. &amp; Thatje, S. (2003) Biffarius pacificus, a new species of the Callianassidae (Crustacea: Decapoda: Thalassinidea) from northern Chile. Scientia Marina, 67, 293 - 298. http: // dx. doi. org / 10.3989 / scimar. 2003.67 n 3293" type="journal article" year="2003">Guzmán &amp; Thatje, 2003</bibRefCitation>
</taxonomicName>
to
<taxonomicName id="4C1037EFFFB51160FD97F9E502A8771E" box="[567,682,1569,1594]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FD97F9E502A8771E" box="[567,682,1569,1594]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAF4C6CFFB51160FF67F981011477CA" blockId="1.[151,1437,597,2026]" pageId="1" pageNumber="504">
The status of the subfamily
<taxonomicName id="4C1037EFFFB51160FD8AF98102B6777A" box="[554,692,1605,1630]" class="Malacostraca" family="Callianassidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="subFamily" subFamily="Cheraminae">Cheraminae</taxonomicName>
, originally proposed by
<bibRefCitation id="EF81319DFFB51160FC46F9810526777A" author="Manning" box="[998,1316,1605,1630]" pageId="1" pageNumber="504" refString="Manning, R. B. &amp; Felder, D. L. (1991) Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington, 104, 764 - 792." type="journal article" year="1991">Manning &amp; Felder (1991)</bibRefCitation>
, is rather questionable, although
<bibRefCitation id="EF81319DFFB51160FE02F9AD028077A6" box="[418,642,1641,1666]" pageId="1" pageNumber="504" refString="Tudge, C. C., Poore, G. C. B. &amp; Lemaitre, R. (2000) Preliminary phylogenetic analysis of generic relationships within the Callianassidae and Ctenochelidae (Decapoda: Thalassinidea: Callianassoidea). Journal of Crustacean Biology, 20, 129 - 149." type="journal article">Tudge et al. (2000)</bibRefCitation>
supported the recognition of this subfamily. Molecular phylogenetic analysis by
<bibRefCitation id="EF81319DFFB51160FE81F94902377782" author="Felder" box="[289,565,1677,1702]" pageId="1" pageNumber="504" refString="Felder, D. L. &amp; Robles, R. (2009) Molecular phylogeny of the family Callianassidae based on preliminary analyses of two mitochondrial genes. In: Martin, J. W., Crandall, K. A. &amp; Felder, D. L. (Eds.), Crustacean Issues 18. Decapod Crustacean Phylogenetics. Taylor &amp; Francis / CRC Press, Boca Raton, Florida, pp. 327 - 342." type="journal article" year="2009">Felder &amp; Robles (2009)</bibRefCitation>
estimated that two species referred to
<taxonomicName id="4C1037EFFFB51160FC5AF949046F7782" box="[1018,1133,1677,1702]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FC5AF949046F7782" box="[1018,1133,1677,1702]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
</taxonomicName>
are subordinated within a group of species referred to
<taxonomicName id="4C1037EFFFB51160FE7CF975028377EE" box="[476,641,1713,1738]" class="Malacostraca" family="Callianassidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="subFamily" subFamily="Callianassinae">Callianassinae</taxonomicName>
. We follow
<bibRefCitation id="EF81319DFFB51160FCADF975041C77EE" author="Felder" box="[781,1054,1713,1738]" pageId="1" pageNumber="504" refString="Felder, D. L. &amp; Robles, R. (2009) Molecular phylogeny of the family Callianassidae based on preliminary analyses of two mitochondrial genes. In: Martin, J. W., Crandall, K. A. &amp; Felder, D. L. (Eds.), Crustacean Issues 18. Decapod Crustacean Phylogenetics. Taylor &amp; Francis / CRC Press, Boca Raton, Florida, pp. 327 - 342." type="journal article" year="2009">Felder &amp; Robles (2009)</bibRefCitation>
and abandon
<taxonomicName id="4C1037EFFFB51160FB60F975054577EE" box="[1216,1351,1713,1738]" class="Malacostraca" family="Callianassidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="subFamily" subFamily="Cheraminae">Cheraminae</taxonomicName>
for the time being.
</paragraph>
<paragraph id="8BAF4C6CFFB51163FF67F93D053F70AC" blockId="1.[151,1437,597,2026]" lastBlockId="2.[151,1436,151,789]" lastPageId="2" lastPageNumber="505" pageId="1" pageNumber="504">
Possibly diagnostic characters generally shared by species assigned to
<taxonomicName id="4C1037EFFFB51160FC4AF93D045F7636" box="[1002,1117,1785,1810]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB51160FC4AF93D045F7636" box="[1002,1117,1785,1810]" italics="true" pageId="1" pageNumber="504">Cheramus</emphasis>
</taxonomicName>
include: rostrum spiniform; eyestalk with cornea being subterminal and lateral; antennular peduncle not much longer or stouter than antennal peduncle; distal margin of merus of third maxilliped not strongly produced; chelipeds unequal and dissimilar, of which the merus of major side lacks a prominent ventral hook, but sometimes with a ventral spine; female second pleopod lacking appendix interna; and third to fifth pleopods relatively narrow, with slender, digitiform appendices internae distinctly projecting beyond mesial margins of endopods. The development of the male first and second pleopods is known for the
<typeStatus id="54ABF2CEFFB51160FE62F81601F076CE" box="[450,498,2002,2026]" pageId="1" pageNumber="504">type</typeStatus>
species,
<taxonomicName id="4C1037EFFFB51160FDF9F815034176CE" box="[601,835,2001,2026]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="profundus">
<emphasis id="B964907EFFB51160FDF9F815034176CE" box="[601,835,2001,2026]" italics="true" pageId="1" pageNumber="504">Cheramus profundus</emphasis>
</taxonomicName>
: first pleopod slender, bi-segmented; second pleopod biramous (
<bibRefCitation id="EF81319DFFB61163FEB2FF53019D7194" author="Biffar" box="[274,415,151,176]" pageId="2" pageNumber="505" refString="Biffar, T. A. (1973) The taxonomic status of Callianassa occidentalis Bate, 1888, and C. batei Borradaile, 1903 (Decapoda, Callianassidae). Crustaceana, 24, 224 - 230. http: // dx. doi. org / 10.1163 / 156854073 X 00399" type="journal article" year="1973">Biffar 1973</bibRefCitation>
). It should be noted that male characters remain unknown for many species presently referred to
<taxonomicName id="4C1037EFFFB61163FEB6FF78018B71F1" box="[278,393,188,213]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB61163FEB6FF78018B71F1" box="[278,393,188,213]" italics="true" pageId="2" pageNumber="505">Cheramus</emphasis>
</taxonomicName>
. Furthermore,
<bibRefCitation id="EF81319DFFB61163FD94FF7802C771F0" author="Sakai" box="[564,709,188,213]" pageId="2" pageNumber="505" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616." type="journal article" year="2011">Sakai (2011)</bibRefCitation>
referred several species similar to the species of
<taxonomicName id="4C1037EFFFB61163FB59FF78056E71F1" box="[1273,1388,188,213]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB61163FB59FF78056E71F1" box="[1273,1388,188,213]" italics="true" pageId="2" pageNumber="505">Cheramus</emphasis>
</taxonomicName>
, for which male characters are unknown, to
<taxonomicName id="4C1037EFFFB61163FDD1FF25036371DC" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[625,865,223,248]" class="Malacostraca" family="Callianassidae" genus="Trypaea" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB61163FDD1FF2502CF71DC" box="[625,717,225,248]" italics="true" pageId="2" pageNumber="505">Trypaea</emphasis>
Dana, 1852
</taxonomicName>
, and the classification has been followed by his subsequent works (
<bibRefCitation id="EF81319DFFB61163FED2FEC001F47038" author="Sakai" box="[370,502,260,285]" pageId="2" pageNumber="505" refString="Sakai, K. (2010) Callianassoidea from the Gulf of Tonkin and the Red Sea, in the Zoological Museum of Moscow University (Decapoda, Thalassinidea). Crustaceana, 83, 1431 - 1467. http: // dx. doi. org / 10.1163 / 001121610 X 538174" type="journal article" year="2010">Sakai 2010</bibRefCitation>
;
<bibRefCitation id="EF81319DFFB61163FDA4FEC002FD7038" author="Sakai" box="[516,767,260,285]" pageId="2" pageNumber="505" refString="Sakai, K. &amp; Turkay, M. (2012) A collection of Thalassinidea Latreille, 1831 (Decapoda, Pleocyemata) from the Senckenberg Forschungsinstitut and Natural History Museum, Frankfurt am Main. Crustaceana, 85, 723 - 765. http: // dx. doi. org / 10.1163 / 156854012 X 643735" type="journal article" year="2012">Sakai &amp; Türkay 2012</bibRefCitation>
, 2014; Sakai et al. 2014). Many workers have pointed out that a thorough revision is necessary to diagnose
<taxonomicName id="4C1037EFFFB61163FD69FEE3033E7064" box="[713,828,295,320]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB61163FD69FEE3033E7064" box="[713,828,295,320]" italics="true" pageId="2" pageNumber="505">Cheramus</emphasis>
</taxonomicName>
and other genera of
<taxonomicName id="4C1037EFFFB61163FB8AFEE304CE7064" box="[1066,1228,295,320]" class="Malacostraca" family="Callianassidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="subFamily" subFamily="Callianassinae">Callianassinae</taxonomicName>
(e.g.,
<bibRefCitation id="EF81319DFFB61163FAB5FEE300D17040" author="Tudge" pageId="2" pageNumber="505" refString="Tudge, C. C., Poore, G. C. B. &amp; Lemaitre, R. (2000) Preliminary phylogenetic analysis of generic relationships within the Callianassidae and Ctenochelidae (Decapoda: Thalassinidea: Callianassoidea). Journal of Crustacean Biology, 20, 129 - 149." type="journal article" year="2000">Tudge et al. 2000</bibRefCitation>
;
<bibRefCitation id="EF81319DFFB61163FF7EFE8801A27040" author="Dworschak" box="[222,416,332,357]" pageId="2" pageNumber="505" refString="Dworschak, P. C. (2003) A new species of ghost shrimp from the Gulf of Aqaba, Red Sea (Crustacea: Decapoda: Callianassidae). Annalen des Naturhistorisches Museums in Wien, 104 (B), 415 - 428." type="journal article" year="2003">Dworschak 2003</bibRefCitation>
;
<bibRefCitation id="EF81319DFFB61163FE0CFE8902547040" author="Ngoc-Ho" box="[428,598,332,357]" pageId="2" pageNumber="505" refString="Ngoc-Ho, N. (2003) European and Mediterranean Thalassinidea (Crustacea, Decapoda). Zoosystema, 25, 439 - 555." type="journal article" year="2003">Ngoc-Ho 2003</bibRefCitation>
,
<bibRefCitation id="EF81319DFFB61163FDC2FE88029F7040" author="Ngoc-Ho" box="[610,669,332,356]" pageId="2" pageNumber="505" refString="Ngoc-Ho, N. (2014) Six species of Axiidea and Gebiidea from the Indo-West Pacific (Crustacea, Decapoda). Zoosystema, 36, 545 - 561. http: // dx. doi. org / 10.5252 / z 2014 n 3 a 1" type="journal article" year="2014">2014</bibRefCitation>
;
<bibRefCitation id="EF81319DFFB61163FD09FE89032F7040" author="Poore" box="[681,813,332,356]" pageId="2" pageNumber="505" refString="Poore, G. C. B. (2008) Thalassinidean shrimps (Crustacea: Decapoda) from north-western Australia, including five new species. In: Jones, D. S. (Ed.), Crustaceans Collected by the Western Australian Museum / Woodside Energy Ltd. Partnership to Explore the Marine Biodiversity of the Dampier Archipelago, Western Australia, 1998 - 2002. Records of the Western Australian Museum, Supplement 73, 161 - 179." type="book chapter" year="2008">Poore 2008</bibRefCitation>
;
<bibRefCitation id="EF81319DFFB61163FC98FE8804357040" author="Felder" box="[824,1079,332,357]" pageId="2" pageNumber="505" refString="Felder, D. L. &amp; Robles, R. (2009) Molecular phylogeny of the family Callianassidae based on preliminary analyses of two mitochondrial genes. In: Martin, J. W., Crandall, K. A. &amp; Felder, D. L. (Eds.), Crustacean Issues 18. Decapod Crustacean Phylogenetics. Taylor &amp; Francis / CRC Press, Boca Raton, Florida, pp. 327 - 342." type="journal article" year="2009">Felder &amp; Robles 2009</bibRefCitation>
). The generic assignment of the two new species made in this study is also merely provisional, because no male specimens were available.
</paragraph>
<paragraph id="8BAF4C6CFFB61163FF67FE50033F7231" blockId="2.[151,1436,151,789]" pageId="2" pageNumber="505">
Nevertheless, we can say that Sakai &amp; Türkay’s (2014) assignment of
<taxonomicName id="4C1037EFFFB61163FC72FE50049D7088" box="[978,1183,404,428]" class="Malacostraca" family="Callianassidae" genus="Biffarius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B964907EFFB61163FC72FE50049D7088" box="[978,1183,404,428]" italics="true" pageId="2" pageNumber="505">Biffarius pacificus</emphasis>
</taxonomicName>
to
<taxonomicName id="4C1037EFFFB61163FB62FE5005377089" box="[1218,1333,404,429]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB61163FB62FE5005377089" box="[1218,1333,404,429]" italics="true" pageId="2" pageNumber="505">Cheramus</emphasis>
</taxonomicName>
is highly questionable.
<bibRefCitation id="EF81319DFFB61163FE93FE7301C070F4" author="Sakai" box="[307,450,439,464]" pageId="2" pageNumber="505" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616." type="journal article" year="2011">Sakai (2011)</bibRefCitation>
transferred
<taxonomicName id="4C1037EFFFB61163FDE9FE7D02CF70F4" box="[585,717,440,464]" class="Malacostraca" family="Callianassidae" genus="Biffarius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B964907EFFB61163FDE9FE7D02CF70F4" box="[585,717,440,464]" italics="true" pageId="2" pageNumber="505">B. pacificus</emphasis>
</taxonomicName>
to
<taxonomicName id="4C1037EFFFB61163FD52FE73048470F4" authority="Manning &amp; Felder, 1992" authorityName="Manning &amp; Felder" authorityYear="1992" box="[754,1158,439,464]" class="Malacostraca" family="Callianassidae" genus="Gilvossius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB61163FD52FE73036570F4" box="[754,871,439,464]" italics="true" pageId="2" pageNumber="505">Gilvossius</emphasis>
Manning &amp; Felder, 1992
</taxonomicName>
, but this action was only based on character of the male second pleopod, which was not properly described by
<bibRefCitation id="EF81319DFFB61163FBD9FE18059E70D0" author="Guzman" box="[1145,1436,476,501]" pageId="2" pageNumber="505" refString="Guzman, G. L. &amp; Thatje, S. (2003) Biffarius pacificus, a new species of the Callianassidae (Crustacea: Decapoda: Thalassinidea) from northern Chile. Scientia Marina, 67, 293 - 298. http: // dx. doi. org / 10.3989 / scimar. 2003.67 n 3293" type="journal article" year="2003">Guzmán &amp; Thatje (2003)</bibRefCitation>
(Sakai estimated that the species does not have the male second pleopod). Later, Sakai &amp; Türkay (2014) found that
<taxonomicName id="4C1037EFFFB61163FF37FDE1011C7318" box="[151,286,548,572]" class="Malacostraca" family="Callianassidae" genus="Biffarius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B964907EFFB61163FF37FDE1011C7318" box="[151,286,548,572]" italics="true" pageId="2" pageNumber="505">B. pacificus</emphasis>
</taxonomicName>
actually has a male second pleopod, and used this evidence to reassign
<taxonomicName id="4C1037EFFFB61163FBCFFDE104F57318" box="[1135,1271,548,572]" class="Malacostraca" family="Callianassidae" genus="Biffarius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B964907EFFB61163FBCFFDE104F57318" box="[1135,1271,548,572]" italics="true" pageId="2" pageNumber="505">B. pacificus</emphasis>
</taxonomicName>
to
<taxonomicName id="4C1037EFFFB61163FA82FDE005977319" box="[1314,1429,548,573]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB61163FA82FDE005977319" box="[1314,1429,548,573]" italics="true" pageId="2" pageNumber="505">Cheramus</emphasis>
</taxonomicName>
. Furthermore, the authors claimed that the absence of appendices interna and masculina on the male second pleopod is sufficient to differentiate
<taxonomicName id="4C1037EFFFB61163FE78FDA8024973A1" box="[472,587,620,645]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB61163FE78FDA8024973A1" box="[472,587,620,645]" italics="true" pageId="2" pageNumber="505">Cheramus</emphasis>
</taxonomicName>
from
<taxonomicName id="4C1037EFFFB61163FD35FDA8030B73A1" box="[661,777,620,645]" class="Malacostraca" family="Callianassidae" genus="Gilvossius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB61163FD35FDA8030B73A1" box="[661,777,620,645]" italics="true" pageId="2" pageNumber="505">Gilvossius</emphasis>
</taxonomicName>
, but this character is common in
<taxonomicName id="4C1037EFFFB61163FB33FDA8053A73A1" box="[1171,1336,620,645]" class="Malacostraca" family="Callianassidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="subFamily" subFamily="Callianassinae">Callianassinae</taxonomicName>
, and not appropriate in this case. According to the original description by
<bibRefCitation id="EF81319DFFB61163FC20FD5404A1738C" author="Guzman" box="[896,1187,655,680]" pageId="2" pageNumber="505" refString="Guzman, G. L. &amp; Thatje, S. (2003) Biffarius pacificus, a new species of the Callianassidae (Crustacea: Decapoda: Thalassinidea) from northern Chile. Scientia Marina, 67, 293 - 298. http: // dx. doi. org / 10.3989 / scimar. 2003.67 n 3293" type="journal article" year="2003">Guzmán &amp; Thatje (2003)</bibRefCitation>
,
<taxonomicName id="4C1037EFFFB61163FB11FD540583738C" box="[1201,1409,656,680]" class="Malacostraca" family="Callianassidae" genus="Biffarius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B964907EFFB61163FB11FD540583738C" box="[1201,1409,656,680]" italics="true" pageId="2" pageNumber="505">Biffarius pacificus</emphasis>
</taxonomicName>
is diagnosed by having an obsolescent rostrum and stubby appendices internae on the third to fifth pleopods slightly projecting beyond mesial margins of endopods, being distinct from other species assigned to
<taxonomicName id="4C1037EFFFB61163FB1CFD13052D73D4" box="[1212,1327,727,752]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB61163FB1CFD13052D73D4" box="[1212,1327,727,752]" italics="true" pageId="2" pageNumber="505">Cheramus</emphasis>
</taxonomicName>
. It seems reasonable to retain the taxon in
<taxonomicName id="4C1037EFFFB61163FDA4FD3802647230" box="[516,614,764,788]" class="Malacostraca" family="Callianassidae" genus="Biffarius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B964907EFFB61163FDA4FD3802647230" box="[516,614,764,788]" italics="true" pageId="2" pageNumber="505">Biffarius</emphasis>
</taxonomicName>
for the time being.
</paragraph>
</subSubSection>
</treatment>
</document>
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