treatments-xml/data/03/B9/AB/03B9AB31FFE94437FE21FD7A0FBA8731.xml

<document id="5DD8A9E84C0C763D2ED722EFAB7F0ED6" ID-DOI="10.5852/ejt.2020.720.1121" ID-GBIF-Dataset="3e326079-e0cd-4a5f-9b58-5d24eccb331b" ID-ISSN="2118-9773" ID-Zenodo-Dep="4106763" ID-ZooBank="F0B07E69-BB75-4AB2-ABD4-339254FBC390" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1603123887628" checkinUser="valdenar" docAuthor="Risaro, Jessica, Williams, Gary C., Pereyra, Daniela &amp; Lauretta, Daniel" docDate="2020" docId="03B9AB31FFE94437FE21FD7A0FBA8731" docLanguage="en" docName="ejt-720_risaro_williams_pereyra.pdf" docOrigin="European Journal of Taxonomy 720" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Umbellula pomona Risaro, Williams &amp; Lauretta 2020, sp. nov." docType="treatment" docUuid="77FCDEF4-AAE3-4BB5-BAB2-1EDAA4B19439" docUuidSource="ZooBank" docVersion="15" lastPageNumber="132" masterDocId="FF80D349FFEF443CFFEAFFFB0B3B8536" masterDocTitle="Umbellula pomona sp. nov., a new sea pen from Mar del Plata Submarine Canyon (Cnidaria: Octocorallia: Pennatulacea)" masterLastPageNumber="143" masterPageNumber="121" pageNumber="127" updateTime="1733486937508" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9E135CD4579E08769085E8C320AEA7A3">Umbellula pomona sp. nov., a new sea pen from Mar del Plata Submarine Canyon (Cnidaria: Octocorallia: Pennatulacea)</mods:title>
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<mods:namePart id="9BB4EE6D0258ED7C58F4A441B4B7895F">Risaro, Jessica</mods:namePart>
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<mods:affiliation id="20022C448FC3046431399B1201D44AB3">Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” - CONICET, Av. Ángel Gallardo 470, Buenos Aires, Argentina.</mods:affiliation>
<mods:nameIdentifier id="C9104939662808F598CB014D8A7A90DD" type="email">jessicarisaro@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4B8920FB46C52C12D3D39B5CD725BCA3">Williams, Gary C.</mods:namePart>
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<mods:affiliation id="7EA674E7F6B11DF8AEF845A599EA0DB2">Department of Invertebrate Zoology and Geology, California Academy of Sciences, Golden Gate Park, San Francisco, California 94118, USA.</mods:affiliation>
<mods:nameIdentifier id="09C3D2D724428B22D56B9F1F2936FAD6" type="email">gwilliams@calacademy.org</mods:nameIdentifier>
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<mods:namePart id="6409FDC57D4C399F332BF321B5B09AD6">Pereyra, Daniela</mods:namePart>
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<mods:affiliation id="C7491AE4276F312CEA2B1B6843B209B7">Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” - CONICET, Av. Ángel Gallardo 470, Buenos Aires, Argentina.</mods:affiliation>
<mods:nameIdentifier id="CD91CD8C586554527E1B5A8A7C23DE67" type="email">mdr.pereyra@gmail.com</mods:nameIdentifier>
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<mods:namePart id="778C48938C20C01145900650E2FE11EB">Lauretta, Daniel</mods:namePart>
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<mods:affiliation id="EEEC05C3CE52855455B9CBECE70E938D">Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” - CONICET, Av. Ángel Gallardo 470, Buenos Aires, Argentina.</mods:affiliation>
<mods:nameIdentifier id="96A1B5004D5FEB0B5B266AA9847B20F0" type="email">dlauretta@gmail.com</mods:nameIdentifier>
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<mods:title id="BE8E6D8C2833F1EFE98134C5FA7F159A">European Journal of Taxonomy</mods:title>
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<mods:date id="181093232E0514200C34345714958B3F">2020</mods:date>
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<subSubSection id="C30A49ACFFE9443AFE21FD7A0F5287AA" box="[459,1129,641,668]" pageId="6" pageNumber="127" type="nomenclature">
<paragraph id="8BAF1A27FFE9443AFE21FD7A0F5287AA" blockId="6.[349,1238,641,738]" box="[459,1129,641,668]" pageId="6" pageNumber="127">
<heading id="D0E7AD4BFFE9443AFE21FD7A0F5287AA" box="[459,1129,641,668]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="127" reason="2">
<taxonomicName id="4C1061A4FFE9443AFE21FD7A0F3C87AA" ID-CoL="8RW37" authority="Risaro, Williams &amp; Lauretta" authorityName="Risaro, Williams &amp; Lauretta" authorityYear="2020" box="[459,1031,641,668]" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="6" pageNumber="127" phylum="Cnidaria" rank="species" species="pomona" status="sp. nov.">
<emphasis id="B964C635FFE9443AFE21FD7A098887AD" bold="true" box="[459,691,641,668]" italics="true" pageId="6" pageNumber="127">Umbellula pomona</emphasis>
Risaro, Williams &amp; Lauretta
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<taxonomicNameLabel id="A2577B4EFFE9443AFBE4FD790F5287AA" box="[1038,1129,642,668]" pageId="6" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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urn:lsid:zoobank.org:act: 
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<paragraph id="8BAF1A27FFE9443AFD0DFD3C087687D4" blockId="6.[349,1238,641,738]" box="[743,845,711,738]" pageId="6" pageNumber="127">
<figureCitation id="132B06A2FFE9443AFD0DFD3C087687D4" box="[743,845,711,738]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="7.[189,232,907,933]" captionStartId-1="7.[189,232,1918,1944]" captionStartId-2="8.[189,232,1952,1978]" captionStartId-3="9.[189,232,1958,1984]" captionStartId-4="10.[189,232,1958,1984]" captionStartId-5="11.[189,232,1958,1984]" captionTargetBox-0="[195,1392,271,864]" captionTargetBox-1="[239,1349,1135,1879]" captionTargetBox-2="[189,1392,830,1909]" captionTargetBox-3="[195,1391,842,1912]" captionTargetBox-4="[195,1393,840,1913]" captionTargetBox-5="[200,1392,834,1913]" captionTargetId-0="figure-12@7.[189,1398,265,870]" captionTargetId-1="figure-85@7.[234,1354,1130,1884]" captionTargetId-2="figure-235@8.[189,1398,824,1909]" captionTargetId-3="figure-224@9.[189,1398,834,1919]" captionTargetId-4="figure-271@10.[189,1398,834,1919]" captionTargetId-5="figure-163@11.[189,1398,828,1913]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionText-0="Fig. 2. A. General aspect of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. A. Holotype (MACN-IN 42608). B. Detail of the terminal cluster of paratype A (MACN-IN 42609), showing three autozooids that form the terminal cluster and amplifications of sclerites (up) and siphonozooids (down). Abbreviations: CP = central polyp; LP = lateral polyp; Pd = peduncle; R = rachis; T = tentacle; S = siphonozooids; Scl = sclerites." captionText-1="Fig. 3. Transversal section of the central axis of paratype B of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42609). A. Transversal section of the central axis between the peduncle and the rachis. B. Transversal section of the central axis near the terminal cluster." captionText-2="Fig. 4. Variability of sizes and ornamentations of the rachis’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." captionText-3="Fig. 5. Variability of sizes and ornamentations of the polyps’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." captionText-4="Fig. 6. Variability of sizes and ornamentations of the tentacles’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." captionText-5="Fig. 7. Variability of sizes and ornamentations of the pinnules’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." figureDoi-0="http://doi.org/10.5281/zenodo.4106767" figureDoi-1="http://doi.org/10.5281/zenodo.4106769" figureDoi-2="http://doi.org/10.5281/zenodo.4106771" figureDoi-3="http://doi.org/10.5281/zenodo.4106773" figureDoi-4="http://doi.org/10.5281/zenodo.4106775" figureDoi-5="http://doi.org/10.5281/zenodo.4106777" httpUri-0="https://zenodo.org/record/4106767/files/figure.png" httpUri-1="https://zenodo.org/record/4106769/files/figure.png" httpUri-2="https://zenodo.org/record/4106771/files/figure.png" httpUri-3="https://zenodo.org/record/4106773/files/figure.png" httpUri-4="https://zenodo.org/record/4106775/files/figure.png" httpUri-5="https://zenodo.org/record/4106777/files/figure.png" pageId="6" pageNumber="127">Figs 2–7</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="8BAF1A27FFE9443AFF57FCF20AF28615" blockId="6.[189,457,777,803]" box="[189,457,777,803]" pageId="6" pageNumber="127">
<heading id="D0E7AD4BFFE9443AFF57FCF20AF28615" bold="true" box="[189,457,777,803]" fontSize="11" level="3" pageId="6" pageNumber="127" reason="3">
<emphasis id="B964C635FFE9443AFF57FCF20AF28615" bold="true" box="[189,457,777,803]" pageId="6" pageNumber="127">Differential diagnosis</emphasis>
</heading>
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<taxonomicName id="4C1061A4FFE9443AFF57FCCC0A998667" authority="Risaro &amp; Williams &amp; Pereyra &amp; Lauretta, 2020" authorityName="Risaro &amp; Williams &amp; Pereyra &amp; Lauretta" authorityYear="2020" box="[189,418,823,849]" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="6" pageNumber="127" phylum="Cnidaria" rank="species" species="pomona" status="sp. nov.">
<emphasis id="B964C635FFE9443AFF57FCCC0A998667" box="[189,418,823,849]" italics="true" pageId="6" pageNumber="127">Umbellula pomona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2577B4EFFE9443AFE41FCC309328664" box="[427,521,824,850]" pageId="6" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
is a spiculated 
<taxonomicName id="4C1061A4FFE9443AFD2DFCCC08788667" authorityName="Cuvier" authorityYear="1798" box="[711,835,823,849]" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="6" pageNumber="127" phylum="Cnidaria" rank="genus">
<emphasis id="B964C635FFE9443AFD2DFCCC08788667" box="[711,835,823,849]" italics="true" pageId="6" pageNumber="127">Umbellula</emphasis>
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with three autozooids in its terminal cluster, a central well-developed polyp and two lateral, symmetric and smaller ones. It presents large sclerites in all its tissues as well as siphonozooids all along the rachis. Its central axis is circular in cross section all along its extension, and does not vary throughout the colony.
</paragraph>
</subSubSection>
<subSubSection id="C30A49ACFFE9443AFF57FC1809D38178" pageId="6" pageNumber="127" type="etymology">
<paragraph id="8BAF1A27FFE9443AFF57FC180A7F86CB" blockId="6.[189,324,995,1021]" box="[189,324,995,1021]" pageId="6" pageNumber="127">
<heading id="D0E7AD4BFFE9443AFF57FC180A7F86CB" bold="true" box="[189,324,995,1021]" fontSize="11" level="3" pageId="6" pageNumber="127" reason="3">
<emphasis id="B964C635FFE9443AFF57FC180A7F86CB" bold="true" box="[189,324,995,1021]" pageId="6" pageNumber="127">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF1A27FFE9443AFF57FBEA09D38178" blockId="6.[189,1399,1041,1103]" pageId="6" pageNumber="127">
The species is named after the birthplace of the first author (JR), Pomona (
<collectingRegion id="49D4D4C5FFE9443AFBDAFBEA0F8B811A" box="[1072,1200,1041,1068]" country="Argentina" name="Rio Negro" pageId="6" pageNumber="127">Río Negro</collectingRegion>
, 
<collectingCountry id="F3075AB7FFE9443AFB51FBEA0E0E811A" box="[1211,1333,1041,1068]" name="Argentina" pageId="6" pageNumber="127">Argentina</collectingCountry>
). The word ‘pomona’ is used as a noun in opposition.
</paragraph>
</subSubSection>
<subSubSection id="C30A49ACFFE9443AFF57FB8E090180C0" pageId="6" pageNumber="127" type="materials_examined">
<paragraph id="8BAF1A27FFE9443AFF57FB8E0A9181B9" blockId="6.[189,426,1141,1167]" box="[189,426,1141,1167]" pageId="6" pageNumber="127">
<heading id="D0E7AD4BFFE9443AFF57FB8E0A9181B9" bold="true" box="[189,426,1141,1167]" fontSize="11" level="3" pageId="6" pageNumber="127" reason="3">
<emphasis id="B964C635FFE9443AFF57FB8E0A9181B9" bold="true" box="[189,426,1141,1167]" pageId="6" pageNumber="127">Material examined</emphasis>
</heading>
</paragraph>
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<emphasis id="B964C635FFE9443AFF00FB5F0A678188" bold="true" box="[234,348,1188,1214]" pageId="6" pageNumber="127">
<typeStatus id="54ABA485FFE9443AFF00FB5F0A678188" box="[234,348,1188,1214]" pageId="6" pageNumber="127" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAF1A27FFE9443AFF57FB3309B4801E" blockId="6.[189,1399,1188,1320]" pageId="6" pageNumber="127">
<materialsCitation id="3B78107AFFE9443AFF57FB3309B4801E" ID-GBIF-Occurrence="2884010301" collectingDate="2013-09" collectionCode="MACN-IN" collectorName="Daniel Lauretta" elevation="2934" latitude="-38.031883" location="Mar del Plata Submarine Canyon" longLatPrecision="1" longitude="-53.65447" pageId="6" pageNumber="127" specimenCode="MACN-IN 42608" specimenCount="1" typeStatus="holotype">
<locationDeviation id="9A1D0305FFE9443AFF57FB330AD481D4" box="[189,495,1224,1250]" pageId="6" pageNumber="127">SW ATLANTIC OCEAN</locationDeviation>
• 
<specimenCount id="9D16D1AEFFE9443AFDE6FB3309D781D4" box="[524,748,1223,1250]" pageId="6" pageNumber="127">one complete spec</specimenCount>
. (preserved in 96% ethanol); 
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, “Talud Continental III” exped., stn N° 45; 
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, 
<geoCoordinate id="EE247CE0FFE9443AFC4DFB120F708033" box="[935,1099,1257,1285]" degrees="53" direction="west" minutes="39.268" orientation="longitude" pageId="6" pageNumber="127" precision="1" value="-53.65447">53°39.268′ W</geoCoordinate>
; 
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deep; 
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<collectingDate id="EFEAC50FFFE9443AFB10FB100E498033" box="[1274,1394,1259,1285]" pageId="6" pageNumber="127" value="2013-09">Sep. 2013</collectingDate>
</date>
; 
<collectorName id="26E57FF1FFE9443AFF57FAF50A4E801E" box="[189,373,1294,1320]" pageId="6" pageNumber="127">Daniel Lauretta</collectorName>
leg.; 
<specimenCode id="DBB6B25CFFE9443AFE5CFAF509B0801E" box="[438,651,1294,1320]" collectionCode="MACN-IN" pageId="6" pageNumber="127">MACN-IN 42608</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BAF1A27FFE9443AFF00FAB4090180C0" blockId="6.[189,1398,1359,1527]" pageId="6" pageNumber="127">
<materialsCitation id="3B78107AFFE9443AFF00FAB4090D80C0" ID-GBIF-Occurrence="2884010302" collectingDate="2013-09" collectionCode="MACN-IN" collectorName="Daniel Lauretta" elevation="3282" latitude="-38.0885" location="Mar del Plata Submarine Canyon" longLatPrecision="1" longitude="-53.666466" pageId="6" pageNumber="127" specimenCode="MACN-IN 42609" specimenCount="3" typeStatus="paratype">
<emphasis id="B964C635FFE9443AFF00FAB40A5C805F" bold="true" box="[234,359,1359,1385]" pageId="6" pageNumber="127">
<typeStatus id="54ABA485FFE9443AFF00FAB40A5C805F" box="[234,359,1359,1385]" pageId="6" pageNumber="127" type="paratype">Paratypes</typeStatus>
</emphasis>
<locationDeviation id="9A1D0305FFE9443AFF57FA880AC880BB" box="[189,499,1395,1421]" pageId="6" pageNumber="127">SW ATLANTIC OCEAN</locationDeviation>
• 
<specimenCount id="9D16D1AEFFE9443AFDFFFA88094B80BB" box="[533,624,1395,1421]" pageId="6" pageNumber="127">3 specs</specimenCount>
(two adult, without the peduncle: paratypes A and B; and one juvenile like, complete: paratype C; preserved in 96% ethanol); 
<location id="8ECF4CFCFFE9443AFC3CFA6D0E498086" LSID="urn:lsid:plazi:treatment:03B9AB31FFE94437FE21FD7A0FBA8731:8ECF4CFCFFE9443AFC3CFA6D0E498086" box="[982,1394,1430,1456]" latitude="-38.0885" longLatPrecision="1" longitude="-53.666466" name="Mar del Plata Submarine Canyon" pageId="6" pageNumber="127">Mar del Plata Submarine Canyon</location>
, “Talud Continental III” exped., stn N° 46; 
<geoCoordinate id="EE247CE0FFE9443AFD58FA4C080680E5" box="[690,829,1463,1491]" degrees="38" direction="south" minutes="5.310" orientation="latitude" pageId="6" pageNumber="127" precision="1" value="-38.0885">38°5.310′ S</geoCoordinate>
, 
<geoCoordinate id="EE247CE0FFE9443AFCA0FA4C08D480E5" box="[842,1007,1463,1491]" degrees="53" direction="west" minutes="39.988" orientation="longitude" pageId="6" pageNumber="127" precision="1" value="-53.666466">53°39.988′ W</geoCoordinate>
; 
<quantity id="4CE8B7C2FFE9443AFC11FA420F6E80E5" box="[1019,1109,1465,1491]" metricMagnitude="3" metricUnit="m" metricValue="3.282" pageId="6" pageNumber="127" unit="m" value="3282.0">
<elevation id="003DFD14FFE9443AFC11FA420F6E80E5" box="[1019,1109,1465,1491]" metricMagnitude="3" metricUnit="m" metricValue="3.282" pageId="6" pageNumber="127" unit="m" value="3282.0">3282 m</elevation>
</quantity>
deep; 
<date id="FFAE3CE7FFE9443AFB49FA420E2780E5" box="[1187,1308,1465,1491]" pageId="6" pageNumber="127" value="2013-09">
<collectingDate id="EFEAC50FFFE9443AFB49FA420E2780E5" box="[1187,1308,1465,1491]" pageId="6" pageNumber="127" value="2013-09">Sep. 2013</collectingDate>
</date>
; 
<collectorName id="26E57FF1FFE9443AFAC2FA420A2480C1" pageId="6" pageNumber="127">Daniel Lauretta</collectorName>
leg.; 
<specimenCode id="DBB6B25CFFE9443AFE8BFA27090D80C0" box="[353,566,1500,1526]" collectionCode="MACN-IN" pageId="6" pageNumber="127">MACN-IN 42609</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A49ACFFE94437FF57F9E60FBA8731" lastPageId="11" lastPageNumber="132" pageId="6" pageNumber="127" type="description">
<paragraph id="8BAF1A27FFE9443AFF57F9E60993830E" blockId="6.[189,680,1565,1592]" box="[189,680,1565,1592]" pageId="6" pageNumber="127">
<emphasis id="B964C635FFE9443AFF57F9E60A758301" bold="true" box="[189,334,1565,1591]" pageId="6" pageNumber="127">Description</emphasis>
(
<typeStatus id="54ABA485FFE9443AFEB7F9E60AFD8301" box="[349,454,1565,1591]" pageId="6" pageNumber="127" type="holotype">holotype</typeStatus>
MACN-IN 42608)
</paragraph>
<paragraph id="8BAF1A27FFE94434FF57F9B70AF084BB" blockId="6.[189,1399,1612,1991]" lastBlockId="8.[187,1397,265,397]" lastPageId="8" lastPageNumber="129" pageId="6" pageNumber="127">
The colony looks rugous in all its extension, especially the terminal cluster of autozooids. The color of the polyps, rachis and the peduncle is white or light yellow (preserved). The 
<typeStatus id="54ABA485FFE9443AFB81F9940FE983BF" box="[1131,1234,1647,1673]" pageId="6" pageNumber="127" type="holotype">holotype</typeStatus>
is 
<quantity id="4CE8B7C2FFE9443AFB1CF9940E6383BC" box="[1270,1368,1647,1674]" metricMagnitude="-1" metricUnit="m" metricValue="2.1399999999999997" pageId="6" pageNumber="127" unit="mm" value="214.0">214 mm</quantity>
in length. It has a terminal cluster of three autozooids with tentacles, one central and larger polyp growing on the distal-most region of the rachis, and two smaller but well-developed lateral polyps growing at the base of the central one (
<figureCitation id="132B06A2FFE9443AFDECF922097483C5" box="[518,591,1753,1779]" captionStart="Fig" captionStartId="7.[189,232,907,933]" captionTargetBox="[195,1392,271,864]" captionTargetId="figure-12@7.[189,1398,265,870]" captionTargetPageId="7" captionText="Fig. 2. A. General aspect of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. A. Holotype (MACN-IN 42608). B. Detail of the terminal cluster of paratype A (MACN-IN 42609), showing three autozooids that form the terminal cluster and amplifications of sclerites (up) and siphonozooids (down). Abbreviations: CP = central polyp; LP = lateral polyp; Pd = peduncle; R = rachis; T = tentacle; S = siphonozooids; Scl = sclerites." figureDoi="http://doi.org/10.5281/zenodo.4106767" httpUri="https://zenodo.org/record/4106767/files/figure.png" pageId="6" pageNumber="127">Fig. 2</figureCitation>
). The central polyp is 
<quantity id="4CE8B7C2FFE9443AFCB0F92208FF83C5" box="[858,964,1753,1780]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="6" pageNumber="127" unit="mm" value="25.3">25.3 mm</quantity>
in length and the two lateral polyps’ lengths are 3.0 mm and 
<quantity id="4CE8B7C2FFE9443AFE34F90709068221" box="[478,573,1788,1815]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="127" unit="mm" value="2.6">2.6 mm</quantity>
. The tentacles of the two types of polyps are larger than its body, the measures are 
<quantity id="4CE8B7C2FFE9443AFEB4F8E40AFC820C" box="[350,455,1823,1850]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="6" pageNumber="127" unit="mm" value="12.3">12.3 mm</quantity>
for the central autozooid, and 
<quantity id="4CE8B7C2FFE9443AFCDBF8E408B0820C" box="[817,907,1823,1850]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="127" unit="mm" value="1.1">1.1 mm</quantity>
and 
<quantity id="4CE8B7C2FFE9443AFC2FF8E40F24820C" box="[965,1055,1823,1850]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="127" unit="mm" value="1.7">1.7 mm</quantity>
for the lateral ones. The axis is 
<quantity id="4CE8B7C2FFE9443AFF33F8B80A00826B" box="[217,315,1859,1885]" metricMagnitude="-1" metricUnit="m" metricValue="1.78" pageId="6" pageNumber="127" unit="mm" value="178.0">178 mm</quantity>
in length and it is circular in cross-section, approximately 
<quantity id="4CE8B7C2FFE9443AFC1CF8B80F5B826B" box="[1014,1120,1859,1885]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="6" pageNumber="127" unit="mm" value="0.74">0.74 mm</quantity>
in diameter. The rachis is approximately 
<quantity id="4CE8B7C2FFE9443AFE63F89D0AD882B6" box="[393,483,1894,1920]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="127" unit="mm" value="0.8">0.8 mm</quantity>
in diameter in the middle zone between the peduncle and the terminal cluster. The peduncle is 
<quantity id="4CE8B7C2FFE9443AFE97F8720ADE8295" box="[381,485,1929,1955]" metricMagnitude="-3" metricUnit="m" metricValue="8.68" pageId="6" pageNumber="127" unit="mm" value="8.68">8.68 mm</quantity>
in length, and its appearance is soft and smooth. The autozooids of the cluster grow with a kind of orientation determining a ‘dorsal’ region where the axis inserts and a ‘ventral’ region towards which the polyps come together (
<figureCitation id="132B06A2FFE74434FD1EFEF208008415" box="[756,827,265,291]" captionStart="Fig" captionStartId="7.[189,232,907,933]" captionTargetBox="[195,1392,271,864]" captionTargetId="figure-12@7.[189,1398,265,870]" captionTargetPageId="7" captionText="Fig. 2. A. General aspect of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. A. Holotype (MACN-IN 42608). B. Detail of the terminal cluster of paratype A (MACN-IN 42609), showing three autozooids that form the terminal cluster and amplifications of sclerites (up) and siphonozooids (down). Abbreviations: CP = central polyp; LP = lateral polyp; Pd = peduncle; R = rachis; T = tentacle; S = siphonozooids; Scl = sclerites." figureDoi="http://doi.org/10.5281/zenodo.4106767" httpUri="https://zenodo.org/record/4106767/files/figure.png" pageId="8" pageNumber="129">Fig. 2</figureCitation>
). The siphonozooids are numerous and resemble small white dots, and are distributed all along the rachis from the base of the autozooids to the middle of the rachis. These polyps are circular and inconspicuous; and have a diameter of about 393 ± 93 μm (299–486 μm, N = 36).
</paragraph>
<caption id="DF6F4AAFFFE8443BFF57FC7009188104" ID-DOI="http://doi.org/10.5281/zenodo.4106767" ID-Zenodo-Dep="4106767" httpUri="https://zenodo.org/record/4106767/files/figure.png" pageId="7" pageNumber="128" startId="7.[189,232,907,933]" targetBox="[195,1392,271,864]" targetPageId="7">
<paragraph id="8BAF1A27FFE8443BFF57FC7009188104" blockId="7.[189,1399,907,1075]" pageId="7" pageNumber="128">
<emphasis id="B964C635FFE8443BFF57FC700A348693" bold="true" box="[189,271,907,933]" pageId="7" pageNumber="128">Fig. 2.</emphasis>
A. General aspect of 
<taxonomicName id="4C1061A4FFE8443BFDC8FC700F578690" authority="Risaro, Williams &amp; Lauretta" authorityName="Risaro, Williams &amp; Lauretta" authorityYear="2020" box="[546,1132,907,934]" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="7" pageNumber="128" phylum="Cnidaria" rank="species" species="pomona" status="sp. nov.">
<emphasis id="B964C635FFE8443BFDC8FC7008338693" box="[546,776,907,933]" italics="true" pageId="7" pageNumber="128">Umbellula pomona</emphasis>
Risaro, Williams &amp; Lauretta
</taxonomicName>
<taxonomicNameLabel id="A2577B4EFFE8443BFB9DFC770FED8690" box="[1143,1238,908,934]" pageId="7" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B964C635FFE8443BFB0BFC700FCD8693" bold="true" box="[1249,1270,907,933]" pageId="7" pageNumber="128">A</emphasis>
. Holotype (MACN-IN 42608). 
<emphasis id="B964C635FFE8443BFE58FC540AFD86FF" bold="true" box="[434,454,943,969]" pageId="7" pageNumber="128">B</emphasis>
. Detail of the terminal cluster of paratype A (MACN-IN 42609), showing three autozooids that form the terminal cluster and amplifications of sclerites (up) and siphonozooids (down). Abbreviations: CP = central polyp; LP = lateral polyp; Pd = peduncle; R = rachis; T = tentacle; S = siphonozooids; Scl = sclerites.
</paragraph>
</caption>
<caption id="DF6F4AAFFFE8443BFF57F8850F4E82E9" ID-DOI="http://doi.org/10.5281/zenodo.4106769" ID-Zenodo-Dep="4106769" httpUri="https://zenodo.org/record/4106769/files/figure.png" pageId="7" pageNumber="128" startId="7.[189,232,1918,1944]" targetBox="[239,1349,1135,1879]" targetPageId="7">
<paragraph id="8BAF1A27FFE8443BFF57F8850F4E82E9" blockId="7.[189,1399,1918,2016]" pageId="7" pageNumber="128">
<emphasis id="B964C635FFE8443BFF57F8850A3082AE" bold="true" box="[189,267,1918,1944]" pageId="7" pageNumber="128">Fig. 3.</emphasis>
Transversal section of the central axis of paratype B of 
<taxonomicName id="4C1061A4FFE8443BFC4CF8840A24828A" authority="Risaro, Williams &amp; Lauretta" authorityName="Risaro, Williams &amp; Lauretta" authorityYear="2020" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="7" pageNumber="128" phylum="Cnidaria" rank="species" species="pomona" status="sp. nov.">
<emphasis id="B964C635FFE8443BFC4CF8840FB282AE" box="[934,1161,1919,1945]" italics="true" pageId="7" pageNumber="128">Umbellula pomona</emphasis>
Risaro, Williams &amp; Lauretta
</taxonomicName>
<taxonomicNameLabel id="A2577B4EFFE8443BFEC2F8590ABF828A" box="[296,388,1954,1980]" pageId="7" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
(MACN-IN 42609). 
<emphasis id="B964C635FFE8443BFD6EF85909A2828A" bold="true" box="[644,665,1954,1980]" pageId="7" pageNumber="128">A</emphasis>
. Transversal section of the central axis between the peduncle and the rachis. 
<emphasis id="B964C635FFE8443BFE85F83E0AB882E9" bold="true" box="[367,387,1989,2015]" pageId="7" pageNumber="128">B</emphasis>
. Transversal section of the central axis near the terminal cluster.
</paragraph>
</caption>
<paragraph id="8BAF1A27FFE74435FF51FE4F0ABD84C0" blockId="8.[187,1397,436,744]" lastBlockId="9.[189,1399,265,503]" lastPageId="9" lastPageNumber="130" pageId="8" pageNumber="129">
Sclerites are conspicuous, along much of the length of the colony (polyps, rachis, tentacles and pinnules). They are translucent and colorless, rod-shaped and spindle-shaped with spines (
<figureCitation id="132B06A2FFE74434FBBFFE2C0FFB84C7" box="[1109,1216,471,497]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="8.[189,232,1952,1978]" captionStartId-1="9.[189,232,1958,1984]" captionStartId-2="10.[189,232,1958,1984]" captionStartId-3="11.[189,232,1958,1984]" captionTargetBox-0="[189,1392,830,1909]" captionTargetBox-1="[195,1391,842,1912]" captionTargetBox-2="[195,1393,840,1913]" captionTargetBox-3="[200,1392,834,1913]" captionTargetId-0="figure-235@8.[189,1398,824,1909]" captionTargetId-1="figure-224@9.[189,1398,834,1919]" captionTargetId-2="figure-271@10.[189,1398,834,1919]" captionTargetId-3="figure-163@11.[189,1398,828,1913]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="Fig. 4. Variability of sizes and ornamentations of the rachis’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." captionText-1="Fig. 5. Variability of sizes and ornamentations of the polyps’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." captionText-2="Fig. 6. Variability of sizes and ornamentations of the tentacles’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." captionText-3="Fig. 7. Variability of sizes and ornamentations of the pinnules’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." figureDoi-0="http://doi.org/10.5281/zenodo.4106771" figureDoi-1="http://doi.org/10.5281/zenodo.4106773" figureDoi-2="http://doi.org/10.5281/zenodo.4106775" figureDoi-3="http://doi.org/10.5281/zenodo.4106777" httpUri-0="https://zenodo.org/record/4106771/files/figure.png" httpUri-1="https://zenodo.org/record/4106773/files/figure.png" httpUri-2="https://zenodo.org/record/4106775/files/figure.png" httpUri-3="https://zenodo.org/record/4106777/files/figure.png" pageId="8" pageNumber="129">Figs 4–7</figureCitation>
). Those present in the rachis are spindle-shaped, have triangular protuberances all along their length (
<figureCitation id="132B06A2FFE74434FB59FE010FC08723" box="[1203,1275,506,533]" captionStart="Fig" captionStartId="8.[189,232,1952,1978]" captionTargetBox="[189,1392,830,1909]" captionTargetId="figure-235@8.[189,1398,824,1909]" captionTargetPageId="8" captionText="Fig. 4. Variability of sizes and ornamentations of the rachis’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." figureDoi="http://doi.org/10.5281/zenodo.4106771" httpUri="https://zenodo.org/record/4106771/files/figure.png" pageId="8" pageNumber="129">Fig. 4</figureCitation>
) and their sizes are 333.5 ± 71.1 μm (262.4–404.6 μm, N = 10), the body wall of the polyps has different 
<typeStatus id="54ABA485FFE74434FAFDFDE50E6E870E" box="[1303,1365,542,568]" pageId="8" pageNumber="129">types</typeStatus>
of sclerites, two of them are rough and rod-shaped, but differ in their measurements, while the others are spindle-shaped with spines along their lengths and also have different sizes (
<figureCitation id="132B06A2FFE74434FBD5FD9F0FBC8749" box="[1087,1159,612,639]" captionStart="Fig" captionStartId="9.[189,232,1958,1984]" captionTargetBox="[195,1391,842,1912]" captionTargetId="figure-224@9.[189,1398,834,1919]" captionTargetPageId="9" captionText="Fig. 5. Variability of sizes and ornamentations of the polyps’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." figureDoi="http://doi.org/10.5281/zenodo.4106773" httpUri="https://zenodo.org/record/4106773/files/figure.png" pageId="8" pageNumber="129">Fig. 5</figureCitation>
). The largest 
<typeStatus id="54ABA485FFE74434FACDFD9E0E618749" box="[1319,1370,613,639]" pageId="8" pageNumber="129">type</typeStatus>
is rod-shaped and approximates 1446 ± 32.7 μm (1413.3–1478.7 μm, N = 5) in length (
<figureCitation id="132B06A2FFE74434FB42FD7C0E3C8794" box="[1192,1287,647,674]" captionStart="Fig" captionStartId="9.[189,232,1958,1984]" captionTargetBox="[195,1391,842,1912]" captionTargetId="figure-224@9.[189,1398,834,1919]" captionTargetPageId="9" captionText="Fig. 5. Variability of sizes and ornamentations of the polyps’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." figureDoi="http://doi.org/10.5281/zenodo.4106773" httpUri="https://zenodo.org/record/4106773/files/figure.png" pageId="8" pageNumber="129">Fig. 5A</figureCitation>
); another 
<typeStatus id="54ABA485FFE74434FF51FD500BCB87F3" box="[187,240,683,709]" pageId="8" pageNumber="129">type</typeStatus>
, smaller and spine-shaped, approximates 687 ± 45.6 μm (641.4–732.6 μm, N = 5) (
<figureCitation id="132B06A2FFE74434FB5CFD510FC287F3" box="[1206,1273,682,709]" captionStart="Fig" captionStartId="9.[189,232,1958,1984]" captionTargetBox="[195,1391,842,1912]" captionTargetId="figure-224@9.[189,1398,834,1919]" captionTargetPageId="9" captionText="Fig. 5. Variability of sizes and ornamentations of the polyps’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." figureDoi="http://doi.org/10.5281/zenodo.4106773" httpUri="https://zenodo.org/record/4106773/files/figure.png" pageId="8" pageNumber="129">Fig. 5</figureCitation>
B–C). The other 
<typeStatus id="54ABA485FFE74434FEEAFD350A0587DE" box="[256,318,718,744]" pageId="8" pageNumber="129">types</typeStatus>
are much smaller, and one 
<typeStatus id="54ABA485FFE74434FD62FD35098087DE" box="[648,699,718,744]" pageId="8" pageNumber="129">type</typeStatus>
is rod shaped, like the largest one (
<figureCitation id="132B06A2FFE74434FB89FD350FF987DE" box="[1123,1218,717,744]" captionStart="Fig" captionStartId="9.[189,232,1958,1984]" captionTargetBox="[195,1391,842,1912]" captionTargetId="figure-224@9.[189,1398,834,1919]" captionTargetPageId="9" captionText="Fig. 5. Variability of sizes and ornamentations of the polyps’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." figureDoi="http://doi.org/10.5281/zenodo.4106773" httpUri="https://zenodo.org/record/4106773/files/figure.png" pageId="8" pageNumber="129">Fig. 5D</figureCitation>
), and the other 
<typeStatus id="54ABA485FFE64435FF57FEF20BCB8415" box="[189,240,265,291]" pageId="9" pageNumber="130">type</typeStatus>
is spine-shaped (
<figureCitation id="132B06A2FFE64435FE2BFEF2093C8412" box="[449,519,265,292]" captionStart="Fig" captionStartId="9.[189,232,1958,1984]" captionTargetBox="[195,1391,842,1912]" captionTargetId="figure-224@9.[189,1398,834,1919]" captionTargetPageId="9" captionText="Fig. 5. Variability of sizes and ornamentations of the polyps’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." figureDoi="http://doi.org/10.5281/zenodo.4106773" httpUri="https://zenodo.org/record/4106773/files/figure.png" pageId="9" pageNumber="130">Fig. 5</figureCitation>
E–F); these two 
<typeStatus id="54ABA485FFE64435FD25FEF208368415" box="[719,781,265,291]" pageId="9" pageNumber="130">types</typeStatus>
of sclerites have similar sizes, length approximates 534 ± 43.4 μm (490.6–577.4 μm, N = 5). Finally, the tentacles have rod-shaped sclerites (
<figureCitation id="132B06A2FFE64435FB03FED70E0B8471" box="[1257,1328,300,327]" captionStart="Fig" captionStartId="10.[189,232,1958,1984]" captionTargetBox="[195,1393,840,1913]" captionTargetId="figure-271@10.[189,1398,834,1919]" captionTargetPageId="10" captionText="Fig. 6. Variability of sizes and ornamentations of the tentacles’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." figureDoi="http://doi.org/10.5281/zenodo.4106775" httpUri="https://zenodo.org/record/4106775/files/figure.png" pageId="9" pageNumber="130">Fig. 6</figureCitation>
) with two sizes: the larger ones are 1223.2 ± 280.1 μm (943.1–1503.3 μm, N = 5) (
<figureCitation id="132B06A2FFE64435FBD3FEB40F46845C" box="[1081,1149,335,362]" captionStart="Fig" captionStartId="10.[189,232,1958,1984]" captionTargetBox="[195,1393,840,1913]" captionTargetId="figure-271@10.[189,1398,834,1919]" captionTargetPageId="10" captionText="Fig. 6. Variability of sizes and ornamentations of the tentacles’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." figureDoi="http://doi.org/10.5281/zenodo.4106775" httpUri="https://zenodo.org/record/4106775/files/figure.png" pageId="9" pageNumber="130">Fig. 6</figureCitation>
A–C) and the smaller ones are 632.8 ± 96.2 μm (536.6–729 μm, N = 5) in length (
<figureCitation id="132B06A2FFE64435FC40FE8908C884BB" box="[938,1011,370,397]" captionStart="Fig" captionStartId="10.[189,232,1958,1984]" captionTargetBox="[195,1393,840,1913]" captionTargetId="figure-271@10.[189,1398,834,1919]" captionTargetPageId="10" captionText="Fig. 6. Variability of sizes and ornamentations of the tentacles’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." figureDoi="http://doi.org/10.5281/zenodo.4106775" httpUri="https://zenodo.org/record/4106775/files/figure.png" pageId="9" pageNumber="130">Fig. 6</figureCitation>
D–F), and those of the pinnules are rod-shaped with some protuberances along their lengths and their sizes are approximately 269.1 ± 50.3 μm (218.8–319.4 μm, N = 10) (
<figureCitation id="132B06A2FFE64435FD80FE42098984E5" box="[618,690,441,467]" captionStart="Fig" captionStartId="11.[189,232,1958,1984]" captionTargetBox="[200,1392,834,1913]" captionTargetId="figure-163@11.[189,1398,828,1913]" captionTargetPageId="11" captionText="Fig. 7. Variability of sizes and ornamentations of the pinnules’ sclerites of the holotype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42608)." figureDoi="http://doi.org/10.5281/zenodo.4106777" httpUri="https://zenodo.org/record/4106777/files/figure.png" pageId="9" pageNumber="130">Fig. 7</figureCitation>
). In both tissues the sclerites are placed along the main axis, not transversally.
</paragraph>
<caption id="DF6F4AAFFFE74434FF57F85B08FE82E9" ID-DOI="http://doi.org/10.5281/zenodo.4106771" ID-Zenodo-Dep="4106771" httpUri="https://zenodo.org/record/4106771/files/figure.png" pageId="8" pageNumber="129" startId="8.[189,232,1952,1978]" targetBox="[189,1392,830,1909]" targetPageId="8">
<paragraph id="8BAF1A27FFE74434FF57F85B08FE82E9" blockId="8.[189,1399,1952,2015]" pageId="8" pageNumber="129">
<emphasis id="B964C635FFE74434FF57F85B0A35828C" bold="true" box="[189,270,1952,1978]" pageId="8" pageNumber="129">Fig. 4.</emphasis>
Variability of sizes and ornamentations of the rachis’ sclerites of the holotype of 
<taxonomicName id="4C1061A4FFE74434FB11F85B095482E8" authority="Risaro, Williams &amp; Lauretta" authorityName="Risaro, Williams &amp; Lauretta" authorityYear="2020" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="8" pageNumber="129" phylum="Cnidaria" rank="species" species="pomona" status="sp. nov.">
<emphasis id="B964C635FFE74434FB11F85B0A2782E8" italics="true" pageId="8" pageNumber="129">Umbellula pomona</emphasis>
Risaro, Williams &amp; Lauretta
</taxonomicName>
<taxonomicNameLabel id="A2577B4EFFE74434FD9CF83F09EA82E8" box="[630,721,1988,2014]" pageId="8" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
(MACN-IN 42608).
</paragraph>
</caption>
<paragraph id="8BAF1A27FFE64435FF57FDE60A788701" blockId="9.[189,323,541,567]" box="[189,323,541,567]" pageId="9" pageNumber="130">
<heading id="D0E7AD4BFFE64435FF57FDE60A788701" bold="true" box="[189,323,541,567]" fontSize="11" level="3" pageId="9" pageNumber="130" reason="3">
<emphasis id="B964C635FFE64435FF57FDE60A788701" bold="true" box="[189,323,541,567]" pageId="9" pageNumber="130">Variability</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF1A27FFE64436FF57FDB7096E87DB" blockId="9.[189,1399,588,755]" lastBlockId="10.[189,1399,265,749]" lastPageId="10" lastPageNumber="131" pageId="9" pageNumber="130">
Of the three 
<typeStatus id="54ABA485FFE64435FEB2FDB60AF38751" box="[344,456,589,615]" pageId="9" pageNumber="130" type="paratype">paratypes</typeStatus>
(A, B and C), one (
<typeStatus id="54ABA485FFE64435FD54FDB7081E8750" box="[702,805,588,614]" pageId="9" pageNumber="130" type="paratype">paratype</typeStatus>
C) is significantly smaller than the others, so we consider it could be a juvenile (
<figureCitation id="132B06A2FFE64435FDD0FD9409B887BF" box="[570,643,623,649]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[195,1392,911,1873]" captionTargetId="figure-286@12.[189,1398,906,1879]" captionTargetPageId="12" captionText="Fig. 8. General aspect of the unique specimen of a juvenile-like paratype of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42609, paratype C). Abbreviations: CP = central polyp; LP = lateral polyp; R = rachis; PD = peduncle; T = tentacles." figureDoi="http://doi.org/10.5281/zenodo.4106779" httpUri="https://zenodo.org/record/4106779/files/figure.png" pageId="9" pageNumber="130">Fig. 8</figureCitation>
). The color of the three colonies is white or light yellow when preserved, all of them have three autozooids with the same grade of development as the 
<typeStatus id="54ABA485FFE64435FB0EFD690E70879A" box="[1252,1355,658,684]" pageId="9" pageNumber="130" type="holotype">holotype</typeStatus>
(as it can be seen in 
<figureCitation id="132B06A2FFE64435FE6DFD4D0ADA87E6" box="[391,481,694,720]" captionStart="Fig" captionStartId="7.[189,232,907,933]" captionTargetBox="[195,1392,271,864]" captionTargetId="figure-12@7.[189,1398,265,870]" captionTargetPageId="7" captionText="Fig. 2. A. General aspect of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. A. Holotype (MACN-IN 42608). B. Detail of the terminal cluster of paratype A (MACN-IN 42609), showing three autozooids that form the terminal cluster and amplifications of sclerites (up) and siphonozooids (down). Abbreviations: CP = central polyp; LP = lateral polyp; Pd = peduncle; R = rachis; T = tentacle; S = siphonozooids; Scl = sclerites." figureDoi="http://doi.org/10.5281/zenodo.4106767" httpUri="https://zenodo.org/record/4106767/files/figure.png" pageId="9" pageNumber="130">Fig. 2B</figureCitation>
) and its central axis is round in all its extension (
<figureCitation id="132B06A2FFE64435FBC4FD4D0F4D87E6" box="[1070,1142,694,720]" captionStart="Fig" captionStartId="7.[189,232,1918,1944]" captionTargetBox="[239,1349,1135,1879]" captionTargetId="figure-85@7.[234,1354,1130,1884]" captionTargetPageId="7" captionText="Fig. 3. Transversal section of the central axis of paratype B of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. (MACN-IN 42609). A. Transversal section of the central axis between the peduncle and the rachis. B. Transversal section of the central axis near the terminal cluster." figureDoi="http://doi.org/10.5281/zenodo.4106769" httpUri="https://zenodo.org/record/4106769/files/figure.png" pageId="9" pageNumber="130">Fig. 3</figureCitation>
). The large 
<typeStatus id="54ABA485FFE64435FAECFD4D0E4D87E6" box="[1286,1398,694,720]" pageId="9" pageNumber="130" type="paratype">paratypes</typeStatus>
(A and B) lack their peduncle. Their general aspect is just like the 
<typeStatus id="54ABA485FFE64435FC20FD220F0F87C5" box="[970,1076,729,755]" pageId="9" pageNumber="130" type="holotype">holotype</typeStatus>
, they are rugous in all their length because of the presence of conspicuous sclerites in the autozooids and rachis (
<figureCitation id="132B06A2FFE54436FB77FEF20FC28415" box="[1181,1273,265,291]" captionStart="Fig" captionStartId="7.[189,232,907,933]" captionTargetBox="[195,1392,271,864]" captionTargetId="figure-12@7.[189,1398,265,870]" captionTargetPageId="7" captionText="Fig. 2. A. General aspect of Umbellula pomona Risaro, Williams &amp; Lauretta sp. nov. A. Holotype (MACN-IN 42608). B. Detail of the terminal cluster of paratype A (MACN-IN 42609), showing three autozooids that form the terminal cluster and amplifications of sclerites (up) and siphonozooids (down). Abbreviations: CP = central polyp; LP = lateral polyp; Pd = peduncle; R = rachis; T = tentacle; S = siphonozooids; Scl = sclerites." figureDoi="http://doi.org/10.5281/zenodo.4106767" httpUri="https://zenodo.org/record/4106767/files/figure.png" pageId="10" pageNumber="131">Fig. 2B</figureCitation>
). The total length of the large 
<typeStatus id="54ABA485FFE54436FE4AFED6092B8471" box="[416,528,301,327]" pageId="10" pageNumber="131" type="paratype">paratypes</typeStatus>
is 
<quantity id="4CE8B7C2FFE54436FDDCFED709A28471" box="[566,665,300,327]" metricMagnitude="-1" metricUnit="m" metricValue="1.34" pageId="10" pageNumber="131" unit="mm" value="134.0">134 mm</quantity>
and 
<quantity id="4CE8B7C2FFE54436FD3EFED708018471" box="[724,826,300,327]" metricMagnitude="-1" metricUnit="m" metricValue="2.31" pageId="10" pageNumber="131" unit="mm" value="231.0">231 mm</quantity>
, their central autozooids are 
<quantity id="4CE8B7C2FFE54436FB79FED70FDC8471" box="[1171,1255,300,327]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="10" pageNumber="131" unit="mm" value="36.0">36 mm</quantity>
and 
<quantity id="4CE8B7C2FFE54436FAC9FED70E4C8471" box="[1315,1399,300,327]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="10" pageNumber="131" unit="mm" value="34.0">34 mm</quantity>
long (length of the tentacles: 
<quantity id="4CE8B7C2FFE54436FDE7FEB40965845C" box="[525,606,335,362]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="131" unit="mm" value="20.0">20 mm</quantity>
and 
<quantity id="4CE8B7C2FFE54436FD79FEB409D3845C" box="[659,744,335,362]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="131" unit="mm" value="12.0">12 mm</quantity>
, length of the columns: 
<quantity id="4CE8B7C2FFE54436FC10FEB40F70845C" box="[1018,1099,335,362]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="131" unit="mm" value="16.0">16 mm</quantity>
and 
<quantity id="4CE8B7C2FFE54436FB6BFEB40FEE845C" box="[1153,1237,335,362]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="131" unit="mm" value="22.0">22 mm</quantity>
, respectively) while the dimensions of the lateral ones are 
<quantity id="4CE8B7C2FFE54436FD2AFE88082184BB" box="[704,794,371,397]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="10" pageNumber="131" unit="mm" value="4.8">4.8 mm</quantity>
and 
<quantity id="4CE8B7C2FFE54436FCB9FE89089684BB" box="[851,941,370,397]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="10" pageNumber="131" unit="mm" value="5.6">5.6 mm</quantity>
in one 
<typeStatus id="54ABA485FFE54436FBE9FE880F5384BB" box="[1027,1128,371,397]" pageId="10" pageNumber="131" type="paratype">paratype</typeStatus>
and 
<quantity id="4CE8B7C2FFE54436FB4BFE890FC084BB" box="[1185,1275,370,397]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="10" pageNumber="131" unit="mm" value="6.4">6.4 mm</quantity>
and 
<quantity id="4CE8B7C2FFE54436FAD9FE880E4C84BB" box="[1331,1399,371,397]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="131" unit="mm" value="7.0">7 mm</quantity>
in the other. Their rachis are 
<quantity id="4CE8B7C2FFE54436FDFAFE6D09588486" box="[528,611,406,432]" metricMagnitude="-2" metricUnit="m" metricValue="9.7" pageId="10" pageNumber="131" unit="mm" value="97.0">97 mm</quantity>
and 
<quantity id="4CE8B7C2FFE54436FD76FE6D09C58486" box="[668,766,406,432]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="131" unit="mm" value="200.0">200 mm</quantity>
in length and their diameters are 
<quantity id="4CE8B7C2FFE54436FB63FE6D0FD88486" box="[1161,1251,406,432]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="131" unit="mm" value="0.9">0.9 mm</quantity>
and 
<quantity id="4CE8B7C2FFE54436FAF7FE6D0E4C8486" box="[1309,1399,406,432]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="131" unit="mm" value="2.2">2.2 mm</quantity>
at thier widest sections. Their axis’ diameter is 
<quantity id="4CE8B7C2FFE54436FD00FE42086F84E5" box="[746,852,441,467]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="10" pageNumber="131" unit="mm" value="0.83">0.83 mm</quantity>
and 
<quantity id="4CE8B7C2FFE54436FC64FE4208D084E5" box="[910,1003,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="131" unit="mm" value="1.3">1.3 mm</quantity>
. Their siphonozooids are 313 μm and 370 μm (mean) in diameter and look like the siphonozooids of the 
<typeStatus id="54ABA485FFE54436FC12FE270F5884C0" box="[1016,1123,476,502]" pageId="10" pageNumber="131" type="holotype">holotype</typeStatus>
. Finally, the sclerites of these 
<typeStatus id="54ABA485FFE54436FEEAFDFB0A4B872C" box="[256,368,512,538]" pageId="10" pageNumber="131" type="paratype">paratypes</typeStatus>
look alike and have similar sizes as the sclerites of the 
<typeStatus id="54ABA485FFE54436FC1BFE040F67872F" box="[1009,1116,511,537]" pageId="10" pageNumber="131" type="holotype">holotype</typeStatus>
. The juvenile paratype’s (C) total length is 
<quantity id="4CE8B7C2FFE54436FE7EFDD80AC3870B" box="[404,504,547,573]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="131" unit="mm" value="110.0">110 mm</quantity>
, its central autozooid is 
<quantity id="4CE8B7C2FFE54436FCFFFDD8084B870B" box="[789,880,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="10" pageNumber="131" unit="mm" value="7.9">7.9 mm</quantity>
in length (column and tentacles are 
<quantity id="4CE8B7C2FFE54436FAF6FDD80E4C870B" box="[1308,1399,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="10" pageNumber="131" unit="mm" value="4.4">4.4 mm</quantity>
and 
<quantity id="4CE8B7C2FFE54436FF18FDBD0A6B8756" box="[242,336,582,608]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="10" pageNumber="131" unit="mm" value="3.4">3.4 mm</quantity>
long, respectively), and the length of the lateral ones is 
<quantity id="4CE8B7C2FFE54436FBECFDBD0F5F8756" box="[1030,1124,582,608]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="131" unit="mm" value="2.9">2.9 mm</quantity>
(column and tentacles are 1.4 and 
<quantity id="4CE8B7C2FFE54436FEA6FD920A9C87B5" box="[332,423,617,644]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="131" unit="mm" value="1.5">1.5 mm</quantity>
in length, respectively). The peduncle’s length is 
<quantity id="4CE8B7C2FFE54436FC15FD920F6087B5" box="[1023,1115,617,643]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="10" pageNumber="131" unit="mm" value="4.6">4.6 mm</quantity>
and the rachis’ 
<quantity id="4CE8B7C2FFE54436FAF1FD920E4887B5" box="[1307,1395,617,644]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="10" pageNumber="131" unit="mm" value="95.0">95 mm</quantity>
, while its diameter is 
<quantity id="4CE8B7C2FFE54436FE54FD7709118791" box="[446,554,652,679]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="10" pageNumber="131" unit="mm" value="0.41">0.41 mm</quantity>
at its widest section. Its central axis is 
<quantity id="4CE8B7C2FFE54436FBFAFD770F478791" box="[1040,1148,652,679]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="131" unit="mm" value="0.35">0.35 mm</quantity>
in diameter. Finally, the siphonozooids of this 
<typeStatus id="54ABA485FFE54436FE05FD4B096F87FC" box="[495,596,688,714]" pageId="10" pageNumber="131" type="paratype">paratype</typeStatus>
are tiny spots with the same aspect and distribution as those on the 
<typeStatus id="54ABA485FFE54436FF57FD290A1F87DA" box="[189,292,722,748]" pageId="10" pageNumber="131" type="holotype">holotype</typeStatus>
and the largest 
<typeStatus id="54ABA485FFE54436FE37FD28096A87DB" box="[477,593,723,749]" pageId="10" pageNumber="131" type="paratype">paratypes</typeStatus>
.
</paragraph>
<caption id="DF6F4AAFFFE64435FF57F85D08FE82D2" ID-DOI="http://doi.org/10.5281/zenodo.4106773" ID-Zenodo-Dep="4106773" httpUri="https://zenodo.org/record/4106773/files/figure.png" pageId="9" pageNumber="130" startId="9.[189,232,1958,1984]" targetBox="[195,1391,842,1912]" targetPageId="9">
<paragraph id="8BAF1A27FFE64435FF57F85D08FE82D2" blockId="9.[189,1399,1958,2020]" pageId="9" pageNumber="130">
<emphasis id="B964C635FFE64435FF57F85D0A3582F6" bold="true" box="[189,270,1958,1984]" pageId="9" pageNumber="130">Fig. 5.</emphasis>
Variability of sizes and ornamentations of the polyps’ sclerites of the holotype of 
<taxonomicName id="4C1061A4FFE64435FB11F85D095482D2" authority="Risaro, Williams &amp; Lauretta" authorityName="Risaro, Williams &amp; Lauretta" authorityYear="2020" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="9" pageNumber="130" phylum="Cnidaria" rank="species" species="pomona" status="sp. nov.">
<emphasis id="B964C635FFE64435FB11F85D0A2782D5" italics="true" pageId="9" pageNumber="130">Umbellula pomona</emphasis>
Risaro, Williams &amp; Lauretta
</taxonomicName>
<taxonomicNameLabel id="A2577B4EFFE64435FD9CF83109EA82D2" box="[630,721,1994,2020]" pageId="9" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
(MACN-IN 42608).
</paragraph>
</caption>
<caption id="DF6F4AAFFFE54436FF57F85D08FE82D2" ID-DOI="http://doi.org/10.5281/zenodo.4106775" ID-Zenodo-Dep="4106775" httpUri="https://zenodo.org/record/4106775/files/figure.png" pageId="10" pageNumber="131" startId="10.[189,232,1958,1984]" targetBox="[195,1393,840,1913]" targetPageId="10">
<paragraph id="8BAF1A27FFE54436FF57F85D08FE82D2" blockId="10.[189,1399,1958,2020]" pageId="10" pageNumber="131">
<emphasis id="B964C635FFE54436FF57F85D0A3782F6" bold="true" box="[189,268,1958,1984]" pageId="10" pageNumber="131">Fig. 6.</emphasis>
Variability of sizes and ornamentations of the tentacles’ sclerites of the holotype of 
<taxonomicName id="4C1061A4FFE54436FB11F85D095482D2" authority="Risaro, Williams &amp; Lauretta" authorityName="Risaro, Williams &amp; Lauretta" authorityYear="2020" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="10" pageNumber="131" phylum="Cnidaria" rank="species" species="pomona" status="sp. nov.">
<emphasis id="B964C635FFE54436FB11F85D0A2782D5" italics="true" pageId="10" pageNumber="131">Umbellula pomona</emphasis>
Risaro, Williams &amp; Lauretta
</taxonomicName>
<taxonomicNameLabel id="A2577B4EFFE54436FD9CF83109EA82D2" box="[630,721,1994,2020]" pageId="10" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
(MACN-IN 42608).
</paragraph>
</caption>
<paragraph id="8BAF1A27FFE44437FF57FEF30AE08413" blockId="11.[189,475,264,293]" box="[189,475,264,293]" pageId="11" pageNumber="132">
<emphasis id="B964C635FFE44437FF57FEF30AE08413" bold="true" box="[189,475,264,293]" italics="true" pageId="11" pageNumber="132">Phylogenetic analysis</emphasis>
</paragraph>
<paragraph id="8BAF1A27FFE44437FF57FEC60FBA8731" blockId="11.[189,1399,317,519]" pageId="11" pageNumber="132">
Both phylogenetics reconstructions (i.e., BA and ML) agree in the basic topology of the trees (for simplicity we only show BA). Both 
<typeStatus id="54ABA485FFE44437FD6AFE9B0988844C" box="[640,691,352,378]" pageId="11" pageNumber="132">type</typeStatus>
specimens of 
<taxonomicName id="4C1061A4FFE44437FC83FE9A08CB844C" authority="Risaro &amp; Williams &amp; Pereyra &amp; Lauretta, 2020" authorityName="Risaro &amp; Williams &amp; Pereyra &amp; Lauretta" authorityYear="2020" box="[873,1008,353,378]" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="11" pageNumber="132" phylum="Cnidaria" rank="species" species="pomona" status="sp. nov.">
<emphasis id="B964C635FFE44437FC83FE9A08CB844C" box="[873,1008,353,378]" italics="true" pageId="11" pageNumber="132">U. pomona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2577B4EFFE44437FC11FE9B0F61844C" box="[1019,1114,352,378]" pageId="11" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
were grouped together with low support values within the same group (possibly because we only have one gene sequence for each species). 
<taxonomicName id="4C1061A4FFE44437FE83FE5D0ADE84F6" authorityName="Cuvier" authorityYear="1798" box="[361,485,422,448]" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="11" pageNumber="132" phylum="Cnidaria" rank="genus">
<emphasis id="B964C635FFE44437FE83FE5D0ADE84F6" box="[361,485,422,448]" italics="true" pageId="11" pageNumber="132">Umbellula</emphasis>
</taxonomicName>
spp. were recovered in two clusters, 
<taxonomicName id="4C1061A4FFE44437FC74FE5D0F2184F6" authorityName="Cuvier" authorityYear="1798" box="[926,1050,422,448]" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="11" pageNumber="132" phylum="Cnidaria" rank="genus">
<emphasis id="B964C635FFE44437FC74FE5D0F2184F6" box="[926,1050,422,448]" italics="true" pageId="11" pageNumber="132">Umbellula</emphasis>
</taxonomicName>
clade I including most of the included 
<taxonomicName id="4C1061A4FFE44437FEC6FE320A9384D5" authorityName="Cuvier" authorityYear="1798" box="[300,424,457,483]" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="11" pageNumber="132" phylum="Cnidaria" rank="genus">
<emphasis id="B964C635FFE44437FEC6FE320A9384D5" box="[300,424,457,483]" italics="true" pageId="11" pageNumber="132">Umbellula</emphasis>
</taxonomicName>
species and a second one (
<taxonomicName id="4C1061A4FFE44437FD1EFE32084B84D5" authorityName="Cuvier" authorityYear="1798" box="[756,880,457,483]" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="11" pageNumber="132" phylum="Cnidaria" rank="genus">
<emphasis id="B964C635FFE44437FD1EFE32084B84D5" box="[756,880,457,483]" italics="true" pageId="11" pageNumber="132">Umbellula</emphasis>
</taxonomicName>
clade II) including only 
<taxonomicName id="4C1061A4FFE44437FB75FE310E5484D5" authorityName="Pasternak" authorityYear="1964" box="[1183,1391,457,483]" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="11" pageNumber="132" phylum="Cnidaria" rank="species" species="monocephalus">
<emphasis id="B964C635FFE44437FB75FE310E5484D5" box="[1183,1391,457,483]" italics="true" pageId="11" pageNumber="132">U. monocephalus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C1061A4FFE44437FF57FE160AA48730" authority="Risaro &amp; Williams &amp; Pereyra &amp; Lauretta, 2020" authorityName="Risaro &amp; Williams &amp; Pereyra &amp; Lauretta" authorityYear="2020" box="[189,415,493,519]" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="11" pageNumber="132" phylum="Cnidaria" rank="species" species="pomona" status="sp. nov.">
<emphasis id="B964C635FFE44437FF57FE160AA48730" box="[189,415,493,519]" italics="true" pageId="11" pageNumber="132">Umbellula pomona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2577B4EFFE44437FE4CFE16093A8731" box="[422,513,493,519]" pageId="11" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C1061A4FFE44437FDD3FE16098E8731" authorityName="Cuvier" authorityYear="1798" box="[569,693,493,519]" class="Anthozoa" family="Umbellulidae" genus="Umbellula" kingdom="Animalia" order="Pennatulacea" pageId="11" pageNumber="132" phylum="Cnidaria" rank="genus">
<emphasis id="B964C635FFE44437FDD3FE16098E8731" box="[569,693,493,519]" italics="true" pageId="11" pageNumber="132">Umbellula</emphasis>
</taxonomicName>
sp. 2 from 
<bibRefCitation id="EF8167D6FFE44437FCD1FE160F218731" author="Dolan" box="[827,1050,493,519]" etAl="et al." firstAuthor="Dolan" pageId="11" pageNumber="132" pagination="610 - 618" refId="ref8861" refString="Dolan E., Tyler P. A., Yesson C. &amp; Rogers A. D. 2013. Phylogeny and systematics of deep-sea sea pens (Anthozoa: Octocorallia: Pennatulacea). Molecular Phylogenetics and Evolution 69: 610 - 618. https: // doi. org / 10.1016 / j. ympev. 2013.07.018" type="journal article" year="2013">
Dolan 
<emphasis id="B964C635FFE44437FC60FE1608FE8731" box="[906,965,493,519]" italics="true" pageId="11" pageNumber="132">et al.</emphasis>
(2013)
</bibRefCitation>
(
<figureCitation id="132B06A2FFE44437FBC1FE160F488731" box="[1067,1139,493,519]" captionStart="Fig" captionStartId="13.[189,232,1913,1939]" captionTargetBox="[164,1769,645,1493]" captionTargetId="figure-12@13.[163,1775,643,1497]" captionTargetPageId="13" captionText="Fig. 9. Phylogenetic reconstruction of the concatenated set of the two mitochondrial genes (mtMutS and ND2) based on the Bayesian analysis. Nodes include the Bayesian posterior probabilities. Arrows point to the new specimens of Umbellula Cuvier, 1798 included in the analysis." figureDoi="http://doi.org/10.5281/zenodo.14288728" httpUri="https://zenodo.org/record/14288728/files/figure.png" pageId="11" pageNumber="132">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>