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<document id="89EBBC56E15656573AD94B1D38A5C62A" ID-CLB-Dataset="21443" ID-DOI="10.11646/zootaxa.4816.3.4" ID-GBIF-Dataset="8de569a5-ae44-4994-8313-57163e9a4514" ID-ISSN="1175-5326" ID-Zenodo-Dep="3954447" ID-ZooBank="FA112B5F-E75E-4E6F-8E4C-E0D87674FCCA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595339868132" checkinUser="plazi" docAuthor="Ahyong, Shane T. &amp; Ng, Peter K. L." docDate="2020" docId="03B087FADC1FFFCAC798FD1BFBC52ACA" docLanguage="en" docName="zootaxa.4816.3.4.pdf" docOrigin="Zootaxa 4816 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arcotheres pollus Ahyong &amp; Ng 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="337" masterDocId="FF89FF82DC1EFFCEC70FFFABFFCA2C29" masterDocTitle="New species of pinnotherid crabs from Southeast Asia and Papua New Guinea (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="349" masterPageNumber="333" pageNumber="334" updateTime="1698843768168" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="94BE6C07790958F65E50BD539C0376D8">
<mods:title id="24C0329B6FCBF1D167F83B327DE58F05">New species of pinnotherid crabs from Southeast Asia and Papua New Guinea (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:name id="5594DA97D417582445889A5AF603C009" type="personal">
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<mods:roleTerm id="8A1209D3240BD38E1A5CCE719FFA16B8">Author</mods:roleTerm>
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<mods:namePart id="D422DE10D36619F51AA755E0FB1697C1">Ahyong, Shane T.</mods:namePart>
<mods:affiliation id="46EA745D809B995395FAEA72176A465A">Australian Museum Research Institute, Australian Museum, 1 William Street, Sydney NSW 2010, Australia, and School of Biological, Earth &amp; Environmental Sciences, University of New South Wales, NSW 2052, Australia.</mods:affiliation>
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<mods:namePart id="1246A692AAC0ADB325732C2FB24DAE58">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="E2E3E666E7626F4EA5AAC96913C18AAD">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<mods:title id="FBEC774495C40BB22B23F489082D4E0C">Zootaxa</mods:title>
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<mods:part id="062E0CF4BE001488BFC583C868EDED0D">
<mods:date id="0DE7B5E7BF104E5F6FD8FED3A850E896">2020</mods:date>
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<mods:number id="CA2050C303284AD79D225B60DBBFCF11">2020-07-17</mods:number>
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<mods:number id="C68B527543ACEC2C1E7778BC75E34E74">4816</mods:number>
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<mods:number id="E7A33D638F107BE09290E8273C3BD6CB">3</mods:number>
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<mods:identifier id="4F48EAEF56F37446ABE91C6F717FA03A" type="CLB-Dataset">21443</mods:identifier>
<mods:identifier id="9C266086BBF0D7D21FB9288767AE7DC7" type="DOI">10.11646/zootaxa.4816.3.4</mods:identifier>
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<treatment id="03B087FADC1FFFCAC798FD1BFBC52ACA" ID-DOI="http://doi.org/10.5281/zenodo.4330026" ID-GBIF-Taxon="165857110" ID-Zenodo-Dep="4330026" LSID="urn:lsid:plazi:treatment:03B087FADC1FFFCAC798FD1BFBC52ACA" httpUri="http://treatment.plazi.org/id/03B087FADC1FFFCAC798FD1BFBC52ACA" lastPageId="4" lastPageNumber="337" pageId="1" pageNumber="334">
<subSubSection id="C3036567DC1FFFCFC798FD1BFE1B2EE3" box="[151,465,688,715]" pageId="1" pageNumber="334" type="nomenclature">
<paragraph id="8BA636ECDC1FFFCFC798FD1BFE1B2EE3" blockId="1.[151,465,688,750]" box="[151,465,688,715]" pageId="1" pageNumber="334">
<heading id="D0EE8180DC1FFFCFC798FD1BFE1B2EE3" bold="true" box="[151,465,688,715]" fontSize="11" level="1" pageId="1" pageNumber="334" reason="1">
<emphasis id="B96DEAFEDC1FFFCFC798FD1BFE1B2EE3" bold="true" box="[151,465,688,715]" pageId="1" pageNumber="334">
<taxonomicName id="4C194D6FDC1FFFCFC798FD1BFEA62EE2" ID-CoL="84GPZ" authority="Ahyong &amp; Ng, 2020" authorityName="Ahyong &amp; Ng" authorityYear="2020" box="[151,364,688,715]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="334" phylum="Arthropoda" rank="species" species="pollus" status="sp. nov.">
<emphasis id="B96DEAFEDC1FFFCFC798FD1BFEA62EE2" bold="true" box="[151,364,688,715]" italics="true" pageId="1" pageNumber="334">Arcotheres pollus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E5785DC1FFFCFC67CFD1BFE1B2EE3" box="[371,465,688,714]" pageId="1" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3036567DC1FFFCFC798FD7FFED92EC7" box="[151,275,724,750]" pageId="1" pageNumber="334" type="description">
<paragraph id="8BA636ECDC1FFFCFC798FD7FFED92EC7" blockId="1.[151,465,688,750]" box="[151,275,724,750]" pageId="1" pageNumber="334">
(
<figureCitation id="13222A69DC1FFFCFC790FD7FFF272EC7" box="[159,237,724,750]" captionStart="FIGURE 1" captionStartId="2.[151,250,1836,1861]" captionTargetBox="[154,1431,320,1806]" captionTargetId="figure@2.[151,1436,309,1811]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres pollus sp. nov., male holotype, cl 2.3 mm, cw 2.4 mm, ZRC 1965.11.24.50. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–5; J–M, right P2–5; N, pleon; O, left G1, sternal view; P, left G1, pleonal view; Q, left G2, pleonal view. Scales: A = 1.0 mm; B = 0.2 mm; C–N = 1.0 mm; O–Q = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3954449" httpUri="https://zenodo.org/record/3954449/files/figure.png" pageId="1" pageNumber="334">Figs. 1</figureCitation>
,
<figureCitation id="13222A69DC1FFFCFC7F4FD7FFEC02EC7" box="[251,266,724,750]" captionStart="FIGURE 2" captionStartId="3.[151,250,1697,1722]" captionTargetBox="[159,1423,184,1670]" captionTargetId="figure@3.[151,1436,181,1672]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres pollus sp. nov., ovigerous female paratype, cl 3.6 mm, cw 4.6 mm, ZRC 1965.11.24.49. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–5; J–M, right P2–5. Scales: A = 2.0 mm; B = 0.5 mm; C–M = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3954451" httpUri="https://zenodo.org/record/3954451/files/figure.png" pageId="1" pageNumber="334">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3036567DC1FFFCFC798FCB7FD572F8B" pageId="1" pageNumber="334" type="materials_examined">
<paragraph id="8BA636ECDC1FFFCFC798FCB7FD572F8B" blockId="1.[151,1437,795,2011]" pageId="1" pageNumber="334">
<emphasis id="B96DEAFEDC1FFFCFC798FCB7FEF72F1F" bold="true" box="[151,317,795,822]" pageId="1" pageNumber="334">Type material</emphasis>
.
<typeStatus id="54A2884EDC1FFFCFC646FCB7FE2B2F1F" box="[329,481,796,822]" pageId="1" pageNumber="334" type="holotype">HOLOTYPE</typeStatus>
:
<collectionCode id="ED08AE29DC1FFFCFC6E4FCB7FDEB2F1F" box="[491,545,796,822]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="334">ZRC</collectionCode>
1965.11.24.49, male (cl
<quantity id="4CE19B09DC1FFFCFC43CFCB7FC402F1E" box="[819,906,796,823]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="334" unit="mm" value="2.3">2.3 mm</quantity>
, cw
<quantity id="4CE19B09DC1FFFCFC4B4FCB7FBD92F1E" box="[955,1043,796,823]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="334" unit="mm" value="2.4">2.4 mm</quantity>
), Paye or Paway Island, Mergui Archipelago,
<collectingCountry id="F30E767CDC1FFFCFC61DFCEAFE4B2F72" box="[274,385,833,859]" name="Myanmar" pageId="1" pageNumber="334">Myanmar</collectingCountry>
,
<geoCoordinate id="EE2D502BDC1FFFCFC684FC94FDDC2F73" box="[395,534,831,858]" degrees="11" direction="north" minutes="25" orientation="latitude" pageId="1" pageNumber="334" precision="15" seconds="00" value="11.416667">11°25’00”N</geoCoordinate>
,
<geoCoordinate id="EE2D502BDC1FFFCFC52EFCEBFD632F73" box="[545,681,832,858]" degrees="98" direction="east" minutes="31" orientation="longitude" pageId="1" pageNumber="334" precision="15" seconds="00" value="98.51667">98°31’00”E</geoCoordinate>
, coral reefs, shore collecting, from
<taxonomicName id="4C194D6FDC1FFFCFC33CFCEBFAF72F73" authority="Lamarck" authorityName="Lamarck" authorityYear="1819" box="[1075,1341,832,858]" class="Bivalvia" family="Malleidae" genus="Malleus" kingdom="Animalia" order="Pterioida" pageId="1" pageNumber="334" phylum="Mollusca" rank="species" species="albus">
<emphasis id="B96DEAFEDC1FFFCFC33CFCEBFB1B2F73" box="[1075,1233,832,858]" italics="true" pageId="1" pageNumber="334">Malleus albus</emphasis>
Lamarck
</taxonomicName>
,
<collectionCode id="ED08AE29DC1FFFCFC247FCEBFA562F73" box="[1352,1436,832,858]" pageId="1" pageNumber="334">RIMSS</collectionCode>
<taxonomicName id="4C194D6FDC1FFFCFC798FCCEFED42F57" box="[151,286,869,894]" family="Nemichthyidae" genus="Investigator" higherTaxonomySource="CoL" kingdom="Animalia" order="Anguilliformes" pageId="1" pageNumber="334" phylum="Chordata" rank="genus">
<emphasis id="B96DEAFEDC1FFFCFC798FCCEFED42F57" box="[151,286,869,894]" italics="true" pageId="1" pageNumber="334">Investigator</emphasis>
</taxonomicName>
, stn 593,
<date id="FFA7102CDC1FFFCFC687FCCFFD702F57" box="[392,698,868,894]" pageId="1" pageNumber="334" value="1914-02-24" valueMax="1914-03-03" valueMin="1914-02-24">24 February–3 March 1914</date>
.
<typeStatus id="54A2884EDC1FFFCFC5CBFCCFFC9E2F57" box="[708,852,868,894]" pageId="1" pageNumber="334" type="paratype">PARATYPE</typeStatus>
:
<collectionCode id="ED08AE29DC1FFFCFC453FCCFFC582F57" box="[860,914,868,894]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="334">ZRC</collectionCode>
1965.11.24.50, 1 ovigerous female (cl
<quantity id="4CE19B09DC1FFFCFC24EFCCFFA522F56" box="[1345,1432,868,895]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="334" unit="mm" value="3.6">3.6 mm</quantity>
, cw
<quantity id="4CE19B09DC1FFFCFC7B0FC23FED32F8A" box="[191,281,904,931]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="334" unit="mm" value="4.6">4.6 mm</quantity>
), collected together with
<typeStatus id="54A2884EDC1FFFCFC53CFC23FD532F8B" box="[563,665,904,930]" pageId="1" pageNumber="334" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036567DC1FFFCAC7C8FC00FB232EAE" lastPageId="4" lastPageNumber="337" pageId="1" pageNumber="334" type="description">
<paragraph id="8BA636ECDC1FFFCFC7C8FC00FC4A287F" blockId="1.[151,1437,795,2011]" pageId="1" pageNumber="334">
<emphasis id="B96DEAFEDC1FFFCFC7C8FC00FE9B2FEF" bold="true" box="[199,337,939,966]" pageId="1" pageNumber="334">Description</emphasis>
. MALE: Carapace irregularly subcircular. Front produced, anterior margin straight in dorsal view. Anterolateral margins poorly defined. Dorsum smooth, very few, minute, scattered setae; dorsal midline convex in profile. Epistome with narrow, triangular, interantennular septum; median buccal margin with acute median point. Antennular sinus smaller than orbit; antennules folded slightly obliquely. Antennal articles 1 and 2 fused to epistome. Eyes visible in dorsal view, filling orbit, cornea pigmented.
</paragraph>
<paragraph id="8BA636ECDC1FFFCFC7C8FBCBFC7E2922" blockId="1.[151,1437,795,2011]" pageId="1" pageNumber="334">Maxilliped 3 ischiomerus length about twice width; inner proximal margin convex; inner distal margin with blunt, obtuse angle; outer margin strongly convex. Carpus shorter than propodus. Propodus spatulate, gently tapering in distal half, apex rounded, dorsally and distal tally setose, length 4× dactylus length. Dactylus digitiform, distally setose, inserted slightly distal to propodal midlength, apex not reaching end of propodus. Exopod inner margin straight, outer margin convex; flagellum with 2 articles, distally setose.</paragraph>
<paragraph id="8BA636ECDC1FFFCFC7C8FABFFECB29CB" blockId="1.[151,1437,795,2011]" pageId="1" pageNumber="334">Chelipeds stout, inflated, symmetrical from left to right, surfaces setose. Dactylus and pollex relatively straight, crossing distally, without gape. Dactylus occlusal margin with triangular tooth proximal to midlength, margin straight in distal half, minutely denticulate. Pollex occlusal margin with blunt triangular tooth proximal to midlength, straight in distal half, sparsely setose; with fringe of setae on inner ventral margin extending to proximal end of palm. Propodus palm dorsal margin 1.1× height, as long as dactylus; ventral margin almost straight. Carpus unarmed.</paragraph>
<paragraph id="8BA636ECDC1FFFCFC7C8FA47FEDB2A93" blockId="1.[151,1437,795,2011]" pageId="1" pageNumber="334">
Pereopods 2–5 (walking legs 1–4) unarmed, symmetrical from left to right; covered in short, upright setae. Relative lengths in decreasing order: P4
<emphasis id="B96DEAFEDC1FFFCFC56EF9BAFDBE2A03" box="[609,628,1553,1578]" italics="true" pageId="1" pageNumber="334">&gt;</emphasis>
P3
<emphasis id="B96DEAFEDC1FFFCFC5AEF9BAFD7E2A03" box="[673,692,1553,1578]" italics="true" pageId="1" pageNumber="334">&gt;</emphasis>
P2
<emphasis id="B96DEAFEDC1FFFCFC5EFF9BAFD392A03" box="[736,755,1553,1578]" italics="true" pageId="1" pageNumber="334">&gt;</emphasis>
P5. P4 merus 0.5× pcl. P2–3 dactylus length subequal, 0.6× propodus length, evenly arcuate, distally spiniform. P4 dactylus 0.7× propodus length, slightly curved, styliform. P5 dactylus 1.1× propodus length, styliform but thicker and straighter than P4 dactylus; surfaces covered in upright setae, of similar length on flexor and extensor margins; flexor margin unarmed. Relative dactylus lengths: P5&gt; P4&gt; P3
<emphasis id="B96DEAFEDC1FFFCFC7DCF90AFF2C2A93" box="[211,230,1697,1722]" italics="true" pageId="1" pageNumber="334">=</emphasis>
P2.
</paragraph>
<paragraph id="8BA636ECDC1FFFCFC7C8F96FFD0B2B2A" blockId="1.[151,1437,795,2011]" pageId="1" pageNumber="334">Thoracic sternum anterior margin shallowly concave medially; sternites 1–3 indistinguishably fused; sternopleonal cavity reaching level of sternite ¾ suture.</paragraph>
<paragraph id="8BA636ECDC1FFFCFC7C8F8A7FDF42B63" blockId="1.[151,1437,795,2011]" pageId="1" pageNumber="334">Pleon slender, margins evenly converging distally; comprising 6 free somites and telson, widest at somites 1–3; telson wider than long, apex rounded.</paragraph>
<paragraph id="8BA636ECDC1FFFCFC7C8F8FFFC772B9F" blockId="1.[151,1437,795,2011]" pageId="1" pageNumber="334">Gonopod 1 elongate, slender, strongly arcuate, dorsoventrally compressed, about ¾ length of pleon, tapering to simple apex; margins setose, longest on outer margin. Gonopod 2 endopod simple, about ¼ length of gonopod 1; exopod short, slender, about ¼ length of endopod. Pleopods 3–5 absent.</paragraph>
<paragraph id="8BA636ECDC1FFFCCC7C8F86BFB882D36" blockId="1.[151,1437,795,2011]" lastBlockId="2.[151,1436,153,287]" lastPageId="2" lastPageNumber="335" pageId="1" pageNumber="334">FEMALE: Carapace ovate, slightly wider than long. Front weakly produced, anterior margin broadly rounded in dorsal view.Anterolateral margins poorly defined. Dorsum smooth, very few, minute, scattered setae; dorsal midline convex in profile. Epistome with narrow triangular interantennular septum; median buccal margin with obtuse median point. Antennular sinus of similar size to orbit; antennules folded slightly obliquely. Antennal articles 1 and 2 fused to epistome. Eyes not visible in dorsal view, filling orbit, cornea pigmented.</paragraph>
<caption id="DF666664DC1CFFCCC798F887FEF22B99" ID-DOI="http://doi.org/10.5281/zenodo.3954449" ID-Zenodo-Dep="3954449" httpUri="https://zenodo.org/record/3954449/files/figure.png" pageId="2" pageNumber="335" startId="2.[151,250,1836,1861]" targetBox="[154,1431,320,1806]" targetPageId="2">
<paragraph id="8BA636ECDC1CFFCCC798F887FEF22B99" blockId="2.[151,1436,1836,1969]" pageId="2" pageNumber="335">
<emphasis id="B96DEAFEDC1CFFCCC798F887FED92B6C" bold="true" box="[151,275,1836,1861]" pageId="2" pageNumber="335">FIGURE 1.</emphasis>
<taxonomicName id="4C194D6FDC1CFFCCC616F886FE012B6C" authority="Ahyong &amp; Ng, 2020" authorityName="Ahyong &amp; Ng" authorityYear="2020" box="[281,459,1837,1861]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="335" phylum="Arthropoda" rank="species" species="pollus" status="sp. nov.">
<emphasis id="B96DEAFEDC1CFFCCC616F886FE012B6C" box="[281,459,1837,1861]" italics="true" pageId="2" pageNumber="335">Arcotheres pollus</emphasis>
</taxonomicName>
<emphasis id="B96DEAFEDC1CFFCCC6DDF886FDE92B6C" bold="true" box="[466,547,1837,1861]" pageId="2" pageNumber="335">
<taxonomicNameLabel id="A25E5785DC1CFFCCC6DDF886FDE92B6C" box="[466,547,1837,1861]" pageId="2" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype, cl 2.3 mm, cw 2.4 mm, ZRC 1965.11.24.50. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–5; J–M, right P2–5; N, pleon; O, left G1, sternal view; P, left G1, pleonal view; Q, left G2, pleonal view. Scales: A = 1.0 mm; B = 0.2 mm; C–N = 1.0 mm; O–Q = 0.4 mm.
</paragraph>
</caption>
<caption id="DF666664DC1DFFCDC798F90AFD1C2B28" ID-DOI="http://doi.org/10.5281/zenodo.3954451" ID-Zenodo-Dep="3954451" httpUri="https://zenodo.org/record/3954451/files/figure.png" pageId="3" pageNumber="336" startId="3.[151,250,1697,1722]" targetBox="[159,1423,184,1670]" targetPageId="3">
<paragraph id="8BA636ECDC1DFFCDC798F90AFD1C2B28" blockId="3.[151,1436,1697,1794]" pageId="3" pageNumber="336">
<emphasis id="B96DEAFEDC1DFFCDC798F90AFEDC2A93" bold="true" box="[151,278,1697,1722]" pageId="3" pageNumber="336">FIGURE 2.</emphasis>
<taxonomicName id="4C194D6FDC1DFFCDC610F909FE192A93" authority="Ahyong &amp; Ng, 2020" authorityName="Ahyong &amp; Ng" authorityYear="2020" box="[287,467,1698,1722]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="pollus" status="sp. nov.">
<emphasis id="B96DEAFEDC1DFFCDC610F909FE192A93" box="[287,467,1698,1722]" italics="true" pageId="3" pageNumber="336">Arcotheres pollus</emphasis>
</taxonomicName>
<emphasis id="B96DEAFEDC1DFFCDC6D3F909FDFA2A93" bold="true" box="[476,560,1698,1722]" pageId="3" pageNumber="336">
<taxonomicNameLabel id="A25E5785DC1DFFCDC6D3F909FDFA2A93" box="[476,560,1698,1722]" pageId="3" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ovigerous female paratype, cl 3.6 mm, cw 4.6 mm, ZRC 1965.11.24.49. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–5; J–M, right P2–5. Scales: A = 2.0 mm; B = 0.5 mm; C–M = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BA636ECDC1DFFCDC7C8F89BFC7E2BF2" blockId="3.[151,1437,1840,2011]" pageId="3" pageNumber="336">Maxilliped 3 ischiomerus length about twice width; inner proximal margin convex; inner distal margin with blunt, obtuse angle; outer margin strongly convex. Carpus shorter than propodus. Propodus spatulate, gently tapering in distal half, apex rounded, dorsally and distally setose, length 0.5× dactylus length. Dactylus digitiform, distally setose, inserted slightly distal to propodal midlength, apex reaching end of propodus. Exopod inner margin straight, outer margin convex; flagellum with 2 articles, distally setose.</paragraph>
<paragraph id="8BA636ECDC1AFFCAC7C8FF32FB9E2D6A" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="337">Chelipeds symmetrical from left to right, surfaces setose. Dactylus and pollex relatively straight, crossing distally, without gape. Dactylus occlusal margin with blunt triangular tooth proximal to midlength, gently sinuous margin in distal half, sparsely setose. Pollex occlusal margin with blunt triangular tooth proximal to midlength, gently curved distal half, sparsely setose; with fringe of short setae on inner ventral margin. Propodus palm dorsal margin 1.3× height, 0.9× length of dactylus; ventral margin almost straight. Carpus unarmed.</paragraph>
<paragraph id="8BA636ECDC1AFFCAC7C8FEE6FB132E16" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="337">
Pereopods 2–5 (walking legs 1–4) unarmed, covered in short, upright setae, most pronounced on flexor and extensor margins of merus, carpus, propodus, and dactylus. P2, 3 and 5 similar, symmetrical from left to right; right P4 length 1.2× longer than left; relative lengths in decreasing order: P4
<emphasis id="B96DEAFEDC1AFFCAC4BBFE3DFC0D2D86" box="[948,967,406,431]" italics="true" pageId="4" pageNumber="337">&gt;</emphasis>
P5
<emphasis id="B96DEAFEDC1AFFCAC4FEFE3DFBCE2D86" box="[1009,1028,406,431]" italics="true" pageId="4" pageNumber="337">&gt;</emphasis>
P3
<emphasis id="B96DEAFEDC1AFFCAC320FE3DFB882D86" box="[1071,1090,406,431]" italics="true" pageId="4" pageNumber="337">&gt;</emphasis>
P2. Longer P4 merus 0.5× pcl. P2–3 dactylus 0.5× propodus length, evenly arcuate, distally spiniform, P3 dactylus slightly longer that of P2. P4 dactylus styliform, unarmed, 0.9× propodus length (right), 0.8× propodus length (left). P5 dactylus 1.1× propodus length, styliform but thicker than P4 dactylus; surfaces covered in upright setae, slightly longer on flexor margin; flexor margin unarmed; apex a short slender spine. Relative dactylus lengths: P5&gt; P4&gt; P3
<emphasis id="B96DEAFEDC1AFFCAC395FD8DFB672E16" box="[1178,1197,550,575]" italics="true" pageId="4" pageNumber="337">&gt;</emphasis>
P2.
</paragraph>
<paragraph id="8BA636ECDC1AFFCAC7C8FDE2FAFA2E4A" blockId="4.[151,1437,153,1763]" box="[199,1328,585,611]" pageId="4" pageNumber="337">Thoracic sternum anterior margin shallowly concave medially; sternites 1–3 indistinguishably fused.</paragraph>
<paragraph id="8BA636ECDC1AFFCAC7C8FDC6FB232EAE" blockId="4.[151,1437,153,1763]" box="[199,1257,621,647]" pageId="4" pageNumber="337">Pleon of 6 free somites and telson, extending to buccal region, covering bases of walking legs.</paragraph>
</subSubSection>
<subSubSection id="C3036567DC1AFFCAC7C8FD3AFD272E85" box="[199,749,657,684]" pageId="4" pageNumber="337" type="biology_ecology">
<paragraph id="8BA636ECDC1AFFCAC7C8FD3AFD272E85" blockId="4.[151,1437,153,1763]" box="[199,749,657,684]" pageId="4" pageNumber="337">
<emphasis id="B96DEAFEDC1AFFCAC7C8FD3AFF352E82" bold="true" box="[199,255,657,683]" pageId="4" pageNumber="337">Host</emphasis>
.
<taxonomicName id="4C194D6FDC1AFFCAC602FD3AFD232E85" authority="Lamarck, 1819" authorityName="Lamarck" authorityYear="1819" box="[269,745,657,684]" class="Bivalvia" family="Malleidae" genus="Malleus" kingdom="Animalia" order="Pterioida" pageId="4" pageNumber="337" phylum="Mollusca" rank="species" species="albus">
<emphasis id="B96DEAFEDC1AFFCAC602FD3AFE612E82" box="[269,427,657,683]" italics="true" pageId="4" pageNumber="337">Malleus albus</emphasis>
Lamarck, 1819 (Malleidae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036567DC1AFFCAC7C8FD1EFB4C2EE6" box="[199,1158,693,719]" pageId="4" pageNumber="337" type="etymology">
<paragraph id="8BA636ECDC1AFFCAC7C8FD1EFB4C2EE6" blockId="4.[151,1437,153,1763]" box="[199,1158,693,719]" pageId="4" pageNumber="337">
<emphasis id="B96DEAFEDC1AFFCAC7C8FD1EFE822EE6" bold="true" box="[199,328,693,719]" pageId="4" pageNumber="337">Etymology</emphasis>
. Named
<taxonomicName id="4C194D6FDC1AFFCAC6A1FD1EFE382EE6" authorityName="Ahyong &amp; Ng" authorityYear="2020" box="[430,498,693,719]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="pollus">
<emphasis id="B96DEAFEDC1AFFCAC6A1FD1EFE382EE6" box="[430,498,693,719]" italics="true" pageId="4" pageNumber="337">pollus</emphasis>
</taxonomicName>
, small, alluding to the diminutive adult size of the species.
</paragraph>
</subSubSection>
<subSubSection id="C3036567DC1AFFCAC7C8FD72FCAA2A96" pageId="4" pageNumber="337" type="discussion">
<paragraph id="8BA636ECDC1AFFCAC7C8FD72FBA42856" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="337">
<emphasis id="B96DEAFEDC1AFFCAC7C8FD72FEFE2EDA" bold="true" box="[199,308,729,755]" pageId="4" pageNumber="337">Remarks</emphasis>
.
<taxonomicName id="4C194D6FDC1AFFCAC64CFD72FDCD2EDA" authority="Ahyong &amp; Ng, 2020" authorityName="Ahyong &amp; Ng" authorityYear="2020" box="[323,519,729,755]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="pollus" status="sp. nov.">
<emphasis id="B96DEAFEDC1AFFCAC64CFD72FDCD2EDA" box="[323,519,729,755]" italics="true" pageId="4" pageNumber="337">Arcotheres pollus</emphasis>
</taxonomicName>
<emphasis id="B96DEAFEDC1AFFCAC500FD72FDA02EDA" bold="true" box="[527,618,729,755]" pageId="4" pageNumber="337">
<taxonomicNameLabel id="A25E5785DC1AFFCAC500FD72FDA02EDA" box="[527,618,729,755]" pageId="4" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is readily separated from all known congeners by the finely setose carapace (glabrous in congeners), distinctly setose pereopods in females with setose cheliped (P1) surfaces and setose flexor and extensor margins of the merus, carpus, propodus and dactylus of P2–5. The cheliped setation in females of other species of
<taxonomicName id="4C194D6FDC1AFFCAC663FCEEFE2F2F76" authorityName="Manning" authorityYear="1993" box="[364,485,837,863]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC1AFFCAC663FCEEFE2F2F76" box="[364,485,837,863]" italics="true" pageId="4" pageNumber="337">Arcotheres</emphasis>
</taxonomicName>
is limited to the fringe on the lower margin of the palm, a small brush of setae sometimes on the inner margin of the carpus, and scattered setae on the occlusal margins and tips of the fingers (
<bibRefCitation id="EF884B1DDC1AFFCAC230FCC2FF182F8E" author="Campos, E." pageId="4" pageNumber="337" pagination="493 - 497" refId="ref9391" refString="Campos, E. (2001) A new crab species of the genus Arcotheres Manning, 1993, from Thailand (Crustacea, Brachyura, Pinnotheridae). Zoosystema, 23, 493 - 497." type="journal article" year="2001">Campos 2001</bibRefCitation>
;
<bibRefCitation id="EF884B1DDC1AFFCAC7D0FC26FE1F2F8E" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[223,469,909,935]" pageId="4" pageNumber="337" pagination="191 - 226" refId="ref9096" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007 a) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng 2007a</bibRefCitation>
;
<bibRefCitation id="EF884B1DDC1AFFCAC6ECFC26FD9D2F8E" author="Ng, P. K. L." box="[483,599,909,935]" pageId="4" pageNumber="337" pagination="474 - 485" refId="ref10601" refString="Ng, P. K. L. (2018 b) Arcotheres placunicola, a new species of pea crab (Crustacea: Brachyura: Pinnotheridae) from the windowpane shell, Placuna ephippium Philipsson, 1788 (Placunidae) in Singapore. Raffles Bulletin of Zoology, 66, 474 - 485." type="journal article" year="2018">Ng 2018b</bibRefCitation>
;
<bibRefCitation id="EF884B1DDC1AFFCAC56AFC26FC8A2F8E" author="Trivedi, J. N. &amp; Campos, E. &amp; Vachhrajani, K. D." box="[613,832,909,935]" pageId="4" pageNumber="337" pagination="195 - 200" refId="ref11972" refString="Trivedi, J. N., Campos, E. &amp; Vachhrajani, K. D. (2018) A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India. Zootaxa, 4433 (1), 195 - 200. https: // doi. org / 10.11646 / zootaxa. 4433.1.13" type="journal article" year="2018">
Trivedi
<emphasis id="B96DEAFEDC1AFFCAC5CFFC25FD302F8E" box="[704,762,909,935]" italics="true" pageId="4" pageNumber="337">et al.</emphasis>
2018
</bibRefCitation>
). On the walking legs of females of other species of
<taxonomicName id="4C194D6FDC1AFFCAC798FC1AFEDA2FE2" authorityName="Manning" authorityYear="1993" box="[151,272,945,971]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC1AFFCAC798FC1AFEDA2FE2" box="[151,272,945,971]" italics="true" pageId="4" pageNumber="337">Arcotheres</emphasis>
</taxonomicName>
, the P5 dactylus is always setose, but P2–5 are otherwise almost glabrous, at most with a few setae on the flexor margin of the dactylus and P2–4, and sometimes with setae on the flexor margin of P2–5 propodi. The male of
<taxonomicName id="4C194D6FDC1AFFCAC7FCFC52FE9C283A" authorityName="Ahyong &amp; Ng" authorityYear="2020" box="[243,342,1017,1043]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="pollus">
<emphasis id="B96DEAFEDC1AFFCAC7FCFC52FE9C283A" box="[243,342,1017,1043]" italics="true" pageId="4" pageNumber="337">A. pollus</emphasis>
</taxonomicName>
is setose as in the female, but too few males of other species of
<taxonomicName id="4C194D6FDC1AFFCAC329FC52FB55283A" authorityName="Manning" authorityYear="1993" box="[1062,1183,1017,1043]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC1AFFCAC329FC52FB55283A" box="[1062,1183,1017,1043]" italics="true" pageId="4" pageNumber="337">Arcotheres</emphasis>
</taxonomicName>
are known to permit a comprehensive comparison.As with the female, the carapace and cheliped setation will separate male
<taxonomicName id="4C194D6FDC1AFFCAC3F1FBB6FAAA281E" authorityName="Ahyong &amp; Ng" authorityYear="2020" box="[1278,1376,1053,1079]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="pollus">
<emphasis id="B96DEAFEDC1AFFCAC3F1FBB6FAAA281E" box="[1278,1376,1053,1079]" italics="true" pageId="4" pageNumber="337">A. pollus</emphasis>
</taxonomicName>
from known males of other species of
<taxonomicName id="4C194D6FDC1AFFCAC502FBEAFD4C2872" authorityName="Manning" authorityYear="1993" box="[525,646,1089,1115]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC1AFFCAC502FBEAFD4C2872" box="[525,646,1089,1115]" italics="true" pageId="4" pageNumber="337">Arcotheres</emphasis>
</taxonomicName>
, which lack dorsal carapace setae and or setae on the outer surface of the palm. As in other known males of
<taxonomicName id="4C194D6FDC1AFFCAC54DFBCEFD712856" authorityName="Manning" authorityYear="1993" box="[578,699,1125,1151]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC1AFFCAC54DFBCEFD712856" box="[578,699,1125,1151]" italics="true" pageId="4" pageNumber="337">Arcotheres</emphasis>
</taxonomicName>
, the G2 exopod is present in
<taxonomicName id="4C194D6FDC1AFFCAC30CFBCEFBAC2856" authorityName="Ahyong &amp; Ng" authorityYear="2020" box="[1027,1126,1125,1151]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="pollus">
<emphasis id="B96DEAFEDC1AFFCAC30CFBCEFBAC2856" box="[1027,1126,1125,1151]" italics="true" pageId="4" pageNumber="337">A. pollus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA636ECDC1AFFCAC7C8FB22FED529EA" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="337">
<taxonomicName id="4C194D6FDC1AFFCAC7C8FB22FE46288A" authorityName="Ahyong &amp; Ng" authorityYear="2020" box="[199,396,1161,1187]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="pollus">
<emphasis id="B96DEAFEDC1AFFCAC7C8FB22FE46288A" box="[199,396,1161,1187]" italics="true" pageId="4" pageNumber="337">Arcotheres pollus</emphasis>
</taxonomicName>
is morphologically nearest to those species in the genus with females having a transversely ovate carapace (
<figureCitation id="13222A69DC1AFFCAC641FB06FE6D28EE" box="[334,423,1197,1223]" captionStart="FIGURE 2" captionStartId="3.[151,250,1697,1722]" captionTargetBox="[159,1423,184,1670]" captionTargetId="figure@3.[151,1436,181,1672]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres pollus sp. nov., ovigerous female paratype, cl 3.6 mm, cw 4.6 mm, ZRC 1965.11.24.49. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–5; J–M, right P2–5. Scales: A = 2.0 mm; B = 0.5 mm; C–M = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3954451" httpUri="https://zenodo.org/record/3954451/files/figure.png" pageId="4" pageNumber="337">Fig. 2A</figureCitation>
) and long, slender, almost styliform dactyli of P4 and 5 that are about twice the length of the P2 and 3 dactyli (
<figureCitation id="13222A69DC1AFFCAC68DFB7AFDE828C2" box="[386,546,1233,1259]" captionStart="FIGURE 2" captionStartId="3.[151,250,1697,1722]" captionTargetBox="[159,1423,184,1670]" captionTargetId="figure@3.[151,1436,181,1672]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres pollus sp. nov., ovigerous female paratype, cl 3.6 mm, cw 4.6 mm, ZRC 1965.11.24.49. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–5; J–M, right P2–5. Scales: A = 2.0 mm; B = 0.5 mm; C–M = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3954451" httpUri="https://zenodo.org/record/3954451/files/figure.png" pageId="4" pageNumber="337">Fig. 2H, L, M</figureCitation>
):
<taxonomicName id="4C194D6FDC1AFFCAC53AFB7AFC5C28C2" authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[565,918,1232,1259]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="boninensis">
<emphasis id="B96DEAFEDC1AFFCAC53AFB7AFD0628C2" box="[565,716,1233,1259]" italics="true" pageId="4" pageNumber="337">A. boninensis</emphasis>
(
<bibRefCitation id="EF884B1DDC1AFFCAC5D5FB7AFC4428C2" author="Stimpson, W." box="[730,910,1232,1259]" pageId="4" pageNumber="337" pagination="93 - 110" refId="ref11901" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110. https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C194D6FDC1AFFCAC4AEFB7AFB1C28C2" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[929,1238,1232,1259]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="B96DEAFEDC1AFFCAC4AEFB7AFBE128C2" box="[929,1067,1233,1259]" italics="true" pageId="4" pageNumber="337">A. pernicola</emphasis>
(
<bibRefCitation id="EF884B1DDC1AFFCAC336FB7AFB0428C2" author="Burger, O." box="[1081,1230,1232,1259]" pageId="4" pageNumber="337" pagination="361 - 390" refId="ref9358" refString="Burger, O. (1895) Ein Beitrag zur kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C194D6FDC1AFFCAC203FB7AFE8E2926" authority="(Alcock, 1900)" baseAuthorityName="Alcock" baseAuthorityYear="1900" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="purpureus">
<emphasis id="B96DEAFEDC1AFFCAC203FB7AFA5628C2" box="[1292,1436,1233,1259]" italics="true" pageId="4" pageNumber="337">A. purpureus</emphasis>
(
<bibRefCitation id="EF884B1DDC1AFFCAC790FB5EFEF12926" author="Alcock, A." box="[159,315,1269,1295]" pageId="4" pageNumber="337" pagination="279 - 486" refId="ref9266" refString="Alcock, A. (1900) Materials for a carcinological fauna of India. No. 6. Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, Calcutta, 69 (3), 279 - 486." type="journal article" year="1900">Alcock, 1900</bibRefCitation>
)
</taxonomicName>
. Aside from the pereopodal setation,
<taxonomicName id="4C194D6FDC1AFFCAC5EBFB5EFC8D2926" authorityName="Ahyong &amp; Ng" authorityYear="2020" box="[740,839,1269,1295]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="pollus">
<emphasis id="B96DEAFEDC1AFFCAC5EBFB5EFC8D2926" box="[740,839,1269,1295]" italics="true" pageId="4" pageNumber="337">A. pollus</emphasis>
</taxonomicName>
is readily separated from each of the aforementioned species by the shape of the propodus of maxilliped 3, which is distally thicker and less tapering (
<figureCitation id="13222A69DC1AFFCAC3FBFAB2FA81291A" box="[1268,1355,1305,1331]" captionStart="FIGURE 2" captionStartId="3.[151,250,1697,1722]" captionTargetBox="[159,1423,184,1670]" captionTargetId="figure@3.[151,1436,181,1672]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres pollus sp. nov., ovigerous female paratype, cl 3.6 mm, cw 4.6 mm, ZRC 1965.11.24.49. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–5; J–M, right P2–5. Scales: A = 2.0 mm; B = 0.5 mm; C–M = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3954451" httpUri="https://zenodo.org/record/3954451/files/figure.png" pageId="4" pageNumber="337">Fig. 2B</figureCitation>
versus Alcock &amp; McArdle 1902: pl. 57 fig. 6b;
<bibRefCitation id="EF884B1DDC1AFFCAC550FA96FD25297E" author="Burger, O." box="[607,751,1340,1367]" pageId="4" pageNumber="337" pagination="361 - 390" refId="ref9358" refString="Burger, O. (1895) Ein Beitrag zur kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger 1895</bibRefCitation>
: pl 10 fig. 16;
<bibRefCitation id="EF884B1DDC1AFFCAC49CFA96FB96297E" author="Dai, A. &amp; Yang, S." box="[915,1116,1341,1367]" pageId="4" pageNumber="337" refId="ref9428" refString="Dai, A. &amp; Yang, S. (1991) Crabs of the China Seas. China Ocean Press, Beijing, 608 pp., 74 pls." type="book" year="1991">Dai &amp; Yang 1991</bibRefCitation>
: fig. 210.2), and the absence of one or more rows of denticles on the distoflexor margin of the dactylus of P5 (
<figureCitation id="13222A69DC1AFFCAC33FFACAFB5A2952" box="[1072,1168,1377,1403]" captionStart="FIGURE 2" captionStartId="3.[151,250,1697,1722]" captionTargetBox="[159,1423,184,1670]" captionTargetId="figure@3.[151,1436,181,1672]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres pollus sp. nov., ovigerous female paratype, cl 3.6 mm, cw 4.6 mm, ZRC 1965.11.24.49. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–5; J–M, right P2–5. Scales: A = 2.0 mm; B = 0.5 mm; C–M = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3954451" httpUri="https://zenodo.org/record/3954451/files/figure.png" pageId="4" pageNumber="337">Fig. 2M</figureCitation>
).
<taxonomicName id="4C194D6FDC1AFFCAC3ABFACAFA562952" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1188,1436,1377,1403]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="boninensis">
<emphasis id="B96DEAFEDC1AFFCAC3ABFACAFA562952" box="[1188,1436,1377,1403]" italics="true" pageId="4" pageNumber="337">Arcotheres boninensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D6FDC1AFFCAC7C8FA2EFE9829B6" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[199,338,1413,1439]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="B96DEAFEDC1AFFCAC7C8FA2EFE9829B6" box="[199,338,1413,1439]" italics="true" pageId="4" pageNumber="337">A. pernicola</emphasis>
</taxonomicName>
have two short rows of distoflexor spines on the P5 dactylus, and
<taxonomicName id="4C194D6FDC1AFFCAC330FA2EFB1A29B6" baseAuthorityName="Alcock" baseAuthorityYear="1900" box="[1087,1232,1413,1439]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="purpureus">
<emphasis id="B96DEAFEDC1AFFCAC330FA2EFB1A29B6" box="[1087,1232,1413,1439]" italics="true" pageId="4" pageNumber="337">A. purpureus</emphasis>
</taxonomicName>
, one row (
<bibRefCitation id="EF884B1DDC1AFFCAC246FA2EFED929EA" author="Trivedi, J. N. &amp; Gosavi, S. &amp; Vachhrajani, K. D. &amp; Ng, P. K. L." pageId="4" pageNumber="337" pagination="587 - 593" refId="ref12040" refString="Trivedi, J. N., Gosavi, S., Vachhrajani, K. D. &amp; Ng, P. K. L. (2019) Redescription of Arcotheres pernicola (Burger, 1895) (Crustacea: Decapoda: Brachyura: Pinnotheridae) from the oyster Magallana gryphoides (Schlotheim, 1820) in India. Zootaxa, 4706 (4), 587 - 593. https: // doi. org / 10.11646 / zootaxa. 4706.4.8" type="journal article" year="2019">
Trivedi
<emphasis id="B96DEAFEDC1AFFCAC798FA01FF0229EA" box="[151,200,1449,1475]" italics="true" pageId="4" pageNumber="337">et al</emphasis>
. 2019
</bibRefCitation>
).
</paragraph>
<paragraph id="8BA636ECDC1AFFCAC7C8FA66FCAA2A96" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="337">
With males and females already mature by cl
<quantity id="4CE19B09DC1AFFCAC5CCFA66FCD229CE" box="[707,792,1485,1511]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="337" unit="mm" value="2.3">2.3 mm</quantity>
and
<quantity id="4CE19B09DC1AFFCAC442FA66FC6F29CE" box="[845,933,1485,1511]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="337" unit="mm" value="3.6">3.6 mm</quantity>
, respectively,
<taxonomicName id="4C194D6FDC1AFFCAC34DFA66FB6F29CE" authorityName="Ahyong &amp; Ng" authorityYear="2020" box="[1090,1189,1485,1511]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="pollus">
<emphasis id="B96DEAFEDC1AFFCAC34DFA66FB6F29CE" box="[1090,1189,1485,1511]" italics="true" pageId="4" pageNumber="337">A. pollus</emphasis>
</taxonomicName>
is the smallest known species of the genus. The next smallest species of
<taxonomicName id="4C194D6FDC1AFFCAC5D4FA5AFC9E2A22" authorityName="Manning" authorityYear="1993" box="[731,852,1521,1547]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC1AFFCAC5D4FA5AFC9E2A22" box="[731,852,1521,1547]" italics="true" pageId="4" pageNumber="337">Arcotheres</emphasis>
</taxonomicName>
appears to be
<taxonomicName id="4C194D6FDC1AFFCAC4F1FA5AFBBF2A22" baseAuthorityName="Bürger" baseAuthorityYear="1895" box="[1022,1141,1521,1547]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="B96DEAFEDC1AFFCAC4F1FA5AFBBF2A22" box="[1022,1141,1521,1547]" italics="true" pageId="4" pageNumber="337">A. exiguus</emphasis>
</taxonomicName>
, being mature (at least in females) by cl
<quantity id="4CE19B09DC1AFFCAC631F9BEFE5E2A06" box="[318,404,1557,1583]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="4" pageNumber="337" unit="mm" value="4.6">4.6 mm</quantity>
(cf. Ahyong &amp; Ng 2007). The
<typeStatus id="54A2884EDC1AFFCAC5FDF9BEFCE82A06" box="[754,802,1557,1583]" pageId="4" pageNumber="337">type</typeStatus>
material of
<taxonomicName id="4C194D6FDC1AFFCAC4A4F9BEFBC42A06" authorityName="Ahyong &amp; Ng" authorityYear="2020" box="[939,1038,1557,1583]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="pollus">
<emphasis id="B96DEAFEDC1AFFCAC4A4F9BEFBC42A06" box="[939,1038,1557,1583]" italics="true" pageId="4" pageNumber="337">A. pollus</emphasis>
</taxonomicName>
was taken from the hammer oyster (
<taxonomicName id="4C194D6FDC1AFFCAC7AEF992FE8B2A7A" authorityName="Lamarck" authorityYear="1819" box="[161,321,1593,1619]" class="Bivalvia" family="Malleidae" genus="Malleus" kingdom="Animalia" order="Pterioida" pageId="4" pageNumber="337" phylum="Mollusca" rank="species" species="albus">
<emphasis id="B96DEAFEDC1AFFCAC7AEF992FE8B2A7A" box="[161,321,1593,1619]" italics="true" pageId="4" pageNumber="337">Malleus albus</emphasis>
</taxonomicName>
) during shore collecting on coral reef. Other invertebrates taken from the same station, include a sipunculan worm (
<taxonomicName id="4C194D6FDC1AFFCAC663F9F6FCDC2A5E" authority="Keferstein, 1865" authorityName="Keferstein" authorityYear="1865" box="[364,790,1628,1655]" class="Sipunculidea" family="Sipunculidae" genus="Sipunculus" kingdom="Animalia" order="Sipunculiformes" pageId="4" pageNumber="337" phylum="Sipuncula" rank="species" species="robustus">
<emphasis id="B96DEAFEDC1AFFCAC663F9F6FD842A5E" box="[364,590,1629,1655]" italics="true" pageId="4" pageNumber="337">Sipunculus robustus</emphasis>
Keferstein, 1865
</taxonomicName>
;
<bibRefCitation id="EF884B1DDC1AFFCAC42CF9F6FC0A2A51" author="Prashad, B." box="[803,960,1629,1656]" pageId="4" pageNumber="337" pagination="232 - 238" refId="ref11382" refString="Prashad, B. (1936) On the Sipunculoidea from Indian waters in the Indian Museum, Calcutta. Records of the Indian Museum, 38, 232 - 238, pl. IX." type="journal article" year="1936">Prashad 1936</bibRefCitation>
), polychaete worms (
<taxonomicName id="4C194D6FDC1AFFCAC3BBF9F6FE8E2AB2" authority="Leuckart, 1927" authorityName="Leuckart" authorityYear="1927" class="Polychaeta" family="Sabellariidae" genus="Sabellaria" kingdom="Animalia" order="Sabellida" pageId="4" pageNumber="337" phylum="Annelida" rank="species" species="spinulosa">
<emphasis id="B96DEAFEDC1AFFCAC3BBF9F6FA562A5E" box="[1204,1436,1629,1655]" italics="true" pageId="4" pageNumber="337">Sabellaria spinulosa</emphasis>
Leuckart, 1927
</taxonomicName>
,
<taxonomicName id="4C194D6FDC1AFFCAC640F92AFCFA2AB2" authority="Beddard, 1887" authorityName="Beddard" authorityYear="1887" box="[335,816,1665,1691]" class="Polychaeta" family="Sabellidae" genus="Branchiomma" kingdom="Animalia" order="Sabellida" pageId="4" pageNumber="337" phylum="Annelida" rank="species" species="intermedium">
<emphasis id="B96DEAFEDC1AFFCAC640F92AFD482AB2" box="[335,642,1665,1691]" italics="true" pageId="4" pageNumber="337">Branchiomma intermedium</emphasis>
Beddard, 1887
</taxonomicName>
,
<taxonomicName id="4C194D6FDC1AFFCAC434F92AFB0C2AB2" authority="Savigny, 1822" authorityName="Savigny" authorityYear="1822" box="[827,1222,1665,1691]" class="Polychaeta" family="Sabellidae" genus="Sabellastarte" kingdom="Animalia" order="Sabellida" pageId="4" pageNumber="337" phylum="Annelida" rank="species" species="indica">
<emphasis id="B96DEAFEDC1AFFCAC434F92AFBD62AB2" box="[827,1052,1665,1691]" italics="true" pageId="4" pageNumber="337">Sabellastarte indica</emphasis>
Savigny, 1822
</taxonomicName>
;
<bibRefCitation id="EF884B1DDC1AFFCAC3DEF92AFAAB2AB2" author="Fauvel, P." box="[1233,1377,1665,1691]" pageId="4" pageNumber="337" pagination="1 - 262" refId="ref9545" refString="Fauvel, P. (1932) Annelida Polychaeta of the Indian Museum. Memoirs of the Indian Museum, 12 (1), 1 - 262, pls. I-VIII." type="journal article" year="1932">Fauvel 1932</bibRefCitation>
), and a mud shrimp (
<taxonomicName id="4C194D6FDC1AFFCAC64EF90EFD6D2A96" authority="Sankolli, 1972" authorityName="Sankolli" authorityYear="1972" box="[321,679,1701,1727]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="kempi">
<emphasis id="B96DEAFEDC1AFFCAC64EF90EFE302A96" box="[321,506,1701,1727]" italics="true" pageId="4" pageNumber="337">Upogebia kempi</emphasis>
<bibRefCitation id="EF884B1DDC1AFFCAC50EF90EFD6D2A96" author="Sankolli, K. N." box="[513,679,1701,1727]" pageId="4" pageNumber="337" pagination="671 - 682" refId="ref11677" refString="Sankolli, K. N. (1972) The Thalassinoidea (Crustacea, Anomura) of Maharashtra. Journal of the Bombay Natural History Museum, 68 (3), 671 - 682." type="journal article" year="1972">Sankolli, 1972</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EF884B1DDC1AFFCAC5BCF90EFC992A96" author="Sankolli, K. N." box="[691,851,1701,1727]" pageId="4" pageNumber="337" pagination="671 - 682" refId="ref11677" refString="Sankolli, K. N. (1972) The Thalassinoidea (Crustacea, Anomura) of Maharashtra. Journal of the Bombay Natural History Museum, 68 (3), 671 - 682." type="journal article" year="1972">Sankolli 1972</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3036567DC1AFFCAC7C8F963FBC52ACA" box="[199,1039,1736,1763]" pageId="4" pageNumber="337" type="distribution">
<paragraph id="8BA636ECDC1AFFCAC7C8F963FBC52ACA" blockId="4.[151,1437,153,1763]" box="[199,1039,1736,1763]" pageId="4" pageNumber="337">
<emphasis id="B96DEAFEDC1AFFCAC7C8F963FE932ACA" bold="true" box="[199,345,1736,1763]" pageId="4" pageNumber="337">Distribution</emphasis>
. Known only from the
<typeStatus id="54A2884EDC1AFFCAC56FF962FD5A2ACA" box="[608,656,1737,1763]" pageId="4" pageNumber="337">type</typeStatus>
locality, Paway Island,
<collectingCountry id="F30E767CDC1AFFCAC493F962FBC12ACA" box="[924,1035,1737,1763]" name="Myanmar" pageId="4" pageNumber="337">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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