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<document id="E0BF2F69AFEADE59D17F59C2074E242F" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B38FFCEFF79F2BAFEB8F8F8" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena variopaca Perkins 2011, new species" docType="treatment" docVersion="6" lastPageNumber="94" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="94" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="15A30E9177BBEAC10996C14CEF7750C0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="58ED7F59802182BF14EDBDACF17BA0A4">
<mods:title id="5A4D7D8366890CA39DB9CE99DCDD06CA">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
</mods:titleInfo>
<mods:name id="5A404140EFEB2B76B55E8F6E9217317E" type="personal">
<mods:role id="D0B0070A63EF8EDF3CFF9CA55537AC72">
<mods:roleTerm id="145EF1953C3D43181CD5961DD369DAAD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="61DDDDDCECBC2DC28BB65455D3DE306A">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="F0353F7F8AF26CB0AB8148F761D36103">text</mods:typeOfResource>
<mods:relatedItem id="B86FE20003C91BEEEA4BED517AA2C960" type="host">
<mods:titleInfo id="3491B9996065A0ADEB8313AEF02C3BA5">
<mods:title id="A1DFC4B653D8F6C83772EA9F3D067506">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="41645D48ECD9A71392B7839C7D28F2F4">
<mods:date id="A90A3F86CFCBFCAEAAC3B8A89B1A9122">2011</mods:date>
<mods:detail id="76CFF9C3CEBFC7F8B7615ECF06D755FD" type="pubDate">
<mods:number id="9EE96F780E337DF6CCC9CD9DF8DE17A4">2011-07-08</mods:number>
</mods:detail>
<mods:detail id="1CCE97B326C9CB3BDCDEE60559352F74" type="volume">
<mods:number id="0683A56BDD2AC1A8D185ED3CDF4246BB">2944</mods:number>
</mods:detail>
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<mods:number id="990C3C8CCB86D702DA633102B3AFBDF2">1</mods:number>
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<mods:location id="DCDE28971A00072669560F6A3E37537B">
<mods:url id="704110A59731B1F7E146C894FB2957E7">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
</mods:location>
<mods:classification id="EB845BCF1CE1ADCCF371E4C1C1C177DB">journal article</mods:classification>
<mods:identifier id="2245651B853C4325419834A38DE82971" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
<mods:identifier id="618614C69C646A3431D552BFCB626DD0" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="2E1456B913CCF267A4CBE7C0E5740AB4" type="Zenodo-Dep">5283420</mods:identifier>
</mods:mods>
<treatment id="03B087E55B38FFCEFF79F2BAFEB8F8F8" ID-DOI="http://doi.org/10.5281/zenodo.5291660" ID-Zenodo-Dep="5291660" LSID="urn:lsid:plazi:treatment:03B087E55B38FFCEFF79F2BAFEB8F8F8" httpUri="http://treatment.plazi.org/id/03B087E55B38FFCEFF79F2BAFEB8F8F8" lastPageNumber="94" pageId="93" pageNumber="94">
<subSubSection id="C30365785B38FFCEFF79F2BAFD9AFF27" box="[151,556,151,177]" pageId="93" pageNumber="94" type="nomenclature">
<paragraph id="8BA636F35B38FFCEFF79F2BAFD9AFF27" blockId="93.[151,556,151,210]" box="[151,556,151,177]" pageId="93" pageNumber="94">
<heading id="D0EE819F5B38FFCEFF79F2BAFD9AFF27" bold="true" box="[151,556,151,177]" fontSize="11" level="1" pageId="93" pageNumber="94" reason="1">
<emphasis id="B96DEAE15B38FFCEFF79F2BAFD9AFF27" bold="true" box="[151,556,151,177]" pageId="93" pageNumber="94">
<taxonomicName id="4C194D705B38FFCEFF79F2BAFE3BFF27" ID-CoL="3N3KH" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,397,151,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="variopaca" status="sp. nov.">
<emphasis id="B96DEAE15B38FFCEFF79F2BAFE3BFF27" bold="true" box="[151,397,151,177]" italics="true" pageId="93" pageNumber="94">Hydraena variopaca</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B38FFCEFE72F2BAFD9AFF27" box="[412,556,151,177]" pageId="93" pageNumber="94" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B38FFCEFF79F297FE0CFF44" box="[151,442,186,210]" pageId="93" pageNumber="94" type="description">
<paragraph id="8BA636F35B38FFCEFF79F297FE0CFF44" blockId="93.[151,556,151,210]" box="[151,442,186,210]" pageId="93" pageNumber="94">
(
<figureCitation id="13222A765B38FFCEFF71F297FEBDFF44" box="[159,267,186,210]" captionStart="FIGURE 159" captionStartId="264.[151,250,1753,1776]" captionTargetBox="[228,1368,220,1724]" captionTargetId="figure-29@264.[206,1380,193,1732]" captionTargetPageId="264" captionText="FIGURE 159. Hydraena variopaca, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283716" httpUri="https://zenodo.org/record/5283716/files/figure.png" pageId="93" pageNumber="94">Figs. 159</figureCitation>
,
<figureCitation id="13222A765B38FFCEFEF8F297FEF4FF44" box="[278,322,186,210]" captionStart="FIGURES 160–161" captionStartId="265.[151,263,1650,1673]" captionTargetBox="[409,1175,202,1617]" captionTargetId="figure-36@265.[385,1202,193,1629]" captionTargetPageId="265" captionText="FIGURES 160–161. Aedeagi. 160. H. cristatigena, non-type specimen from E. Highlands, Wanatabe Valley, nr. Okapa. 161. H. variopaca, holotype." figureDoi="http://doi.org/10.5281/zenodo.5283720" httpUri="https://zenodo.org/record/5283720/files/figure.png" pageId="93" pageNumber="94">161</figureCitation>
,
<figureCitation id="13222A765B38FFCEFEA0F297FECCFF44" box="[334,378,186,210]" captionStart="FIGURES 340–345" captionStartId="361.[151,263,1650,1673]" captionTargetBox="[178,1409,194,1629]" captionTargetId="figure-30@361.[178,1409,193,1629]" captionTargetPageId="361" captionText="FIGURES 340–345. Scanning electron micrographs of H. bacchusi (340–341), H. variopaca (342) and H. cristatigena (343– 345)." figureDoi="http://doi.org/10.5281/zenodo.5283999" httpUri="https://zenodo.org/record/5283999/files/figure.png" pageId="93" pageNumber="94">342</figureCitation>
,
<figureCitation id="13222A765B38FFCEFE68F297FE07FF44" box="[390,433,186,210]" captionStart="FIGURES 503–506" captionStartId="401.[151,264,1650,1673]" captionTargetBox="[165,1421,222,1609]" captionTargetId="figure-20@401.[165,1421,193,1629]" captionTargetPageId="401" captionText="FIGURES 503–506. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5299476" httpUri="https://zenodo.org/record/5299476/files/figure.png" pageId="93" pageNumber="94">505</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B38FFCEFF79F32EFAA2FD78" pageId="93" pageNumber="94" type="materials_examined">
<paragraph id="8BA636F35B38FFCEFF79F32EFAA2FD78" blockId="93.[151,1436,258,1902]" pageId="93" pageNumber="94">
Type Material.
<materialsCitation id="3B713CAE5B38FFCEFEA1F32EFAF8FEA8" collectingDate="1965-02-05" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1372" latitude="-6.5333333" location="Okapa" longLatPrecision="1304" longitude="145.61667" municipality="Wanitabi Valley" pageId="93" pageNumber="94" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="holotype">
<typeStatus id="54A288515B38FFCEFEA1F32EFE0AFE8D" box="[335,444,259,283]" pageId="93" pageNumber="94" type="holotype">
<emphasis id="B96DEAE15B38FFCEFEA1F32EFE0AFE8D" bold="true" box="[335,444,259,283]" pageId="93" pageNumber="94">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B38FFCEFDCCF32EFCCCFE8D" bold="true" box="[546,890,258,283]" pageId="93" pageNumber="94">
<collectingRegion id="49DDF8115B38FFCEFDCCF32EFCC0FE8D" box="[546,886,258,283]" country="Papua New Guinea" name="Eastern Highlands" pageId="93" pageNumber="94">Eastern Highlands Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BC2AC895B38FFCEFC67F32FFB8AFE8C" box="[905,1084,258,282]" pageId="93" pageNumber="94">Wanitabi Valley</collectingMunicipality>
, nr.
<location id="8EC660285B38FFCEFB85F32FFB01FE8C" LSID="urn:lsid:plazi:treatment:03B087E55B38FFCEFF79F2BAFEB8F8F8:8EC660285B38FFCEFB85F32FFB01FE8C" box="[1131,1207,258,282]" latitude="-6.5333333" longLatPrecision="1304" longitude="145.61667" municipality="Wanitabi Valley" name="Okapa" pageId="93" pageNumber="94" stateProvince="Eastern Highlands">Okapa</location>
, on wet rocks above waterline in small swift river,
<quantity id="4CE19B165B38FFCEFE02F30BFDF5FEA8" box="[492,579,294,318]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="93" pageNumber="94" unit="m" value="1372.0">
<elevation id="0034D1C05B38FFCEFE02F30BFDF5FEA8" box="[492,579,294,318]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="93" pageNumber="94" unit="m" value="1372.0">1372 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B38FFCEFDBEF30BFD1DFEA8" box="[592,683,294,318]" degrees="6" direction="south" minutes="32" orientation="latitude" pageId="93" pageNumber="94" precision="925" value="-6.5333333">6° 32' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B38FFCEFD57F30BFC87FEA8" box="[697,817,294,318]" degrees="145" direction="east" minutes="37" orientation="longitude" pageId="93" pageNumber="94" precision="925" value="145.61667">145° 37' E</geoCoordinate>
,
<date id="FFA710335B38FFCEFCD0F30BFC13FEA8" box="[830,933,294,318]" pageId="93" pageNumber="94" value="1965-02-05">
<collectingDate id="EFE3E9DB5B38FFCEFCD0F30BFC13FEA8" box="[830,933,294,318]" pageId="93" pageNumber="94" value="1965-02-05">5 ii 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B38FFCEFC5CF30AFBEFFEA8" box="[946,1113,294,318]" pageId="93" pageNumber="94">M. E. Bacchus</collectorName>
(MEB 171c) (
<collectionCode id="ED08AE365B38FFCEFB10F30BFAF0FEA8" box="[1278,1350,294,318]" pageId="93" pageNumber="94">NHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5B38FFCEFAB7F30AFCF0FEF4" collectorName="Awande &amp; Okapa" location="Eastern Highlands Province" pageId="93" pageNumber="94" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="paratype">
<emphasis id="B96DEAE15B38FFCEFAB7F30AFF63FEF5" bold="true" pageId="93" pageNumber="94">
<typeStatus id="54A288515B38FFCEFAB7F30AFF63FEF5" pageId="93" pageNumber="94" type="paratype">Paratypes</typeStatus>
</emphasis>
(131):
<emphasis id="B96DEAE15B38FFCEFEC3F366FD32FEF5" bold="true" box="[301,644,330,355]" pageId="93" pageNumber="94">
<collectingRegion id="49DDF8115B38FFCEFEC3F366FDC9FEF5" box="[301,639,330,355]" country="Papua New Guinea" name="Eastern Highlands" pageId="93" pageNumber="94">Eastern Highlands Province</collectingRegion>
:
</emphasis>
<collectorName id="26EC53255B38FFCEFD7CF367FD59FEF4" box="[658,751,330,354]" pageId="93" pageNumber="94">Awande</collectorName>
,
<collectorName id="26EC53255B38FFCEFD14F367FCF0FEF4" box="[762,838,330,354]" pageId="93" pageNumber="94">Okapa</collectorName>
</materialsCitation>
,
<materialsCitation id="3B713CAE5B38FFCEFCBEF366FCBCFE10" collectingDate="1965-05-25" collectionCode="NMNZ" collectorName="R. W. Hornabrook" elevation="1600" latitude="-6.5166664" location="Eastern Highlands" longLatPrecision="1304" longitude="145.58333" pageId="93" pageNumber="94" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="paratype">
<collectingRegion id="49DDF8115B38FFCEFCBEF366FB97FEF4" box="[848,1057,330,354]" country="Papua New Guinea" name="Eastern Highlands" pageId="93" pageNumber="94">Eastern Highlands</collectingRegion>
, [GE est.]
<quantity id="4CE19B165B38FFCEFB78F367FA85FEF4" box="[1174,1331,330,354]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="93" pageNumber="94" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">
<elevation id="0034D1C05B38FFCEFB78F367FA85FEF4" box="[1174,1331,330,354]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="93" pageNumber="94" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">1500–1700 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B38FFCEFAD0F367FA20FEF4" box="[1342,1430,330,354]" degrees="6" direction="south" minutes="31" orientation="latitude" pageId="93" pageNumber="94" precision="925" value="-6.5166664">6° 31' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B38FFCEFF79F343FEA2FE10" box="[151,276,366,390]" degrees="145" direction="east" minutes="35" orientation="longitude" pageId="93" pageNumber="94" precision="925" value="145.58333">145° 35' E</geoCoordinate>
,
<date id="FFA710335B38FFCEFECDF343FE2AFE10" box="[291,412,366,390]" pageId="93" pageNumber="94" value="1965-05-25">
<collectingDate id="EFE3E9DB5B38FFCEFECDF343FE2AFE10" box="[291,412,366,390]" pageId="93" pageNumber="94" value="1965-05-25">25 v 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B38FFCEFE45F342FDC8FE10" box="[427,638,366,390]" pageId="93" pageNumber="94">R. W. Hornabrook</collectorName>
(3
<collectionCode id="ED08AE365B38FFCEFD44F343FCB7FE10" box="[682,769,366,390]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="93" pageNumber="94" type="Museum">NMNZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B38FFCEFCF6F343FBA3FE3C" collectingDate="1965-03-17" collectionCode="NMNZ" collectorName="R. W. Hornabrook" elevation="1600" latitude="-6.5333333" location="Okapa" longLatPrecision="1304" longitude="145.61667" municipality="Mjairasa" pageId="93" pageNumber="94" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="paratype">
<collectingMunicipality id="6BC2AC895B38FFCEFCF6F343FCCBFE10" box="[792,893,366,390]" pageId="93" pageNumber="94">Mjairasa</collectingMunicipality>
,
<location id="8EC660285B38FFCEFC65F343FC61FE10" LSID="urn:lsid:plazi:treatment:03B087E55B38FFCEFF79F2BAFEB8F8F8:8EC660285B38FFCEFC65F343FC61FE10" box="[907,983,366,390]" latitude="-6.5333333" longLatPrecision="1304" longitude="145.61667" municipality="Mjairasa" name="Okapa" pageId="93" pageNumber="94" stateProvince="Eastern Highlands">Okapa</location>
, ex stones in waterfall spray, [GE est.]
<quantity id="4CE19B165B38FFCEFF79F3BFFE82FE3C" box="[151,308,402,426]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="93" pageNumber="94" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">
<elevation id="0034D1C05B38FFCEFF79F3BFFE82FE3C" box="[151,308,402,426]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="93" pageNumber="94" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">1500–1700 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B38FFCEFEAEF3BFFE2CFE3C" box="[320,410,402,426]" degrees="6" direction="south" minutes="32" orientation="latitude" pageId="93" pageNumber="94" precision="925" value="-6.5333333">6° 32' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B38FFCEFE48F3BFFDA9FE3C" box="[422,543,402,426]" degrees="145" direction="east" minutes="37" orientation="longitude" pageId="93" pageNumber="94" precision="925" value="145.61667">145° 37' E</geoCoordinate>
,
<date id="FFA710335B38FFCEFDC5F3BFFD11FE3C" box="[555,679,402,426]" pageId="93" pageNumber="94" value="1965-03-17">
<collectingDate id="EFE3E9DB5B38FFCEFDC5F3BFFD11FE3C" box="[555,679,402,426]" pageId="93" pageNumber="94" value="1965-03-17">17 iii 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B38FFCEFD5DF3BEFC37FE3C" box="[691,897,402,426]" pageId="93" pageNumber="94">R. W. Hornabrook</collectorName>
(40
<collectionCode id="ED08AE365B38FFCEFC5BF3BFFBBBFE3C" box="[949,1037,402,426]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="93" pageNumber="94" type="Museum">NMNZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B38FFCEFBCEF3BFFB0FFE58" collectingDate="1965-02-09" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1524" latitude="-6.5333333" location="Okapa" longLatPrecision="1304" longitude="145.61667" pageId="93" pageNumber="94" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="paratype">
<location id="8EC660285B38FFCEFBCEF3BFFBDAFE3C" LSID="urn:lsid:plazi:treatment:03B087E55B38FFCEFF79F2BAFEB8F8F8:8EC660285B38FFCEFBCEF3BFFBDAFE3C" box="[1056,1132,402,426]" latitude="-6.5333333" longLatPrecision="1304" longitude="145.61667" name="Okapa" pageId="93" pageNumber="94" stateProvince="Eastern Highlands">Okapa</location>
, ex muddy gravel banks of fast, clear stream,
<quantity id="4CE19B165B38FFCEFE8CF39BFE0FFE58" box="[354,441,438,462]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="93" pageNumber="94" unit="m" value="1524.0">
<elevation id="0034D1C05B38FFCEFE8CF39BFE0FFE58" box="[354,441,438,462]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="93" pageNumber="94" unit="m" value="1524.0">1524 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B38FFCEFE2BF39BFDABFE58" box="[453,541,438,462]" degrees="6" direction="south" minutes="32" orientation="latitude" pageId="93" pageNumber="94" precision="925" value="-6.5333333">6° 32' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B38FFCEFDC7F39BFD16FE58" box="[553,672,438,462]" degrees="145" direction="east" minutes="37" orientation="longitude" pageId="93" pageNumber="94" precision="925" value="145.61667">145° 37' E</geoCoordinate>
,
<date id="FFA710335B38FFCEFD45F39BFCA6FE58" box="[683,784,438,462]" pageId="93" pageNumber="94" value="1965-02-09">
<collectingDate id="EFE3E9DB5B38FFCEFD45F39BFCA6FE58" box="[683,784,438,462]" pageId="93" pageNumber="94" value="1965-02-09">9 ii 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B38FFCEFCF2F39AFC76FE58" box="[796,960,438,462]" pageId="93" pageNumber="94">M. E. Bacchus</collectorName>
(MEB 183) (2
<collectionCode id="ED08AE365B38FFCEFB85F39BFB07FE58" box="[1131,1201,438,462]" pageId="93" pageNumber="94">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B38FFCEFB2AF39BFA2EFE64" collectingDate="1965-02-10" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1524" latitude="-6.5333333" location="Okapa" longLatPrecision="1304" longitude="145.61667" pageId="93" pageNumber="94" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="paratype">
<location id="8EC660285B38FFCEFB2AF39BFAA6FE58" LSID="urn:lsid:plazi:treatment:03B087E55B38FFCEFF79F2BAFEB8F8F8:8EC660285B38FFCEFB2AF39BFAA6FE58" box="[1220,1296,438,462]" latitude="-6.5333333" longLatPrecision="1304" longitude="145.61667" name="Okapa" pageId="93" pageNumber="94" stateProvince="Eastern Highlands">Okapa</location>
, on logs and vegetable debris in small river,
<quantity id="4CE19B165B38FFCEFE13F3F7FDE3FE64" box="[509,597,474,498]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="93" pageNumber="94" unit="m" value="1524.0">
<elevation id="0034D1C05B38FFCEFE13F3F7FDE3FE64" box="[509,597,474,498]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="93" pageNumber="94" unit="m" value="1524.0">1524 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B38FFCEFD8CF3F7FD08FE64" box="[610,702,474,498]" degrees="6" direction="south" minutes="32" orientation="latitude" pageId="93" pageNumber="94" precision="925" value="-6.5333333">6° 32' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B38FFCEFD22F3F7FCF0FE64" box="[716,838,474,498]" degrees="145" direction="east" minutes="37" orientation="longitude" pageId="93" pageNumber="94" precision="925" value="145.61667">145° 37' E</geoCoordinate>
,
<date id="FFA710335B38FFCEFCBDF3F7FC7FFE64" box="[851,969,474,498]" pageId="93" pageNumber="94" value="1965-02-10">
<collectingDate id="EFE3E9DB5B38FFCEFCBDF3F7FC7FFE64" box="[851,969,474,498]" pageId="93" pageNumber="94" value="1965-02-10">10 ii 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B38FFCEFC38F3F6FBC8FE64" box="[982,1150,474,498]" pageId="93" pageNumber="94">M. E. Bacchus</collectorName>
(MEB 187a) (16
<collectionCode id="ED08AE365B38FFCEFAA7F3F7FA39FE64" box="[1353,1423,474,498]" pageId="93" pageNumber="94">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B38FFCEFF79F3D3FE1EFDAC" collectingDate="1965-02-09" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1524" latitude="-6.5333333" location="Okapa" longLatPrecision="1304" longitude="145.61667" pageId="93" pageNumber="94" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="paratype">
<location id="8EC660285B38FFCEFF79F3D3FF55FD80" LSID="urn:lsid:plazi:treatment:03B087E55B38FFCEFF79F2BAFEB8F8F8:8EC660285B38FFCEFF79F3D3FF55FD80" box="[151,227,510,534]" latitude="-6.5333333" longLatPrecision="1304" longitude="145.61667" name="Okapa" pageId="93" pageNumber="94" stateProvince="Eastern Highlands">Okapa</location>
, on wet rocks above waterline, fast, clear stream,
<quantity id="4CE19B165B38FFCEFCCFF3D3FCCCFD80" box="[801,890,510,534]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="93" pageNumber="94" unit="m" value="1524.0">
<elevation id="0034D1C05B38FFCEFCCFF3D3FCCCFD80" box="[801,890,510,534]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="93" pageNumber="94" unit="m" value="1524.0">1524 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B38FFCEFC66F3D3FC53FD80" box="[904,997,510,534]" degrees="6" direction="south" minutes="32" orientation="latitude" pageId="93" pageNumber="94" precision="925" value="-6.5333333">6° 32' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B38FFCEFC1DF3D3FBD8FD80" box="[1011,1134,510,534]" degrees="145" direction="east" minutes="37" orientation="longitude" pageId="93" pageNumber="94" precision="925" value="145.61667">145° 37' E</geoCoordinate>
,
<date id="FFA710335B38FFCEFB92F3D3FB50FD80" box="[1148,1254,510,534]" pageId="93" pageNumber="94" value="1965-02-09">
<collectingDate id="EFE3E9DB5B38FFCEFB92F3D3FB50FD80" box="[1148,1254,510,534]" pageId="93" pageNumber="94" value="1965-02-09">9 ii 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B38FFCEFB1DF3D2FA2AFD80" box="[1267,1436,510,534]" pageId="93" pageNumber="94">M. E. Bacchus</collectorName>
(MEB 183a) (13
<collectionCode id="ED08AE365B38FFCEFEB4F00FFE16FDAC" box="[346,416,546,570]" pageId="93" pageNumber="94">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B38FFCEFE5AF00FFE63FDC8" collectingDate="1965-02-10" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1524" latitude="-6.5333333" location="Okapa" longLatPrecision="1304" longitude="145.61667" pageId="93" pageNumber="94" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="paratype">
<location id="8EC660285B38FFCEFE5AF00FFDB6FDAC" LSID="urn:lsid:plazi:treatment:03B087E55B38FFCEFF79F2BAFEB8F8F8:8EC660285B38FFCEFE5AF00FFDB6FDAC" box="[436,512,546,570]" latitude="-6.5333333" longLatPrecision="1304" longitude="145.61667" name="Okapa" pageId="93" pageNumber="94" stateProvince="Eastern Highlands">Okapa</location>
, under stones in small river,
<quantity id="4CE19B165B38FFCEFCA6F00FFC16FDAC" box="[840,928,546,570]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="93" pageNumber="94" unit="m" value="1524.0">
<elevation id="0034D1C05B38FFCEFCA6F00FFC16FDAC" box="[840,928,546,570]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="93" pageNumber="94" unit="m" value="1524.0">1524 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B38FFCEFC43F00FFBBEFDAC" box="[941,1032,546,570]" degrees="6" direction="south" minutes="32" orientation="latitude" pageId="93" pageNumber="94" precision="925" value="-6.5333333">6° 32' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B38FFCEFBF8F00FFB39FDAC" box="[1046,1167,546,570]" degrees="145" direction="east" minutes="37" orientation="longitude" pageId="93" pageNumber="94" precision="925" value="145.61667">145° 37' E</geoCoordinate>
,
<date id="FFA710335B38FFCEFB72F00FFAA4FDAC" box="[1180,1298,546,570]" pageId="93" pageNumber="94" value="1965-02-10">
<collectingDate id="EFE3E9DB5B38FFCEFB72F00FFAA4FDAC" box="[1180,1298,546,570]" pageId="93" pageNumber="94" value="1965-02-10">10 ii 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B38FFCEFAF1F00EFF7CFDC8" pageId="93" pageNumber="94">M. E. Bacchus</collectorName>
(MEB 186) (12
<collectionCode id="ED08AE365B38FFCEFE69F06BFE7BFDC8" box="[391,461,582,606]" pageId="93" pageNumber="94">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B38FFCEFE0EF06AFB05FDC8" box="[480,1203,582,606]" collectingDate="1965-02-10" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1524" latitude="-6.5333333" location="Okapa" longLatPrecision="1304" longitude="145.61667" pageId="93" pageNumber="94" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="paratype">
same locality,
<date id="FFA710335B38FFCEFD6CF06BFD41FDC8" box="[642,759,582,606]" pageId="93" pageNumber="94" value="1965-02-10">
<collectingDate id="EFE3E9DB5B38FFCEFD6CF06BFD41FDC8" box="[642,759,582,606]" pageId="93" pageNumber="94" value="1965-02-10">10 ii 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B38FFCEFCEDF06AFC1FFDC8" box="[771,937,582,606]" pageId="93" pageNumber="94">M. E. Bacchus</collectorName>
(MEB 187) (26
<collectionCode id="ED08AE365B38FFCEFB8BF06BFB1DFDC8" box="[1125,1195,582,606]" pageId="93" pageNumber="94">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B38FFCEFB51F06BFE10FD30" collectingDate="1965-02-05" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1372" latitude="-6.5333333" location="Okapa" longLatPrecision="1304" longitude="145.61667" municipality="Wanitabi Valley" pageId="93" pageNumber="94" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="paratype">
<collectingMunicipality id="6BC2AC895B38FFCEFB51F06BFAC2FDC8" box="[1215,1396,582,606]" pageId="93" pageNumber="94">Wanitabi Valley</collectingMunicipality>
, nr.
<location id="8EC660285B38FFCEFF79F047FF55FD14" LSID="urn:lsid:plazi:treatment:03B087E55B38FFCEFF79F2BAFEB8F8F8:8EC660285B38FFCEFF79F047FF55FD14" box="[151,227,618,642]" latitude="-6.5333333" longLatPrecision="1304" longitude="145.61667" municipality="Wanitabi Valley" name="Okapa" pageId="93" pageNumber="94" stateProvince="Eastern Highlands">Okapa</location>
, on wet rocks above waterline in small swift river,
<quantity id="4CE19B165B38FFCEFCC4F047FC34FD14" box="[810,898,618,642]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="93" pageNumber="94" unit="m" value="1372.0">
<elevation id="0034D1C05B38FFCEFCC4F047FC34FD14" box="[810,898,618,642]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="93" pageNumber="94" unit="m" value="1372.0">1372 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B38FFCEFC61F047FC5DFD14" box="[911,1003,618,642]" degrees="6" direction="south" minutes="32" orientation="latitude" pageId="93" pageNumber="94" precision="925" value="-6.5333333">6° 32' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B38FFCEFC17F047FBC4FD14" box="[1017,1138,618,642]" degrees="145" direction="east" minutes="37" orientation="longitude" pageId="93" pageNumber="94" precision="925" value="145.61667">145° 37' E</geoCoordinate>
,
<date id="FFA710335B38FFCEFB91F047FB51FD14" box="[1151,1255,618,642]" pageId="93" pageNumber="94" value="1965-02-05">
<collectingDate id="EFE3E9DB5B38FFCEFB91F047FB51FD14" box="[1151,1255,618,642]" pageId="93" pageNumber="94" value="1965-02-05">5 ii 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B38FFCEFB1BF046FA2AFD14" box="[1269,1436,618,642]" pageId="93" pageNumber="94">M. E. Bacchus</collectorName>
(MEB 171c) (15
<collectionCode id="ED08AE365B38FFCEFEB6F0A3FE28FD30" box="[344,414,654,678]" pageId="93" pageNumber="94">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B38FFCEFE5CF0A3FD83FD5C" collectingDate="2006-11-10" collectionCode="ZSM" collectorName="Balke &amp; Kinibel" country="Papua New Guinea" elevation="1000" latitude="-7.0599666" location="Ande" longLatPrecision="1" longitude="145.73958" municipality="Marawaka" pageId="93" pageNumber="94" specimenCount="1" stateProvince="Gulf" typeStatus="paratype">
<emphasis id="B96DEAE15B38FFCEFE5CF0A3FDD5FD31" bold="true" box="[434,611,654,679]" pageId="93" pageNumber="94">
<collectingRegion id="49DDF8115B38FFCEFE5CF0A3FDE8FD31" box="[434,606,654,679]" country="Papua New Guinea" name="Gulf" pageId="93" pageNumber="94">Gulf Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BC2AC895B38FFCEFD9FF0A3FD5CFD30" box="[625,746,654,678]" pageId="93" pageNumber="94">Marawaka</collectingMunicipality>
, near
<location id="8EC660285B38FFCEFCC0F0A3FCDBFD30" LSID="urn:lsid:plazi:treatment:03B087E55B38FFCEFF79F2BAFEB8F8F8:8EC660285B38FFCEFCC0F0A3FCDBFD30" box="[814,877,654,678]" country="Papua New Guinea" latitude="-7.0599666" longLatPrecision="1" longitude="145.73958" municipality="Marawaka" name="Ande" pageId="93" pageNumber="94" stateProvince="Gulf">Ande</location>
,
<quantity id="4CE19B165B38FFCEFC97F0A3FC66FD30" box="[889,976,654,678]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="93" pageNumber="94" unit="m" value="1000.0">
<elevation id="0034D1C05B38FFCEFC97F0A3FC66FD30" box="[889,976,654,678]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="93" pageNumber="94" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B38FFCEFC33F0A3FBECFD30" box="[989,1114,654,678]" degrees="7" direction="south" minutes="3.598" orientation="latitude" pageId="93" pageNumber="94" precision="1" value="-7.0599666">7° 3.598' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B38FFCEFB89F0A3FAA6FD30" box="[1127,1296,654,678]" degrees="145" direction="east" minutes="44.375" orientation="longitude" pageId="93" pageNumber="94" precision="1" value="145.73958">145° 44.375' E</geoCoordinate>
,
<date id="FFA710335B38FFCEFAF2F0A3FA2EFD30" box="[1308,1432,654,678]" pageId="93" pageNumber="94" value="2006-11-10">
<collectingDate id="EFE3E9DB5B38FFCEFAF2F0A3FA2EFD30" box="[1308,1432,654,678]" pageId="93" pageNumber="94" value="2006-11-10">10 xi 2006</collectingDate>
</date>
,
<collectorName id="26EC53255B38FFCEFF79F09FFF6EFD5C" box="[151,216,690,714]" pageId="93" pageNumber="94">Balke</collectorName>
&amp;
<collectorName id="26EC53255B38FFCEFF13F09FFEE7FD5C" box="[253,337,690,714]" pageId="93" pageNumber="94">Kinibel</collectorName>
(
<collectingCountry id="F30E76635B38FFCEFE8FF09FFE2DFD5C" box="[353,411,690,714]" name="Papua New Guinea" pageId="93" pageNumber="94">PNG</collectingCountry>
89) (1
<collectionCode id="ED08AE365B38FFCEFE01F09FFD9BFD5C" box="[495,557,690,714]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="93" pageNumber="94" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B38FFCEFDAFF09EFAA6FD78" collectingDate="1964-11-07" collectionCode="NHM" collectorName="M. E. Bacchus" country="Papua New Guinea" county="Finisterre Mts." elevation="1432" latitude="-5.8333335" location="Moro" longLatPrecision="1305" longitude="146.11667" municipality="Naho River Valley" pageId="93" pageNumber="94" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<emphasis id="B96DEAE15B38FFCEFDAFF09EFCA9FD5D" bold="true" box="[577,799,690,715]" pageId="93" pageNumber="94">
<collectingRegion id="49DDF8115B38FFCEFDAFF09EFCACFD5D" box="[577,794,690,715]" country="Papua New Guinea" name="Madang" pageId="93" pageNumber="94">Madang Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62C74E7F5B38FFCEFCC1F09FFC61FD5C" box="[815,983,690,714]" pageId="93" pageNumber="94">Finisterre Mts.</collectingCounty>
,
<collectingMunicipality id="6BC2AC895B38FFCEFC0DF09FFB00FD5C" box="[995,1206,690,714]" pageId="93" pageNumber="94">Naho River Valley</collectingMunicipality>
, nr.
<location id="8EC660285B38FFCEFB09F09EFA9EFD5C" LSID="urn:lsid:plazi:treatment:03B087E55B38FFCEFF79F2BAFEB8F8F8:8EC660285B38FFCEFB09F09EFA9EFD5C" box="[1255,1320,691,714]" country="Papua New Guinea" county="Finisterre Mts." latitude="-5.8333335" longLatPrecision="1305" longitude="146.11667" municipality="Naho River Valley" name="Moro" pageId="93" pageNumber="94" stateProvince="Madang">Moro</location>
, ex gravel banks of small fast stream,
<quantity id="4CE19B165B38FFCEFE27F0FBFD96FD78" box="[457,544,726,750]" metricMagnitude="3" metricUnit="m" metricValue="1.432" pageId="93" pageNumber="94" unit="m" value="1432.0">
<elevation id="0034D1C05B38FFCEFE27F0FBFD96FD78" box="[457,544,726,750]" metricMagnitude="3" metricUnit="m" metricValue="1.432" pageId="93" pageNumber="94" unit="m" value="1432.0">1432 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B38FFCEFDC2F0FBFD33FD78" box="[556,645,726,750]" degrees="5" direction="south" minutes="50" orientation="latitude" pageId="93" pageNumber="94" precision="925" value="-5.8333335">5° 50' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B38FFCEFD7CF0FBFD4DFD78" box="[658,763,726,750]" degrees="146" direction="east" minutes="7" orientation="longitude" pageId="93" pageNumber="94" precision="925" value="146.11667">146° 7' E</geoCoordinate>
,
<date id="FFA710335B38FFCEFCE9F0FAFCC5FD78" box="[775,883,726,750]" pageId="93" pageNumber="94" value="1964-11-07">
<collectingDate id="EFE3E9DB5B38FFCEFCE9F0FAFCC5FD78" box="[775,883,726,750]" pageId="93" pageNumber="94" value="1964-11-07">7 xi 1964</collectingDate>
</date>
,
<collectorName id="26EC53255B38FFCEFC90F0FAFB95FD78" box="[894,1059,726,750]" pageId="93" pageNumber="94">M. E. Bacchus</collectorName>
(MEB 91) (3
<collectionCode id="ED08AE365B38FFCEFB2CF0FBFABEFD78" box="[1218,1288,726,750]" pageId="93" pageNumber="94">NHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B38FFCEFF29F0D7FE46FC50" pageId="93" pageNumber="94" type="diagnosis">
<paragraph id="8BA636F35B38FFCEFF29F0D7FE46FC50" blockId="93.[151,1436,258,1902]" pageId="93" pageNumber="94">
<emphasis id="B96DEAE15B38FFCEFF29F0D7FE65FC85" bold="true" box="[199,467,762,787]" pageId="93" pageNumber="94">Differential Diagnosis.</emphasis>
Among members of the
<emphasis id="B96DEAE15B38FFCEFD03F0D6FCCFFC85" box="[749,889,763,787]" italics="true" pageId="93" pageNumber="94">Cristatigena</emphasis>
group, quite similar in dorsal habitus to
<taxonomicName id="4C194D705B38FFCEFAABF0D6FF45FCA0" authorityName="Perkins" authorityYear="2011" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="cristatigena">
<emphasis id="B96DEAE15B38FFCEFAABF0D6FAEFFC84" box="[1349,1369,763,786]" italics="true" pageId="93" pageNumber="94">H</emphasis>
.
<emphasis id="B96DEAE15B38FFCEFA87F0D6FF45FCA0" italics="true" pageId="93" pageNumber="94">cristatigena</emphasis>
</taxonomicName>
(
<figureCitation id="13222A765B38FFCEFEEDF133FEC6FCA0" box="[259,368,798,822]" captionStart="FIGURE 158" captionStartId="263.[151,250,1644,1667]" captionTargetBox="[200,1393,215,1608]" captionTargetId="figure-26@263.[186,1401,193,1623]" captionTargetPageId="263" captionText="FIGURE 158. Hydraena cristatigena, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283712" httpUri="https://zenodo.org/record/5283712/files/figure.png" pageId="93" pageNumber="94">Figs. 158</figureCitation>
,
<figureCitation id="13222A765B38FFCEFE93F133FE1CFCA0" box="[381,426,798,822]" captionStart="FIGURE 159" captionStartId="264.[151,250,1753,1776]" captionTargetBox="[228,1368,220,1724]" captionTargetId="figure-29@264.[206,1380,193,1732]" captionTargetPageId="264" captionText="FIGURE 159. Hydraena variopaca, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283716" httpUri="https://zenodo.org/record/5283716/files/figure.png" pageId="93" pageNumber="94">159</figureCitation>
); differing therefrom by the slightly larger size (ca. 1.88 vs.
<quantity id="4CE19B165B38FFCEFBB0F133FB71FCA0" box="[1118,1223,798,822]" metricMagnitude="-3" metricUnit="m" metricValue="1.81" pageId="93" pageNumber="94" unit="mm" value="1.81">1.81 mm</quantity>
), the slightly more widely spaced metaventral plaques, and the slightly coarser granulation of the elytra. Males are additionally recognized by the simple hind tibiae. The male genitalia of the two species differ in significant details, while in general shape suggest a rather close relationship (
<figureCitation id="13222A765B38FFCEFD85F1A7FCA7FC34" box="[619,785,906,930]" captionStart="FIGURES 160–161" captionStartId="265.[151,263,1650,1673]" captionTargetBox="[409,1175,202,1617]" captionTargetId="figure-36@265.[385,1202,193,1629]" captionTargetPageId="265" captionText="FIGURES 160–161. Aedeagi. 160. H. cristatigena, non-type specimen from E. Highlands, Wanatabe Valley, nr. Okapa. 161. H. variopaca, holotype." figureDoi="http://doi.org/10.5281/zenodo.5283720" httpUri="https://zenodo.org/record/5283720/files/figure.png" pageId="93" pageNumber="94">Figs. 160, 160</figureCitation>
). The ventral surfaces of the head and prothorax are illustrated with an SEM (
<figureCitation id="13222A765B38FFCEFE6FF183FE55FC50" box="[385,483,942,966]" captionStart="FIGURES 340–345" captionStartId="361.[151,263,1650,1673]" captionTargetBox="[178,1409,194,1629]" captionTargetId="figure-30@361.[178,1409,193,1629]" captionTargetPageId="361" captionText="FIGURES 340–345. Scanning electron micrographs of H. bacchusi (340–341), H. variopaca (342) and H. cristatigena (343– 345)." figureDoi="http://doi.org/10.5281/zenodo.5283999" httpUri="https://zenodo.org/record/5283999/files/figure.png" pageId="93" pageNumber="94">Fig. 342</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B38FFCEFF29F1FFFC0FF948" pageId="93" pageNumber="94" type="description">
<paragraph id="8BA636F35B38FFCEFF29F1FFFAD4FBA4" blockId="93.[151,1436,258,1902]" pageId="93" pageNumber="94">
<emphasis id="B96DEAE15B38FFCEFF29F1FFFEEEFC7D" bold="true" box="[199,344,978,1003]" inLineHeading="true" pageId="93" pageNumber="94" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B38FFCEFE42F1FFFDB8FC7C" box="[428,526,978,1002]" pageId="93" pageNumber="94" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.88/0.88; head 0.28/0.46; pronotum 0.43/0.58, PA 0.50, PB 0.48; elytra 1.17/0.88. Dorsum dark brown to piceous, head and pronotal disc slightly darker, palpi light brown, except distal 1/2 of last palpomere darker; legs brown, &quot;knees&quot; dark brown.
</paragraph>
<paragraph id="8BA636F35B38FFCEFF29F612FA98FB70" blockId="93.[151,1436,258,1902]" pageId="93" pageNumber="94">Dorsum of head and pronotum densely punctulate, dull; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 1xpd or slightly larger; punctures of clypeus obsolete. Labrum apicomedially excised, free margins weakly upturned. Mentum sparsely punctate, effacedly microreticulate, weakly shining; postmentum anterior 1/2 with punctation similar to mentum, posterior 1/2 densely micropunctulate. Genae raised, posterior ridge strong. Pronotum with anterior margin weakly emarginate over middle 1/2; all pronotal foveae moderately deep.</paragraph>
<paragraph id="8BA636F35B38FFCEFF29F6DFFCA9FAE0" blockId="93.[151,1436,258,1902]" pageId="93" pageNumber="94">Elytra subrugulose, dull, punctures very indistinct, most punctures with three granules, one at anterior margin and one on each side. Intervals not raised, width ca. 1xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another. Explanate margin well developed, about as wide as widest part of metatibia.</paragraph>
<paragraph id="8BA636F35B38FFCEFF29F7AFFCFEF9E4" blockId="93.[151,1436,258,1902]" pageId="93" pageNumber="94">Ratios of P2 width and plaque shape (P2/w/l/s) ca. 4/1/10/7. P1 laminate; median carina sinuate in profile. P2 moderately narrow, l/w ca. 2/1, sides parallel, apex blunt. Plaques very narrow, weakly raised, subcarinate, located at lateral margins of deep, median depression. Metaventrite tumid lateral to plaques. No midlongitudinal carina between mesoventral intercoxal process and plaques. Hydrofuge pubescence dense. AIS flat, width at straight posterior margin ca. 1.5x P2. Protibia (male) markedly widened, slightly arcuate, medial surface rather flat; mesotibia simple; metatibia slender, gradually increasing in width from base to apex, very weakly emarginate on medial surface slightly before midlength. Abdominal apex symmetrical.</paragraph>
<paragraph id="8BA636F35B38FFCEFF29F453FC0FF948" blockId="93.[151,1436,258,1902]" pageId="93" pageNumber="94">
Aedeagus (
<figureCitation id="13222A765B38FFCEFEA6F453FE1CF900" box="[328,426,1662,1686]" captionStart="FIGURES 160–161" captionStartId="265.[151,263,1650,1673]" captionTargetBox="[409,1175,202,1617]" captionTargetId="figure-36@265.[385,1202,193,1629]" captionTargetPageId="265" captionText="FIGURES 160–161. Aedeagi. 160. H. cristatigena, non-type specimen from E. Highlands, Wanatabe Valley, nr. Okapa. 161. H. variopaca, holotype." figureDoi="http://doi.org/10.5281/zenodo.5283720" httpUri="https://zenodo.org/record/5283720/files/figure.png" pageId="93" pageNumber="94">Fig. 161</figureCitation>
) main piece with large distal lobe, in ventral aspect lobe apically pointed, with arcuate right side and left side with tiny lobe; left paramere small, not bilobed, setae in two clusters; right paramere sharply rounded apically; distal piece, in right lateral view, with knob-like lobe.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B38FFCEFF29F4C6FCC8F894" box="[199,894,1770,1795]" pageId="93" pageNumber="94" type="etymology">
<paragraph id="8BA636F35B38FFCEFF29F4C6FCC8F894" blockId="93.[151,1436,258,1902]" box="[199,894,1770,1795]" pageId="93" pageNumber="94">
<emphasis id="B96DEAE15B38FFCEFF29F4C6FEFBF895" bold="true" box="[199,333,1771,1795]" inLineHeading="true" pageId="93" pageNumber="94" reason="1">Etymology.</emphasis>
Named in reference to the varyingly dull dorsum.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B38FFCEFF29F523FEB8F8F8" pageId="93" pageNumber="94" type="distribution">
<paragraph id="8BA636F35B38FFCEFF29F523FEB8F8F8" blockId="93.[151,1436,258,1902]" pageId="93" pageNumber="94">
<emphasis id="B96DEAE15B38FFCEFF29F523FED6F8B1" bold="true" box="[199,352,1806,1831]" inLineHeading="true" pageId="93" pageNumber="94" reason="1">Distribution.</emphasis>
Currently known from far eastern Area 1 (
<quantity id="4CE19B165B38FFCEFCACF523FC69F8B0" box="[834,991,1806,1830]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="93" pageNumber="94" unit="m" value="1350.0" valueMax="1700.0" valueMin="1000.0">1000–1700 m</quantity>
) and one locality in the Finisterre Mts., Area 5 (
<quantity id="4CE19B165B38FFCEFF1CF51FFEFFF8DC" box="[242,329,1842,1866]" metricMagnitude="3" metricUnit="m" metricValue="1.432" pageId="93" pageNumber="94" unit="m" value="1432.0">1432 m</quantity>
) (
<figureCitation id="13222A765B38FFCEFE8FF51FFE75F8DC" box="[353,451,1842,1866]" captionStart="FIGURES 503–506" captionStartId="401.[151,264,1650,1673]" captionTargetBox="[165,1421,222,1609]" captionTargetId="figure-20@401.[165,1421,193,1629]" captionTargetPageId="401" captionText="FIGURES 503–506. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5299476" httpUri="https://zenodo.org/record/5299476/files/figure.png" pageId="93" pageNumber="94">Fig. 505</figureCitation>
). This is one of ten species with distributions exclusively shared between Areas 1 and 5 (
<figureCitation id="13222A765B38FFCEFF71F57BFEB4F8F8" box="[159,258,1878,1902]" captionStart="FIGURES 421–422" captionStartId="378.[151,263,1752,1775]" captionTargetBox="[228,1326,203,1731]" captionTargetId="figure-46@378.[221,1367,193,1731]" captionTargetPageId="378" captionText="FIGURES 421–422. 421. All known collecting localities for Papua New Guinea Hydraena. 422. Ten species, representing eight species groups, with distributions exclusively shared between Areas of Endemism 1 and 5." figureDoi="http://doi.org/10.5281/zenodo.5284047" httpUri="https://zenodo.org/record/5284047/files/figure.png" pageId="93" pageNumber="94">Fig. 422</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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