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148 lines
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<document id="EDD54DDCC84092CA04852CF3E1C5DB57" ID-DOI="10.17161/jom.v0i46.4839" ID-ISSN="2325-4467" ID-Zenodo-Dep="13145840" ID-ZooBank="urn:lsid:zoobank.org:pub:0626B747-69EF-4C66-A9C5-A2EEB29187C1" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1722454560475" checkinUser="felipe" docAuthor="Engel, Michael S." docDate="2015" docId="03A7CE56FFFDFFC90AF7B795FCFEF9E2" docLanguage="en" docName="JMelittology.2015.46.1-7.pdf" docOrigin="Journal of Melittology 2015 (46)" docSource="http://dx.doi.org/10.17161/jom.v0i46.4839" docStyle="DocumentStyle:20C606B715423E98DFED429F4976A416.4:JMelittology.2013-.journal_article.0cover" docStyleId="20C606B715423E98DFED429F4976A416" docStyleName="JMelittology.2013-.journal_article.0cover" docStyleVersion="4" docTitle="Tarsaliini Engel, 2015, new tribe" docType="treatment" docUuid="7E511B3C-6119-4A89-823B-636F574CBA7B" docUuidSource="ZooBank" docVersion="3" lastPageNumber="4" masterDocId="FF9EB62EFFFEFFCA0B0DB554FFC1FFDB" masterDocTitle="Notes on family-group names for bees (Hymenoptera: Apoidea)" masterLastPageNumber="7" masterPageNumber="1" pageNumber="4" updateTime="1722612847461" updateUser="carolina" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:titleInfo id="7F5016694F5EA9CFB28031CBCB6B1674">
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<mods:title id="5D580B379BA22E97E8E1D7227FD1E4E5">Notes on family-group names for bees (Hymenoptera: Apoidea)</mods:title>
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<mods:name id="7150D759F178596C474889AB3A71E0E2" type="personal">
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<mods:namePart id="C71CF8676C74AFB57BE88A9B6555B6C2">Engel, Michael S.</mods:namePart>
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<mods:affiliation id="7FD62DD2C24E8063C8728D22938F33CF">Division of Entomology, Natural History Museum, and Department of Ecology & Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). & Division of Invertebrate Zoology, American Museum of Natural History, Central Park West</mods:affiliation>
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<mods:title id="99E27A12329C12E3DF94A9B1E69435D8">Journal of Melittology</mods:title>
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</mods:titleInfo>
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<mods:part id="28EE6D5A36B105025858251636178AF4">
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<mods:date id="7163AA6978A24C9AE4B69E61580F7D6C">2015</mods:date>
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<mods:number id="5C58CFE0766AFBEC92D166F5AB8BB21B">2015-03-13</mods:number>
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<mods:number id="20240395917D75F66579644FE63456C4">2015</mods:number>
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<mods:number id="9611737F46FD731576FC1AB34371F264">46</mods:number>
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<mods:url id="CFE914965AB5BD32531F8C6F7EB5A0E1">http://dx.doi.org/10.17161/jom.v0i46.4839</mods:url>
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<mods:identifier id="5E8C320A61D5DCB3FD7EBEEB699C4B1A" type="DOI">10.17161/jom.v0i46.4839</mods:identifier>
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<mods:identifier id="AB60A2331C86FDEEF8826999AC54E651" type="ISSN">2325-4467</mods:identifier>
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<subSubSection id="C3142CCBFFFDFFC90AF7B795FBF4FD22" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="8BB17F40FFFDFFC90AF7B795FCF7FD02" blockId="3.[251,1077,673,761]" box="[506,822,705,729]" pageId="3" pageNumber="4">
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<heading id="D0F9C82CFFFDFFC90AF7B795FCF7FD02" box="[506,822,705,729]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="4" reason="2">
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<taxonomicName id="4C0E04C3FFFDFFC90AF7B795FD7AFD02" ID-CoL="L9N" authorityName="Engel" authorityYear="2015" box="[506,699,705,729]" class="Insecta" family="Apidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Tarsaliini">
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<emphasis id="B97AA352FFFDFFC90AF7B795FDAFFD02" bold="true" box="[506,622,705,729]" pageId="3" pageNumber="4">Tarsaliini</emphasis>
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Engel
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2491E29FFFDFFC909CBB796FCF7FD02" box="[710,822,705,729]" pageId="3" pageNumber="4" rank="tribe">new tribe</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BB17F40FFFDFFC90BF6B7B7FBF4FD22" blockId="3.[251,1077,673,761]" box="[251,1077,739,761]" pageId="3" pageNumber="4">
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ZooBank:
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urn:lsid:zoobank.org:act:
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<uuid id="FFA84595FFFDFFC90956B7B7FBF4FD22" box="[603,1077,739,761]" pageId="3" pageNumber="4">7E511B3C-6119-4A89-823B-636F574CBA7B</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3142CCBFFFDFFC90BDDB676FDB9FCE2" box="[208,632,801,825]" pageId="3" pageNumber="4" type="materials_examined">
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<paragraph id="8BB17F40FFFDFFC90BDDB676FDB9FCE2" blockId="3.[160,1175,801,1593]" box="[208,632,801,825]" pageId="3" pageNumber="4">
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<materialsCitation id="3B66751DFFFDFFC90BDDB676FDB5FCE2" box="[208,628,801,825]" collectionCode="GENUS" pageId="3" pageNumber="4" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54B5C1E2FFFDFFC90BDDB676FEC7FCE2" box="[208,262,802,825]" pageId="3" pageNumber="4">TYPE</typeStatus>
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<collectionCode id="ED1FE785FFFDFFC90A00B675FE96FCE2" box="[269,343,801,825]" pageId="3" pageNumber="4">GENUS</collectionCode>
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:
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<taxonomicName id="4C0E04C3FFFDFFC90A6DB675FDB5FCE2" authority="Morawitz, 1895" authorityName="Morawitz" authorityYear="1895" box="[352,628,801,825]" class="Insecta" family="Apidae" genus="Tarsalia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AA352FFFDFFC90A6DB675FE74FCE2" box="[352,437,801,825]" italics="true" pageId="3" pageNumber="4">Tarsalia</emphasis>
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<bibRefCitation id="EF9F02B1FFFDFFC90AB1B675FDB5FCE2" author="Morawitz, F. F." box="[444,628,801,825]" pageId="3" pageNumber="4" pagination="1 - 76" refId="ref3193" refString="Morawitz, F. F. 1895. Beitrag zur Bienenfauna Turkmeniens. Horae Societatis Entomologicae Rossicae 29: 1 - 76." type="journal article" year="1895">Morawitz, 1895</bibRefCitation>
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</taxonomicName>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3142CCBFFFDFFC90BDDB615FC11FA62" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="8BB17F40FFFDFFC90BDDB615FE87FA42" blockId="3.[160,1175,801,1593]" pageId="3" pageNumber="4">
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DIAGNOSIS: Bees of moderate size (
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<quantity id="4CF6D2A5FFFDFFC90950B615FD09FC82" box="[605,712,833,857]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="3" pageNumber="4" unit="mm" value="10.0" valueMax="13.0" valueMin="7.0">7–13 mm</quantity>
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in length); clypeus often yellow, weakly to moderately protuberant in profile, strongly bent back at side of labrum; lower face often with yellow maculation; labiomaxillary complex with glossa and labial palpus longer than prementum; first two palpomeres of labial palpus long, sheathlike (in contrast to the shortened condition of
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<taxonomicName id="4C0E04C3FFFDFFC90968B695FD1AFC02" box="[613,731,961,985]" class="Insecta" family="Apidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Ancylaini">Ancylaini</taxonomicName>
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); male antennal flagellum not crenulate. Forewing with pterostigma small, as long as wide, scarcely wider than prestigma; pterostigmal margin inside marginal cell weakly or not convex; marginal cell apex not truncate, bent away from anterior wing margin; three submarginal cells present; hind wing with second abscisssa of M+Cu half as long as vein M or less [several
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<taxonomicName id="4C0E04C3FFFDFFC90F0AB115FB95FB82" authorityName="Lepeletier" authorityYear="1841" box="[1031,1108,1089,1113]" class="Insecta" family="Apidae" genus="Ancyla" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AA352FFFDFFC90F0AB115FB95FB82" box="[1031,1108,1089,1113]" italics="true" pageId="3" pageNumber="4">Ancyla</emphasis>
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</taxonomicName>
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Lepeletier de Saint Fargeau have this vein slightly more than half as long as vein M,
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<emphasis id="B97AA352FFFDFFC90F46B136FB51FBA2" box="[1099,1168,1122,1145]" italics="true" pageId="3" pageNumber="4">contra</emphasis>
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<bibRefCitation id="EF9F02B1FFFDFFC90BADB1D5FEB1FB42" author="Michener, C. D." box="[160,368,1153,1177]" pageId="3" pageNumber="4" refId="ref3110" refString="Michener, C. D. 2007. The Bees of the World [2 nd Edition]. Johns Hopkins University Press; Baltimore, MD; xvi + [i] + 953 pp., + 20 pls." type="book" year="2007">Michener’s (2007)</bibRefCitation>
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key to tribes of
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<taxonomicName id="4C0E04C3FFFDFFC90922B1D5FD4BFB42" box="[559,650,1153,1177]" class="Insecta" family="Apidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subFamily" subFamily="Apinae">Apinae</taxonomicName>
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]. Metatibial and metabasitibial scopa large, dense, composed of long, plumose setae; basal area of propodeum with subhorizontal area [particularly well developed in
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<taxonomicName id="4C0E04C3FFFDFFC90979B195FC59FB02" authority="(Warncke)" baseAuthorityName="Warncke" baseAuthorityYear="1979" box="[628,920,1217,1241]" class="Insecta" family="Apidae" genus="Tarsalia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="persica">
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<emphasis id="B97AA352FFFDFFC90979B195FCDDFB02" box="[628,796,1217,1241]" italics="true" pageId="3" pageNumber="4">Tarsalia persica</emphasis>
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(Warncke)
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</taxonomicName>
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], subhorizontal area punctate and setose (area poorly developed, impunctate, and asetose in
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<taxonomicName id="4C0E04C3FFFDFFC90F03B1B5FB4AFB22" box="[1038,1163,1249,1273]" class="Insecta" family="Apidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Ancylaini">Ancylaini</taxonomicName>
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). Metasomal tergum I with weak carina at angle of anterior-facing and dorsal-facing surfaces; female sterna II–
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<collectionCode id="ED1FE785FFFDFFC90AD1B076FE2EFAE2" box="[476,495,1314,1337]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="4" type="Museum">V</collectionCode>
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with scopa composed of dense setae; male tergum VII setose, medially produced and truncate to weakly bidentate, pygidial plate occupying most of dorsum of tergum, laterally defined by carina; male sternum VII and genitalia asymmetrical.
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</paragraph>
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<paragraph id="8BB17F40FFFDFFC90BDDB0F5FC11FA62" blockId="3.[160,1175,801,1593]" box="[208,976,1441,1466]" pageId="3" pageNumber="4">INCLUDED GENERA: Presently only the nominate genus is included.</paragraph>
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</subSubSection>
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<subSubSection id="C3142CCBFFFDFFC90BDDB095FCFEF9E2" pageId="3" pageNumber="4" type="discussion">
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<paragraph id="8BB17F40FFFDFFC90BDDB095FCFEF9E2" blockId="3.[160,1175,801,1593]" pageId="3" pageNumber="4">
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COMMENTS: The genus
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<taxonomicName id="4C0E04C3FFFDFFC90AD1B095FDF0FA02" authorityName="Morawitz" authorityYear="1895" box="[476,561,1473,1497]" class="Insecta" family="Apidae" genus="Tarsalia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AA352FFFDFFC90AD1B095FDF0FA02" box="[476,561,1473,1497]" italics="true" pageId="3" pageNumber="4">Tarsalia</emphasis>
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</taxonomicName>
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has long been grouped with
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<taxonomicName id="4C0E04C3FFFDFFC90884B095FC17FA02" authorityName="Lepeletier" authorityYear="1841" box="[905,982,1473,1497]" class="Insecta" family="Apidae" genus="Ancyla" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AA352FFFDFFC90884B095FC17FA02" box="[905,982,1473,1497]" italics="true" pageId="3" pageNumber="4">Ancyla</emphasis>
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</taxonomicName>
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in the tribe
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<taxonomicName id="4C0E04C3FFFDFFC90F6EB096FF2FFA22" class="Insecta" family="Apidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Ancylaini">Ancylaini</taxonomicName>
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(
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<emphasis id="B97AA352FFFDFFC90BF3B0B6FEDCFA22" box="[254,285,1506,1529]" italics="true" pageId="3" pageNumber="4">e.g</emphasis>
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.,
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<bibRefCitation id="EF9F02B1FFFDFFC90A3DB0B5FE27FA22" author="Michener, C. D." box="[304,486,1505,1529]" pageId="3" pageNumber="4" pagination="151 - 326" refId="ref3008" refString="Michener, C. D. 1944. Comparative external morphology, phylogeny, and a classification of the bees (Hymenoptera). Bulletin of the American Museum of Natural History 82 (6): 151 - 326." type="journal article" year="1944">Michener, 1944</bibRefCitation>
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,
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<bibRefCitation id="EF9F02B1FFFDFFC90AFDB0B5FDE9FA22" author="Michener, C. D." box="[496,552,1505,1529]" pageId="3" pageNumber="4" refId="ref3110" refString="Michener, C. D. 2007. The Bees of the World [2 nd Edition]. Johns Hopkins University Press; Baltimore, MD; xvi + [i] + 953 pp., + 20 pls." type="book" year="2007">2007</bibRefCitation>
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;
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<bibRefCitation id="EF9F02B1FFFDFFC9093EB0B5FD39FA22" author="Engel, M. S. & J. S. Ascher & D. A. Yanega" box="[563,760,1505,1529]" pageId="3" pageNumber="4" pagination="198 - 201" refId="ref2539" refString="Engel, M. S., J. S. Ascher, & D. A. Yanega. 2008. Ancylini Michener, 1944 (Insecta, Hymenoptera): Proposed emendation of spelling to Ancylaini, to remove homonymy with Ancylini Rafinesque, 1815 (Mollusca, Gastropoda). Bulletin of Zoological Nomenclature 65 (3): 198 - 201." type="journal article" year="2008">
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<taxonomicName id="4C0E04C3FFFDFFC9093EB0B5FDB7FA22" authorityName="Engel" authorityYear="2015" box="[563,630,1505,1529]" class="Insecta" family="Apidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Engel">Engel</taxonomicName>
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<emphasis id="B97AA352FFFDFFC90970B0B6FD6BFA22" box="[637,682,1505,1529]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
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., 2008
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</bibRefCitation>
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). However, the monophyly of this grouping is suspect (
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<bibRefCitation id="EF9F02B1FFFDFFC90AADB355FDEFF9C2" author="Baker, D. B." box="[416,558,1537,1561]" pageId="3" pageNumber="4" pagination="823 - 860" refId="ref2283" refString="Baker, D. B. 1998. Taxonomic and phylogenetic problems in Old World eucerine bees, with special reference to the genus Tarsalia Morawitz, 1895 (Hymenoptera: Apoidea: Anthophoridae). Journal of Natural History 32 (6): 823 - 860." type="journal article" year="1998">Baker, 1998</bibRefCitation>
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;
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<bibRefCitation id="EF9F02B1FFFDFFC90931B356FCF2F9C2" author="Praz, C. J. & L. Packer" box="[572,819,1537,1561]" pageId="3" pageNumber="4" pagination="258 - 270" refId="ref3287" refString="Praz, C. J., & L. Packer. 2014. Phylogenetic position of the bee genera Ancyla and Tarsalia (Hymenoptera: Apidae): A remarkable base compositional bias and an early Paleogene geodispersal from North America to the Old World. Molecular Phylogenetics and Evolution 81: 258 - 270." type="journal article" year="2014">Praz & Packer, 2014</bibRefCitation>
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), and it seems apparent that
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<taxonomicName id="4C0E04C3FFFDFFC90BADB375FF34F9E2" authorityName="Morawitz" authorityYear="1895" box="[160,245,1569,1593]" class="Insecta" family="Apidae" genus="Tarsalia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AA352FFFDFFC90BADB375FF34F9E2" box="[160,245,1569,1593]" italics="true" pageId="3" pageNumber="4">Tarsalia</emphasis>
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</taxonomicName>
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is more closely related to
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<taxonomicName id="4C0E04C3FFFDFFC90925B375FD4AF9E2" box="[552,651,1569,1593]" class="Insecta" family="Apidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Eucerini">Eucerini</taxonomicName>
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than to
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<taxonomicName id="4C0E04C3FFFDFFC909E6B375FCF9F9E2" authorityName="Lepeletier" authorityYear="1841" box="[747,824,1569,1593]" class="Insecta" family="Apidae" genus="Ancyla" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AA352FFFDFFC909E6B375FCF9F9E2" box="[747,824,1569,1593]" italics="true" pageId="3" pageNumber="4">Ancyla</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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