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<document id="EFFDB46C0C37035347A064EB39638019" ID-CLB-Dataset="28075" ID-DOI="10.11646/zootaxa.4504.3.6" ID-GBIF-Dataset="3de64436-3ff2-4d0a-8676-b0504170ec50" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606507" ID-ZooBank="E4F4A4FC-2AD0-4438-91D4-BB690C91A5A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553584873659" checkinUser="plazi" docAuthor="Carvalho-Filho, Fernando Da Silva, Gorayeb, Inocêncio De Sousa, Soares, Jéssica Maria Menezes &amp; Souza, Matheus Tavares De" docDate="2018" docId="039A5520B33BDB284EA0F8DD2BB1F83F" docLanguage="en" docName="zootaxa.4504.3.6.pdf" docOrigin="Zootaxa 4504 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Metopia fofo Carvalho-Filho &amp; Gorayeb &amp; Soares &amp; Souza 2018, sp. nov." docType="treatment" docVersion="8" lastPageNumber="409" masterDocId="FFA32D58B33CDB204E37FFA92823FFA2" masterDocTitle="Flesh flies (Diptera: Sarcophagidae) from a white-sand habitat in the Brazilian Amazon, with the description of four new species" masterLastPageNumber="417" masterPageNumber="401" pageNumber="408" updateTime="1698712787524" updateUser="ExternalLinkService">
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<mods:titleInfo id="614AAF4B1788BF84A8A73E2779509008">
<mods:title id="DCF13F07224D602C92ED9F4E3010C75D">Flesh flies (Diptera: Sarcophagidae) from a white-sand habitat in the Brazilian Amazon, with the description of four new species</mods:title>
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<mods:namePart id="3358CF5AF5992B3FF2ADD7E30C8D3539">Carvalho-Filho, Fernando Da Silva</mods:namePart>
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<mods:namePart id="CC723FC5EDCEDEBEC4E0A885B4677A08">Gorayeb, Inocêncio De Sousa</mods:namePart>
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<mods:namePart id="CC065FEF0FD71447EBBF6478B6D57B34">Soares, Jéssica Maria Menezes</mods:namePart>
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<mods:namePart id="7DE2E68098D2A0BDBDD4C3D5ECF854C8">Souza, Matheus Tavares De</mods:namePart>
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<mods:typeOfResource id="A56863994439F16C4C2191F2EB977B71">text</mods:typeOfResource>
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<mods:title id="97C7BDC02B0556B43C80184F3C769103">Zootaxa</mods:title>
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<mods:part id="F76596D5153B9B7E07AD4178F7779BA4">
<mods:date id="D06C646EF1D776A97A40AA927EE78CE1">2018</mods:date>
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<mods:number id="B852493BFFDEE9E8578A0AF2AF1190ED">2018-10-25</mods:number>
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<mods:number id="1BA07C72670B926FA3EB5C742DC289A8">4504</mods:number>
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<mods:detail id="2A8FD88ADC2B5BD735ABA95CD9D31B68" type="issue">
<mods:number id="A2E424649EE673ADBDA262E65F44A5E3">3</mods:number>
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<mods:classification id="B891DA3599C31862B9F25531B9DE5C2F">journal article</mods:classification>
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<mods:identifier id="6F982A76221A65F2D078E4ACC9F6B080" type="DOI">10.11646/zootaxa.4504.3.6</mods:identifier>
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<mods:identifier id="2E0215D0CD2A4DE29EA3DD7894987697" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039A5520B33BDB284EA0F8DD2BB1F83F" ID-DOI="http://doi.org/10.5281/zenodo.5968090" ID-GBIF-Taxon="154495096" ID-Zenodo-Dep="5968090" LSID="urn:lsid:plazi:treatment:039A5520B33BDB284EA0F8DD2BB1F83F" httpUri="http://treatment.plazi.org/id/039A5520B33BDB284EA0F8DD2BB1F83F" lastPageId="8" lastPageNumber="409" pageId="7" pageNumber="408">
<subSubSection id="C329B7BDB33BDB274EA0F8DD29B4F82C" box="[151,407,1908,1934]" pageId="7" pageNumber="408" type="nomenclature">
<paragraph id="8B8CE436B33BDB274EA0F8DD29B4F82C" blockId="7.[151,407,1908,1967]" box="[151,407,1908,1934]" pageId="7" pageNumber="408">
<heading id="D0C4535AB33BDB274EA0F8DD29B4F82C" bold="true" box="[151,407,1908,1934]" fontSize="11" level="1" pageId="7" pageNumber="408" reason="1">
<emphasis id="B9473824B33BDB274EA0F8DD29B4F82C" bold="true" box="[151,407,1908,1934]" pageId="7" pageNumber="408">
<taxonomicName id="4C339FB5B33BDB274EA0F8DD2911F82C" ID-CoL="8GTX3" authority="Carvalho-Filho &amp; Gorayeb &amp; Soares &amp; Souza, 2018" authorityName="Carvalho-Filho &amp; Gorayeb &amp; Soares &amp; Souza" authorityYear="2018" box="[151,306,1908,1934]" class="Insecta" family="Sarcophagidae" genus="Metopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="fofo" status="sp. nov.">
<emphasis id="B9473824B33BDB274EA0F8DD2911F82C" bold="true" box="[151,306,1908,1934]" italics="true" pageId="7" pageNumber="408">Metopia fofo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A274855FB33BDB274F0DF8DD29B4F82C" box="[314,407,1908,1934]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C329B7BDB33BDB274EA0F83F2904F80D" box="[151,295,1942,1967]" pageId="7" pageNumber="408" type="description">
<paragraph id="8B8CE436B33BDB274EA0F83F2904F80D" blockId="7.[151,407,1908,1967]" box="[151,295,1942,1967]" pageId="7" pageNumber="408">
(
<figureCitation id="1308F8B3B33BDB274EA8F83F293CF80D" box="[159,287,1942,1967]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="7" pageNumber="408">Figs 22–26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C329B7BDB33BDB284EA0F8702BADFEE2" lastPageId="8" lastPageNumber="409" pageId="7" pageNumber="408" type="materials_examined">
<paragraph id="8B8CE436B33BDB274EA0F8702DBFF853" blockId="7.[151,1436,2009,2034]" box="[151,1436,2009,2034]" pageId="7" pageNumber="408">
<emphasis id="B9473824B33BDB274EA0F8702966F850" bold="true" box="[151,325,2009,2034]" pageId="7" pageNumber="408">Type material.</emphasis>
<materialsCitation id="3B5BEE6BB33BDB274F7BF8732DBFF853" ID-GBIF-Occurrence="2028137927" box="[332,1436,2009,2034]" collectionCode="MPEG" collectorName="Km &amp; de Cameta" pageId="7" pageNumber="408" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54885A94B33BDB274F7BF87329C3F850" box="[332,480,2010,2034]" pageId="7" pageNumber="408" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D352FBFB33BDB274FDFF87329DEF853" box="[488,509,2010,2033]" pageId="7" pageNumber="408" type="male">♂</specimenCount>
(
<collectionCode id="ED227CF3B33BDB274C3BF8732A43F850" box="[524,608,2010,2034]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="7" pageNumber="408">MPEG</collectionCode>
), labeled as follows: BRASIL-PARÁ, BR-422 /
<collectorName id="26C681E0B33BDB274AB6F8732C88F853" box="[1153,1195,2010,2033]" pageId="7" pageNumber="408">Km</collectorName>
11, sul
<collectorName id="26C681E0B33BDB274B33F8702D58F850" box="[1284,1403,2009,2034]" pageId="7" pageNumber="408">de Cametá</collectorName>
[=
</materialsCitation>
</paragraph>
<paragraph id="8B8CE436B334DB284EA0FF3E2BADFEE2" blockId="8.[151,1436,151,1949]" pageId="8" pageNumber="409">
south of Cametá City], Campina [= white-sand vegetation] /
<geoCoordinate id="EE0782F1B334DB284DB0FF3E2BDDFF12" box="[903,1022,151,176]" direction="south" orientation="latitude" pageId="8" pageNumber="409" precision="15" value="-2.2533333">2°15’12”S</geoCoordinate>
<geoCoordinate id="EE0782F1B334DB284A3AFF3E2CBFFF12" box="[1037,1180,151,176]" direction="west" orientation="longitude" pageId="8" pageNumber="409" precision="15" value="-49.603333">49°36’12”W</geoCoordinate>
/ Arm. Suspensa [= suspended trap], 0 5 a [= to]
<date id="FF8DC2F6B334DB284FEAFF152A7EFF77" box="[477,605,188,213]" pageId="8" pageNumber="409" value="2015-07-20">20.vii.2015</date>
[printed on white label] // BRASIL-PARÁ, Cametá/ Cols: [= collectors] F.S. Carvalho Fh., B.F. dos Anjos, L./ A. Quaresma, D.R. Guimarães/ 0 5 a [= to]
<date id="FF8DC2F6B334DB284A63FF762CE8FF5A" box="[1108,1227,223,248]" pageId="8" pageNumber="409" value="2015-11-20">20.xi.2015</date>
[printed on white label;
<typeStatus id="54885A94B334DB284EEBFEAD291DFEBF" box="[220,318,260,285]" pageId="8" pageNumber="409" type="holotype">holotype</typeStatus>
with left fore leg and right mid leg missing, and with segment 5 of abdomen and terminalia cleared and placed in glycerin in a microvial pinned beneath the specimen].
</paragraph>
</subSubSection>
<subSubSection id="C329B7BDB334DB284EF0FEE52C5AFE0D" pageId="8" pageNumber="409" type="diagnosis">
<paragraph id="8B8CE436B334DB284EF0FEE52C5AFE0D" blockId="8.[151,1436,151,1949]" pageId="8" pageNumber="409">
<emphasis id="B9473824B334DB284EF0FEE52962FEC7" bold="true" box="[199,321,332,357]" pageId="8" pageNumber="409">Diagnosis.</emphasis>
Palpus black and reduced to stubs. Male fore tarsomeres 3–5 with elongate hair-like setae dorsally (
<figureCitation id="1308F8B3B334DB284EA8FEC628D6FE2A" box="[159,245,367,392]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 22</figureCitation>
). Veins R
<subScript id="17B7E673B334DB284F52FED7294EFE2E" attach="left" box="[357,365,382,396]" fontSize="6" pageId="8" pageNumber="409">1</subScript>
and CuA
<subScript id="17B7E673B334DB284FEBFED729C7FE2E" attach="left" box="[476,484,382,396]" fontSize="6" pageId="8" pageNumber="409">1</subScript>
bare. Pregonite elongate and narrow, with pointed apex (
<figureCitation id="1308F8B3B334DB284A41FEC62CEFFE2A" box="[1142,1228,367,392]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 25</figureCitation>
). Phallus elongate and gently curved ventrally (
<figureCitation id="1308F8B3B334DB284FEEFE3F2A0DFE0D" box="[473,558,406,431]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 26</figureCitation>
). Acrophallus elongate and without spines (
<figureCitation id="1308F8B3B334DB284A2FFE3F2C4FFE0D" box="[1048,1132,406,431]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C329B7BDB334DB284EF0FE1329BCF9B3" pageId="8" pageNumber="409" type="description">
<paragraph id="8B8CE436B334DB284EF0FE132ACCFE71" blockId="8.[151,1436,151,1949]" box="[199,751,442,467]" pageId="8" pageNumber="409">
<emphasis id="B9473824B334DB284EF0FE13297AFE71" bold="true" box="[199,345,442,467]" pageId="8" pageNumber="409">Description.</emphasis>
<emphasis id="B9473824B334DB284F68FE132982FE71" box="[351,417,442,467]" italics="true" pageId="8" pageNumber="409">Male.</emphasis>
Body length:
<quantity id="4CCB49D3B334DB284C08FE132AB6FE70" box="[575,661,442,467]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="8" pageNumber="409" unit="mm" value="4.6">4.6 mm</quantity>
(n = 1).
</paragraph>
<paragraph id="8B8CE436B334DB284EF0FE772C13FD51" blockId="8.[151,1436,151,1949]" pageId="8" pageNumber="409">
Head. Frons distinctly protruding giving the head a trapezoidal profile. Fronto-orbital and parafacial plates covered with silver microtrichia. Fronto-orbital plate with some small scattered setulae in addition to 9 frontal setae continuing on to parafacial plate to below level of aristal insertion in lateral view. Frontal vitta black, narrowing towards lunule. Inner vertical setae differentiated, outer vertical setae developed [but both broken]. Ocellar setae of about the same length as frontal setae; 2 reclinate and 2 proclinate fronto-orbital setae arranged in partly overlapping rows with anterior reclinate orbital seta positioned anterior to posterior proclinate seta (see
<bibRefCitation id="EFA299C7B334DB284B71FD3A2910FD6D" author="Pape, T. &amp; Dahlem, G. A." pageId="8" pageNumber="409" pagination="1313 - 1335" refId="ref8565" refString="Pape, T. &amp; Dahlem, G. A. (2010) Sarcophagidae. In: Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, N. E. &amp; Zumbado, M. (Eds.), Manual of Central American Diptera. Uol. 2. NRC Research Press, Ottawa, pp. 1313 - 1335." type="book chapter" year="2010">Pape &amp; Dahlem 2010</bibRefCitation>
: fig. 6). Antenna dark brown, first flagellomere with gray microtrichia; arista thickened in proximal half and as long as first flagellomere. Palpus black, reduced to a rudimentary stub.
</paragraph>
<paragraph id="8B8CE436B334DB284EF0FD572908FB9E" blockId="8.[151,1436,151,1949]" pageId="8" pageNumber="409">
Thorax. Black, with silvery microtrichia. Chaetotaxy: acrostichals 0+1, dorsocentrals 2+3, intra-alars 1+2, supra-alars 2+3, postpronotals 2, postalars 2, notopleurals 2, anepisternals 4–5, katepisternals 3, merals 10, scutellum with 1 pair of basal setae, 1 pair of lateral setae, 1 pair of apical setae and no discal setae.
<emphasis id="B9473824B334DB284B13FCE12D7AFCFD" box="[1316,1369,840,863]" italics="true" pageId="8" pageNumber="409">Legs</emphasis>
. Fore femur with row of setae on posterodorsal and posteroventral surfaces. Fore tarsomere 3 with two long, hair-like setae dorsally (
<figureCitation id="1308F8B3B334DB284F08FC2729B0FC05" box="[319,403,910,935]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 22</figureCitation>
). Fore tarsomere 4 with one long, hair-like seta dorsally (
<figureCitation id="1308F8B3B334DB284A27FC272C46FC05" box="[1040,1125,910,935]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 22</figureCitation>
). Fore tarsomere 5 with one long dorsally seta (
<figureCitation id="1308F8B3B334DB284F58FC1B29E7FC69" box="[367,452,946,971]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 22</figureCitation>
). Mid tibia without anterodorsal and anteroventral setae. Hind femur with row of setae on posterodorsal and posteroventral surfaces.
<emphasis id="B9473824B334DB284C9BFC712AC7FC4D" box="[684,740,984,1007]" italics="true" pageId="8" pageNumber="409">Wing</emphasis>
. Hyaline. Tegula dark brown. Basicosta yellowish. Vein R
<subScript id="17B7E673B334DB284BA3FC4F2DBFFC56" attach="left" box="[1428,1436,998,1012]" fontSize="6" pageId="8" pageNumber="409">1</subScript>
bare. Vein R
<subScript id="17B7E673B334DB284F10FBA5291CFBB8" attach="left" box="[295,319,1036,1050]" fontSize="6" pageId="8" pageNumber="409">4+5</subScript>
with setae dorsally. Vein CuA
<subScript id="17B7E673B334DB284CAAFBA52A86FBB8" attach="left" box="[669,677,1036,1050]" fontSize="6" pageId="8" pageNumber="409">1</subScript>
bare. Costal spine not differentiated. Third costal section without ventral setae.
</paragraph>
<paragraph id="8B8CE436B334DB284EF0FBE12DA2FA4F" blockId="8.[151,1436,151,1949]" pageId="8" pageNumber="409">
Abdomen. Black with silvery-gray microtrichia in about anterior half of each tergite. Microtrichia on tergites 3 and 4 interrupted by a median black stripe. Syntergite 1+2 without marginal setae. Tergites 3 and 4 with a pair of median marginal setae. Tergite 5 with a row of marginal setae.
<emphasis id="B9473824B334DB284D4AFB392BDFFB0B" box="[893,1020,1168,1193]" italics="true" pageId="8" pageNumber="409">Terminalia.</emphasis>
Tergite 6, syntergosternite 7+8 and epandrium brown. Cercus dark brown, shorter than epandrium in lateral view, with distal portion abruptly narrowing towards apex in lateral view (
<figureCitation id="1308F8B3B334DB284C56FB712A94FB53" box="[609,695,1240,1265]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 23</figureCitation>
). Basal portion of cercus covered with many long setae (
<figureCitation id="1308F8B3B334DB284B0CFB712DB3FB53" box="[1339,1424,1240,1265]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 24</figureCitation>
). Ventral margin of cercal prong slightly curved; cercal apex rounded (
<figureCitation id="1308F8B3B334DB284DA2FB552BCBFAB7" box="[917,1000,1276,1301]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 23</figureCitation>
). Cercal prongs, in dorsal view, narrow and parallel (
<figureCitation id="1308F8B3B334DB284F1EFA89295EFA9B" box="[297,381,1312,1337]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 24</figureCitation>
). Surstylus dark brown, about half the length of cercus, with distal half narrowing toward apex and curved dorsally; with microtrichia restricted to basal half (
<figureCitation id="1308F8B3B334DB284D64FAED2B84FAFF" box="[851,935,1348,1373]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 23</figureCitation>
). Pregonite elongate, longer than postgonite, narrowed and curved ventrally (
<figureCitation id="1308F8B3B334DB284C3EFAC12A7CFA23" box="[521,607,1384,1409]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 25</figureCitation>
). Postgonite longer than wide, with a long pre-apical seta and a row of small setae along ventral margin (
<figureCitation id="1308F8B3B334DB284C25FA252A45FA07" box="[530,614,1420,1445]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 25</figureCitation>
). Phallus elongate, gently curved ventrally, with ventral and dorsal plates well developed (
<figureCitation id="1308F8B3B334DB284F61FA19298DFA6B" box="[342,430,1456,1481]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 26</figureCitation>
). Distiphallus with a distinct ventral swelling, with many membranous cuticular spines (
<figureCitation id="1308F8B3B334DB284EA8FA7D28D7FA4F" box="[159,244,1492,1517]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 26</figureCitation>
). Acrophallus elongate and rounded apically, with some wrinkles and without cuticular spines (
<figureCitation id="1308F8B3B334DB284B17FA7D2D56FA4F" box="[1312,1397,1492,1517]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 26</figureCitation>
).
</paragraph>
<paragraph id="8B8CE436B334DB284EF0FA5129BCF9B3" blockId="8.[151,1436,151,1949]" box="[199,415,1528,1553]" pageId="8" pageNumber="409">
<emphasis id="B9473824B334DB284EF0FA512900F9B3" box="[199,291,1528,1553]" italics="true" pageId="8" pageNumber="409">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C329B7BDB334DB284EF0F9B5297EF893" pageId="8" pageNumber="409" type="discussion">
<paragraph id="8B8CE436B334DB284EF0F9B5297EF893" blockId="8.[151,1436,151,1949]" pageId="8" pageNumber="409">
<emphasis id="B9473824B334DB284EF0F9B52918F997" bold="true" box="[199,315,1564,1589]" pageId="8" pageNumber="409">Remarks.</emphasis>
<taxonomicName id="4C339FB5B334DB284F74F9B429F7F997" authority="Carvalho-Filho &amp; Gorayeb &amp; Soares &amp; Souza, 2018" authorityName="Carvalho-Filho &amp; Gorayeb &amp; Soares &amp; Souza" authorityYear="2018" box="[323,468,1565,1589]" class="Insecta" family="Sarcophagidae" genus="Metopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="fofo" status="sp. nov.">
<emphasis id="B9473824B334DB284F74F9B429F7F997" box="[323,468,1565,1589]" italics="true" pageId="8" pageNumber="409">Metopia fofo</emphasis>
</taxonomicName>
<emphasis id="B9473824B334DB284FEBF9B52A15F997" bold="true" box="[476,566,1564,1589]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="A274855FB334DB284FEBF9B52A15F997" box="[476,566,1564,1589]" pageId="8" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
runs to
<taxonomicName id="4C339FB5B334DB284CA3F9B42BF9F997" authority="Sabrosky" authorityName="Sabrosky" box="[660,986,1564,1589]" class="Insecta" family="Sarcophagidae" genus="Metopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="krombeini">
<emphasis id="B9473824B334DB284CA3F9B42B49F997" box="[660,874,1564,1589]" italics="true" pageId="8" pageNumber="409">Metopia krombeini</emphasis>
Sabrosky
</taxonomicName>
in the most recent key to the American species of
<emphasis id="B9473824B334DB284F26F9E8294DF9FA" box="[273,366,1601,1624]" italics="true" pageId="8" pageNumber="409">Metopia</emphasis>
Meigen (
<bibRefCitation id="EFA299C7B334DB284FE9F9E82A44F9FB" author="Pape, T." box="[478,615,1601,1625]" pageId="8" pageNumber="409" pagination="81 - 101" refId="ref8396" refString="Pape, T. (1987 a) Revision of Neotropical Metopia Meigen (Diptera: Sarcophagidae). Systematic Entomology, 12, 81 - 101. https: // doi. org / 10.1111 / j. 1365 - 3113.1987. tb 00550. x" type="journal article" year="1987">Pape 1987a</bibRefCitation>
), but differs from this species in having phallus gently curved ventrally (
<figureCitation id="1308F8B3B334DB284EA8F9CD28D1F9DF" box="[159,242,1636,1661]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 26</figureCitation>
), acrophallus without spines and rounded apically (
<figureCitation id="1308F8B3B334DB284D1AF9CD2BA3F9DF" box="[813,896,1636,1661]" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 26</figureCitation>
), and cercal prong narrowed in dorsal view (
<figureCitation id="1308F8B3B334DB284B58F9CD2894F903" captionStart="FIGURES 22–26" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[206,1397,208,1888]" captionTargetId="figure@12.[191,1401,193,1898]" captionTargetPageId="12" captionText="FIGURES 22–26. Metopia fofo sp. nov., holotype ♂. 22. Right fore tarsus, dorsal view. 23. Epandrium, surstylus and cercus, left lateral view. 24. Cerci, dorsal view. 25. Gonites, left lateral view. 26. Phallus, left lateral view. Abbreviations: acp = acrophallus; dpl = dorsal plate; prg = pregonite; ptg = postgonite; vpl = ventral plate. Scale bars: 0.1 mm (Figs 22–25); 0.2 mm (Fig. 21)." httpUri="https://zenodo.org/record/2606519/files/figure.png" pageId="8" pageNumber="409">Fig. 24</figureCitation>
). In
<taxonomicName id="4C339FB5B334DB284EDEF920295DF903" box="[233,382,1672,1697]" class="Insecta" family="Sarcophagidae" genus="Metopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="krombeini">
<emphasis id="B9473824B334DB284EDEF920295DF903" box="[233,382,1672,1697]" italics="true" pageId="8" pageNumber="409">M. krombeini</emphasis>
</taxonomicName>
the phallus is almost straight, the acrophallus has small cuticular spines and is subrectangular apically, and the cercal prong has an enlarged base and narrow tip (
<bibRefCitation id="EFA299C7B334DB284DBFF9042CE3F967" author="Pape, T." box="[904,1216,1708,1733]" pageId="8" pageNumber="409" pagination="81 - 101" refId="ref8396" refString="Pape, T. (1987 a) Revision of Neotropical Metopia Meigen (Diptera: Sarcophagidae). Systematic Entomology, 12, 81 - 101. https: // doi. org / 10.1111 / j. 1365 - 3113.1987. tb 00550. x" type="journal article" year="1987">Pape 1987a: figs 27, 30, 33</bibRefCitation>
). The phallus of
<taxonomicName id="4C339FB5B334DB284B4AF90428E0F94B" authority="Carvalho-Filho &amp; Gorayeb &amp; Soares &amp; Souza, 2018" authorityName="Carvalho-Filho &amp; Gorayeb &amp; Soares &amp; Souza" authorityYear="2018" class="Insecta" family="Sarcophagidae" genus="Metopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="fofo" status="sp. nov.">
<emphasis id="B9473824B334DB284B4AF90428E0F94B" italics="true" pageId="8" pageNumber="409">M. fofo</emphasis>
</taxonomicName>
<emphasis id="B9473824B334DB284EFCF9792904F94B" bold="true" box="[203,295,1744,1769]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="A274855FB334DB284EFCF9792904F94B" box="[203,295,1744,1769]" pageId="8" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is very similar to that of
<taxonomicName id="4C339FB5B334DB284C79F9782B05F94B" authority="Allen" authorityName="Allen" box="[590,806,1744,1769]" class="Insecta" family="Sarcophagidae" genus="Metopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="lateropili">
<emphasis id="B9473824B334DB284C79F9782AFFF94B" box="[590,732,1744,1769]" italics="true" pageId="8" pageNumber="409">M. lateropili</emphasis>
Allen
</taxonomicName>
, since both share a distinct ventral swelling with many membranous cuticular spines.
<taxonomicName id="4C339FB5B334DB284FCDF95C2AE9F8AF" box="[506,714,1780,1805]" class="Insecta" family="Sarcophagidae" genus="Metopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="lateropili">
<emphasis id="B9473824B334DB284FCDF95C2AE9F8AF" box="[506,714,1780,1805]" italics="true" pageId="8" pageNumber="409">Metopia lateropili</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C339FB5B334DB284D5BF95C2BE1F8AF" authority="Carvalho-Filho &amp; Gorayeb &amp; Soares &amp; Souza, 2018" authorityName="Carvalho-Filho &amp; Gorayeb &amp; Soares &amp; Souza" authorityYear="2018" box="[876,962,1781,1805]" class="Insecta" family="Sarcophagidae" genus="Metopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="fofo" status="sp. nov.">
<emphasis id="B9473824B334DB284D5BF95C2BE1F8AF" box="[876,962,1781,1805]" italics="true" pageId="8" pageNumber="409">M. fofo</emphasis>
</taxonomicName>
<emphasis id="B9473824B334DB284DF9F95D2C0FF8AF" bold="true" box="[974,1068,1780,1805]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="A274855FB334DB284DF9F95D2C0FF8AF" box="[974,1068,1780,1805]" pageId="8" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having palpus elongate, not reduced to a stub.
</paragraph>
</subSubSection>
<subSubSection id="C329B7BDB334DB284EF0F8952AFEF8F7" box="[199,733,1852,1877]" pageId="8" pageNumber="409" type="distribution">
<paragraph id="8B8CE436B334DB284EF0F8952AFEF8F7" blockId="8.[151,1436,151,1949]" box="[199,733,1852,1877]" pageId="8" pageNumber="409">
<emphasis id="B9473824B334DB284EF0F8952943F8F7" bold="true" box="[199,352,1852,1877]" pageId="8" pageNumber="409">Distribution.</emphasis>
NEOTROPICAL—Brazil (
<collectingRegion id="49F72AD4B334DB284CAEF8942AF2F8F7" box="[665,721,1853,1877]" country="Brazil" name="Para" pageId="8" pageNumber="409">Pará</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C329B7BDB334DB284EF0F8C92BB1F83F" pageId="8" pageNumber="409" type="etymology">
<paragraph id="8B8CE436B334DB284EF0F8C92BB1F83F" blockId="8.[151,1436,151,1949]" pageId="8" pageNumber="409">
<emphasis id="B9473824B334DB284EF0F8C9296DF8DB" bold="true" box="[199,334,1888,1913]" pageId="8" pageNumber="409">Etymology.</emphasis>
The specific epithet is derived from the word “fofó”, which is a famous popular festival that takes place before the carnival in the streets of Cametá (the type-locality).
</paragraph>
</subSubSection>
</treatment>
</document>
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