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<document id="3FAFE9FC4F76AF222F482240CAD378DD" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2015" docId="039787812E5FFFD294FD4FDBE8CDEEEF" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) coreanus Tschitscherine 1895" docType="treatment" docVersion="7" lastPageNumber="20" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="19" updateTime="1698562432667" updateUser="plazi">
<mods:mods id="D5F1484BFB0B741390ACACD38D26A1D7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A2AA74D7521A61DE97A7935BD36A0993">
<mods:title id="538F8B7DB256170A7ADB87D07D131471">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="06C01866836FA9F8A2461EF472BB6A2C">Kataev, Boris M.</mods:namePart>
</mods:name>
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<mods:role id="9BB3E3319855D0421452500B4A4C5F8D">
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<mods:namePart id="AE8AF96F62B1A2EE30C252FEDB0A953D">Liang, Hongbin</mods:namePart>
</mods:name>
<mods:typeOfResource id="C1E7722B57131FBBEDF08D5AC4CA9116">text</mods:typeOfResource>
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<mods:title id="A2FF97A1FFE265D495A5BAB390AF5634">Zootaxa</mods:title>
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<mods:part id="B8C370A841609C7F48BC1DEFFE7FBC79">
<mods:date id="F39768D5CC3D29538180047688AE727B">2015</mods:date>
<mods:detail id="2CF77D86BDBD4F490C4A22DCD328B4CD" type="volume">
<mods:number id="A1F9EF727CB387161E9560E5D20272BD">3920</mods:number>
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<mods:detail id="C69F064F6B5DB3647CD2890C57951B41" type="issue">
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<mods:identifier id="CD2B8D7621621F5F4966FAAE641AF802" type="DOI">10.11646/zootaxa.3920.1.1</mods:identifier>
<mods:identifier id="7396BB7AEB2F266B9624C8A398CDA95C" type="GBIF-Dataset">7289c86d-0ed1-4e93-b7b3-bc596042d854</mods:identifier>
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</mods:mods>
<treatment id="039787812E5FFFD294FD4FDBE8CDEEEF" ID-DOI="http://doi.org/10.5281/zenodo.6114266" ID-GBIF-Taxon="127671640" ID-Zenodo-Dep="6114266" LSID="urn:lsid:plazi:treatment:039787812E5FFFD294FD4FDBE8CDEEEF" httpUri="http://treatment.plazi.org/id/039787812E5FFFD294FD4FDBE8CDEEEF" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="C324651C2E5FFFD394FD4FDBE800E931" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8B8136972E5FFFD394FD4FDBEA56E910" blockId="18.[151,881,1520,1580]" box="[151,881,1520,1546]" pageId="18" pageNumber="19">
<heading id="D0C981FB2E5FFFD394FD4FDBEA56E910" bold="true" box="[151,881,1520,1546]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="4C3E4D142E5FFFD394FD4FDBEA56E910" ID-CoL="3JSTJ" authority="Tschitscherine, 1895" authorityName="Tschitscherine" authorityYear="1895" box="[151,881,1520,1546]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="coreanus" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E5FFFD394FD4FDBEA56E910" bold="true" box="[151,881,1520,1546]" pageId="18" pageNumber="19">
<emphasis id="B94AEA852E5FFFD394FD4FDBE82EE910" bold="true" box="[151,265,1520,1546]" italics="true" pageId="18" pageNumber="19">Harpalus</emphasis>
(
<emphasis id="B94AEA852E5FFFD395704FDBE8FDE910" bold="true" box="[282,474,1520,1546]" italics="true" pageId="18" pageNumber="19">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E5FFFD395814FDBEB7DE910" bold="true" box="[491,602,1520,1546]" italics="true" pageId="18" pageNumber="19">coreanus</emphasis>
(
<bibRefCitation id="EFAF4B662E5FFFD396034FDBEA4EE910" author="Tschitscherine" box="[617,873,1520,1546]" pageId="18" pageNumber="19" refString="Tschitscherine, T. (1895) Supplement a la faune des Carabiques de la Coree. Horae Societatis Entomologicae Rossicae, 29, 154 - 188." type="journal article" year="1895">Tschitschérine, 1895</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8136972E5FFFD394FD4C38E800E931" blockId="18.[151,881,1520,1580]" box="[151,295,1555,1580]" pageId="18" pageNumber="19">
(
<figureCitation id="13052A122E5FFFD394F54C38E838E931" box="[159,287,1555,1580]" captionStart="FIGURES 36 – 39" captionStartId="18.[151,263,1386,1409]" captionTargetBox="[236,1350,454,1359]" captionTargetId="figure@18.[230,1356,447,1365]" captionTargetPageId="18" captionText="FIGURES 36 – 39. Harpalus, median lobe of aedeagus. 36 – 37, H. simplicidens (Liaoning); 38 – 39, H. coreanus (Gansu). 36, 38, view from the left side; 37, 39, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287893/files/figure.png" pageId="18" pageNumber="19">Figs 38–39</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E5FFFD394FD4C70EA9CE96B" box="[151,955,1627,1649]" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="8B8136972E5FFFD394FD4C70EA9CE96B" blockId="18.[151,955,1627,1649]" box="[151,955,1627,1649]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="AB2E11B92E5FFFD394FD4C70EA9CE96B" box="[151,955,1627,1649]" pageId="18" pageNumber="19">
<taxonomicName id="4C3E4D142E5FFFD394FD4C70EBFCE96B" ID-CoL="74PCM" authority="Tschitscherine, 1895: 156" authorityName="Tschitscherine" authorityPageNumber="156" authorityYear="1895" box="[151,731,1627,1649]" class="Insecta" family="Carabidae" genus="Ophonus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="coreanus" subGenus="Pardileus">
<emphasis id="B94AEA852E5FFFD394FD4C70E9D4E96A" box="[151,243,1627,1648]" italics="true" pageId="18" pageNumber="19">Ophonus</emphasis>
(
<emphasis id="B94AEA852E5FFFD3956B4C70E845E96A" box="[257,354,1627,1648]" italics="true" pageId="18" pageNumber="19">Pardileus</emphasis>
)
<emphasis id="B94AEA852E5FFFD3951B4C77E8EAE96B" box="[369,461,1628,1649]" italics="true" pageId="18" pageNumber="19">coreanus</emphasis>
<treatmentCitation id="0A9F10862E5FFFD395BE4C70EBFCE96B" author="Tschitscherine" box="[468,731,1627,1649]" page="156" pageId="18" pageNumber="19" year="1895">
<bibRefCitation id="EFAF4B662E5FFFD395BE4C70EB8FE96B" author="Tschitscherine" box="[468,680,1627,1649]" pageId="18" pageNumber="19" refString="Tschitscherine, T. (1895) Supplement a la faune des Carabiques de la Coree. Horae Societatis Entomologicae Rossicae, 29, 154 - 188." type="journal article" year="1895">Tschitschérine, 1895</bibRefCitation>
: 156
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E5FFFD3968E4C77EA31E96B" box="[740,790,1628,1649]" pageId="18" pageNumber="19">Type</typeStatus>
locality:
<collectingCountry id="F32976072E5FFFD3971C4C77EA90E96B" box="[886,951,1628,1649]" name="South Korea" pageId="18" pageNumber="19">Korea</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E5FFFD394FD4CB6EC3FE8A8" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8B8136972E5FFFD394FD4CB6ED9DE9C0" blockId="18.[151,1437,1693,2006]" pageId="18" pageNumber="19">
<emphasis id="B94AEA852E5FFFD394FD4CB6E867E9AC" bold="true" box="[151,320,1693,1718]" pageId="18" pageNumber="19">
<typeStatus id="548588352E5FFFD394FD4CB6E9F5E9AC" box="[151,210,1693,1718]" pageId="18" pageNumber="19">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548588352E5FFFD3953B4CB5E8E6E9AC" box="[337,449,1694,1718]" pageId="18" pageNumber="19" type="lectotype">Lectotype</typeStatus>
[designated by
<bibRefCitation id="EFAF4B662E5FFFD396114CB6EA19E9AC" author="Lafer" box="[635,830,1693,1718]" pageId="18" pageNumber="19" refString="Lafer, G. Sh. (1996) Notes on some species of the subtribe Harpalina (Coleoptera, Carabidae, Harpalinae) from Korea and adjacent areas. Insecta Koreana, 13, 77 - 89." type="journal article" year="1996">Lafer (1996: 80)</bibRefCitation>
]: ♂, labeled &quot;
<collectingCountry id="F32976072E5FFFD397B64CB5ED04E9AC" box="[988,1059,1694,1718]" name="South Korea" pageId="18" pageNumber="19">Korea</collectingCountry>
(Staud.–Rttr.)&quot;, &quot;
<taxonomicName id="4C3E4D142E5FFFD390864CB6EC1DE9AF" box="[1260,1338,1693,1717]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E5FFFD390864CB6EC1DE9AF" box="[1260,1338,1693,1717]" italics="true" pageId="18" pageNumber="19">tridens</emphasis>
</taxonomicName>
A.
<taxonomicName id="4C3E4D142E5FFFD391354CB5E9DAE9C3" class="Insecta" family="Carabidae" genus="Migadophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="variety" variety="coreanus">
M. v.
<emphasis id="B94AEA852E5FFFD394FD4CE9E9DAE9C3" box="[151,253,1730,1753]" italics="true" pageId="18" pageNumber="19">coreanus</emphasis>
</taxonomicName>
m. Typ. Tschitscherin det.&quot;, &quot;
<taxonomicName id="4C3E4D142E5FFFD396204CE9EA99E9C0" authority="Tschit." authorityName="Tschit." box="[586,958,1729,1754]" class="Insecta" family="Carabidae" genus="Pseudoophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="coreanus">
<emphasis id="B94AEA852E5FFFD396204CE9EA4FE9C3" box="[586,872,1730,1754]" italics="true" pageId="18" pageNumber="19">Pseudoophonus coreanus</emphasis>
Tschit.
</taxonomicName>
, 74. Lafer det.&quot; (
<collectionCode id="ED2FAE522E5FFFD390134CE9ED8AE9C0" box="[1145,1197,1730,1754]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="18" pageNumber="19">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E5FFFD394AD4CCEEC3FE8A8" blockId="18.[151,1437,1693,2006]" pageId="18" pageNumber="19">
<emphasis id="B94AEA852E5FFFD394AD4CCEEB1AE9E4" bold="true" box="[199,573,1765,1790]" pageId="18" pageNumber="19">Additional material examined</emphasis>
. More than
<specimenCount id="9D38FD1E2E5FFFD396B44CCEEAABE9E4" box="[734,908,1765,1790]" pageId="18" pageNumber="19" type="generic">450 specimens</specimenCount>
(388 from
<collectingCountry id="F32976072E5FFFD390484CCEED4EE9E4" box="[1058,1129,1765,1790]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
), including the following specimens from Anhui, Henan, Zhejiang, Shanghai, Hubei and Guangdong.
<collectingCountry id="F32976072E5FFFD390564D22EDA2E838" box="[1084,1157,1801,1826]" name="China" pageId="18" pageNumber="19">
<emphasis id="B94AEA852E5FFFD390564D22EDA2E838" bold="true" box="[1084,1157,1801,1826]" pageId="18" pageNumber="19">China</emphasis>
</collectingCountry>
.
<collectionCode id="ED2FAE522E5FFFD390F64D21EDDDE838" box="[1180,1274,1802,1826]" pageId="18" pageNumber="19">ANHUI</collectionCode>
: 1 ♀, Hefei,
<date id="FF8010572E5FFFD394FD4D06E80CE85C" box="[151,299,1837,1862]" pageId="18" pageNumber="19" value="1981-08-15">15.VIII.1981</date>
, Qin Yisheng leg. (
<collectionCode id="ED2FAE522E5FFFD3966B4D05EB12E85C" box="[513,565,1838,1862]" pageId="18" pageNumber="19">IOZ</collectionCode>
).
<collectionCode id="ED2FAE522E5FFFD3962D4D05EB89E85C" box="[583,686,1838,1862]" pageId="18" pageNumber="19">HENAN</collectionCode>
: 1 ♀, Neixiang Co.,
<quantity id="4CC69B722E5FFFD397FA4D06EAFFE85F" box="[912,984,1837,1862]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="m" value="600.0">600 m</quantity>
,
<date id="FF8010572E5FFFD3978E4D05ED48E85C" box="[996,1135,1838,1862]" pageId="18" pageNumber="19" value="1998-07-12">12.VII.1998</date>
, Zhang Youwei leg. (
<collectionCode id="ED2FAE522E5FFFD391364D05ECB7E85C" box="[1372,1424,1838,1862]" pageId="18" pageNumber="19">IOZ</collectionCode>
). ZHEJIANG:
<specimenCount id="9D38FD1E2E5FFFD3955E4D79E841E873" box="[308,358,1874,1898]" pageId="18" pageNumber="19" type="generic">1 ♂</specimenCount>
, Qingyuan Co.,
<date id="FF8010572E5FFFD3965B4D79EBE2E870" box="[561,709,1874,1898]" pageId="18" pageNumber="19" value="1991-08-19">19.VIII.1991</date>
, Wu Hong leg. (
<collectionCode id="ED2FAE522E5FFFD397FD4D79EAECE870" box="[919,971,1874,1898]" pageId="18" pageNumber="19">IOZ</collectionCode>
); 1 ♀, Mokan Shan,
<date id="FF8010572E5FFFD390BE4D7AEC61E870" box="[1236,1350,1873,1898]" pageId="18" pageNumber="19" value="1936-05-18">18.V.1936</date>
(
<collectionCode id="ED2FAE522E5FFFD391364D79ECB7E870" box="[1372,1424,1874,1898]" pageId="18" pageNumber="19">IOZ</collectionCode>
). SHANGHAI:
<specimenCount id="9D38FD1E2E5FFFD395544D5DE856E897" box="[318,369,1910,1934]" pageId="18" pageNumber="19" type="generic">2 ♂</specimenCount>
, 1 ♀, Shanghai,
<date id="FF8010572E5FFFD3962E4D5DEBE0E894" box="[580,711,1910,1934]" pageId="18" pageNumber="19" value="1938-09-14">14.IX.1938</date>
, O. Piel leg. (
<collectionCode id="ED2FAE522E5FFFD397134D5DEA8AE894" box="[889,941,1910,1934]" pageId="18" pageNumber="19">IOZ</collectionCode>
).
<collectionCode id="ED2FAE522E5FFFD397AC4D5DED07E897" box="[966,1056,1910,1933]" pageId="18" pageNumber="19">HUBEI</collectionCode>
: 1 ♀, Shennongjia, Songbai T.,
<date id="FF8010572E5FFFD394FD4DB2E833E8A8" box="[151,276,1945,1970]" pageId="18" pageNumber="19" value="1980-07-05">5.VII.1980</date>
, Yu Peiyu leg. (
<collectionCode id="ED2FAE522E5FFFD395AF4DB1E8DEE8A8" box="[453,505,1946,1970]" pageId="18" pageNumber="19">IOZ</collectionCode>
). GUANGDONG: 1 ♀, Dongguan Co.,
<date id="FF8010572E5FFFD397D04DB1ED3FE8A8" box="[954,1048,1946,1970]" pageId="18" pageNumber="19" value="1981-03">III.1981</date>
, Liu Tanjin leg. (
<collectionCode id="ED2FAE522E5FFFD390B24DB1EC2BE8A8" box="[1240,1292,1946,1970]" pageId="18" pageNumber="19">IOZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E5FFFD294AD4D96EAE4EFED" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8B8136972E5FFFD294AD4D96EAE4EFED" blockId="18.[151,1437,1693,2006]" lastBlockId="19.[151,1436,151,501]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B94AEA852E5FFFD394AD4D96E81DE8CC" bold="true" box="[199,314,1981,2006]" pageId="18" pageNumber="19">Diagnosis</emphasis>
. This species is similar in habitus to
<taxonomicName id="4C3E4D142E5FFFD3968F4D94EA68E8CC" box="[741,847,1983,2006]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pastor">
<emphasis id="B94AEA852E5FFFD3968F4D94EA68E8CC" box="[741,847,1983,2006]" italics="true" pageId="18" pageNumber="19">H. pastor</emphasis>
</taxonomicName>
, but differs from the latter in having pronotal basal angles with sharp denticle at apex (occasionally denticle very small, indistinct) and with oblique convex fold underneath, elytral interval 7 without preapical pores. The internal sac of the median lobe of aedeagus (
<figureCitation id="13052A122E5EFFD2917F4A97ECB3EFCE" box="[1301,1428,188,213]" captionStart="FIGURES 36 – 39" captionStartId="18.[151,263,1386,1409]" captionTargetBox="[236,1350,454,1359]" captionTargetId="figure@18.[230,1356,447,1365]" captionTargetPageId="18" captionText="FIGURES 36 – 39. Harpalus, median lobe of aedeagus. 36 – 37, H. simplicidens (Liaoning); 38 – 39, H. coreanus (Gansu). 36, 38, view from the left side; 37, 39, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287893/files/figure.png" pageId="19" pageNumber="20">Figs 38–39</figureCitation>
) is without any distinct sclerotized elements. Body length
<quantity id="4CC69B722E5EFFD297734AF4EA98EFED" box="[793,959,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.435" metricValueMax="1.59" metricValueMin="1.28" pageId="19" pageNumber="20" unit="mm" value="14.35" valueMax="15.9" valueMin="12.8">12.8–15.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E5EFFD294AD4B2FE8CDEEEF" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8B8136972E5EFFD294AD4B2FEABFEE92" blockId="19.[151,1436,151,501]" pageId="19" pageNumber="20">
<emphasis id="B94AEA852E5EFFD294AD4B2FE87EEE07" bold="true" box="[199,345,260,285]" pageId="19" pageNumber="20">Distribution</emphasis>
.
<collectingCountry id="F32976072E5EFFD295194B2FE89DEE07" box="[371,442,260,285]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
: the north-eastern and eastern parts. The species was previously known from Heilongjiang, Liaoning, Beijing, Hebei, Nei Mongol, Shanxi, Shaanxi, Gansu, Sichuan, Hubei, Henan, Anhui, Shanghai, Zhejiang, Fujian and Guangdong (
<bibRefCitation id="EFAF4B662E5EFFD296CA4B67EA10EE7E" author="Kataev" box="[672,823,332,356]" pageId="19" pageNumber="20" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev 1997</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5EFFD2972D4B66ED03EE7E" author="Kataev" box="[839,1060,332,356]" pageId="19" pageNumber="20" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E5EFFD297CB4B66EAFAEE7E" box="[929,989,332,356]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2003
</bibRefCitation>
); we record it here from Anhui, Henan, Zhejiang, Shanghai, Hubei and Guangdong for the first time.
</paragraph>
<paragraph id="8B8136972E5EFFD294AD4BBFE8CDEEEF" blockId="19.[151,1436,151,501]" pageId="19" pageNumber="20">
The species also occurs in the south of eastern
<collectingCountry id="F32976072E5EFFD296B74BBFEA0FEEB7" box="[733,808,404,429]" name="Russia" pageId="19" pageNumber="20">Russia</collectingCountry>
(Buryatia, Chita and Amur Provinces, Khabarovsk and Maritime Territories), and North and
<collectingCountry id="F32976072E5EFFD296504B9CEBE1EED5" box="[570,710,439,464]" name="South Korea" pageId="19" pageNumber="20">South Korea</collectingCountry>
(
<bibRefCitation id="EFAF4B662E5EFFD296BF4B9CEA70EECA" author="Lafer" box="[725,855,439,464]" pageId="19" pageNumber="20" refString="Lafer, G. Sh. (1989) Fam. Carabidae-The ground-beetles. In: Lehr, P. A. (Ed.), Opredelitel' nasekomykh Dal'nego Vostoka SSSR v shesti tomakh. Tom III. Zhestkokrylye, ili zhuki. Chast' 1. Nauka, Leningrad, pp. 70 - 222. [in Russian]" type="journal article" year="1989">Lafer 1989</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5EFFD297084B9CEDAFEECA" author="Kryzhanovskij" box="[866,1160,439,464]" pageId="19" pageNumber="20" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, B. M., Makarov, K. V., Shilenkov, V. G. (1995) A checklist of the groundbeetles of Russia and adjacent lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers, Sofia - Moscow, 272 pp." type="book" year="1995">
Kryzhanovskij
<emphasis id="B94AEA852E5EFFD290674B92ED18EECA" box="[1037,1087,440,464]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. 1995
</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5EFFD290FE4B93EC02EECA" author="Kataev" box="[1172,1317,440,464]" pageId="19" pageNumber="20" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev 1997</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5EFFD2915B4B93E9D1EEEE" author="Kataev" pageId="19" pageNumber="20" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E5EFFD291E24B92E98AEEEE" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. 2003
</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E5EFFD2956B4BF7E8F9EEEF" author="Paik" box="[257,478,476,501]" pageId="19" pageNumber="20" refString="Paik, J. - Ch. &amp; Moon, Ch. - S. (2005) Some ground bettles (Coleoptera, Carabidae) from Korea (15). Korean Journal of Soil Zoology, 10 (1 - 2), 42 - 54." type="journal article" year="2005">Paik &amp; Moon 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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