140 lines
18 KiB
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140 lines
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<mods:titleInfo id="A3E5A108C226AF4D84E218B99B96FE98">
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<mods:title id="B4537F5C8857CFCC767269B3C03B2338">Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco</mods:title>
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<mods:namePart id="F1432BF2D917C318FBACDF1C536278A8">Zouhri, Samir</mods:namePart>
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<mods:namePart id="B5A352F1C228DCCD199A7FC1AF0C00E3">Gingerich, Philip D.</mods:namePart>
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<mods:namePart id="D6EF11755B94CCEA5B4EE0F7100E4295">Khalloufi, Bouziane</mods:namePart>
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<mods:namePart id="9407FAE23A812571064BDF1131BC360A">Bourdon, Estelle</mods:namePart>
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<mods:namePart id="32728D68D88237B70273980090E1659C">Adnet, Sylvain</mods:namePart>
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<mods:namePart id="08AE0F83AFF802B8FEF05EF1B3B72F2A">Jouve, Stéphane</mods:namePart>
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<mods:namePart id="757656808D6BB267541D28F8F6E5A687">Elboudali, Najia</mods:namePart>
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<mods:namePart id="440EF5FD95DF30032271F2A95F7598E9">Amane, Ayoub</mods:namePart>
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<mods:namePart id="FD8C586B823343642E689EBEF1D23162">Rage, Jean-Claude</mods:namePart>
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<mods:namePart id="A72340908F7CC552F96FCDB90DBDBE4C">Tabuce, Rodolphe</mods:namePart>
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<mods:typeOfResource id="649273211FBAE25D169A0959715C16C2">text</mods:typeOfResource>
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<mods:title id="6D6D340CA45962D497CC2E874F4C17BF">Geodiversitas</mods:title>
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<mods:part id="3B463410E6727ED7A2B9333A566775E4">
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<mods:date id="8DB929285A03AC60258FF72B29242E74">2021</mods:date>
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<mods:number id="BCB34ED42B527185C388C81F74E12700">2021-03-11</mods:number>
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<mods:number id="56D7D49C23D6DA8C671BFF913057847D">43</mods:number>
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<mods:detail id="B127F31ED694E0552613E291DCA42F53" type="issue">
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<mods:number id="4639FFBD223154E1AC5CBC25CA250EF4">5</mods:number>
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<mods:start id="E6AA14CBAC76BB28B9E244A3E9535A71">121</mods:start>
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<mods:classification id="72882DF6DE5CF49D0495029098027594">journal article</mods:classification>
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<mods:identifier id="42E257C40506360A3229E6433EE5FAEC" type="CLB-Dataset">7746</mods:identifier>
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<mods:identifier id="8D4A6EF38E4964B1D86790A9BE32A8AE" type="DOI">10.5252/geodiversitas2021v43a5</mods:identifier>
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<mods:identifier id="61EDB24CB044692CB02417582CD1444F" type="GBIF-Dataset">e42658a2-0e15-4e90-9bc5-0d00cbcbd781</mods:identifier>
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<mods:identifier id="C0B8478E062EC3F4CB069E44199D6E5C" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="FD85802A8A0A40563D07EA041EE50A65" type="Zenodo-Dep">4605963</mods:identifier>
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<mods:identifier id="01DFB5D6ED363B367D83A8901FA30EAB" type="ZooBank">urn:lsid:zoobank.org:pub:697FC553-E37B-4EF9-97A4-950E4DEE246C</mods:identifier>
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</mods:mods>
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<subSection id="E2B494B8FF92FF9B3086FE69FD6712E0" pageId="19" pageNumber="138" type="multiple">
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<treatment id="03923C45FF92FF9B3086FE69FD6712E0" ID-DOI="http://doi.org/10.5281/zenodo.4606649" ID-GBIF-Taxon="180569432" ID-Zenodo-Dep="4606649" LSID="urn:lsid:plazi:treatment:03923C45FF92FF9B3086FE69FD6712E0" httpUri="http://treatment.plazi.org/id/03923C45FF92FF9B3086FE69FD6712E0" lastPageNumber="138" pageId="19" pageNumber="138">
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<subSubSection id="C321DED8FF92FF9B3086FE69FD7817BF" box="[158,749,438,465]" pageId="19" pageNumber="138" type="nomenclature">
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<paragraph id="8B848D53FF92FF9B3086FE69FD7817BF" blockId="19.[158,749,438,465]" box="[158,749,438,465]" pageId="19" pageNumber="138">
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<heading id="D0CC3A3FFF92FF9B3086FE69FD7817BF" box="[158,749,438,465]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="138" reason="2">
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Clade
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<taxonomicName id="4C3BF6D0FF92FF9B30FCFE68FD9F17BF" authority="Huxley, 1875" box="[228,522,438,465]" kingdom="Animalia" pageId="19" pageNumber="138">EUSUCHIA Huxley, 1875</taxonomicName>
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<emphasis id="B94F5141FF92FF9B3209FE68FDD017BE" box="[529,581,439,464]" italics="true" pageId="19" pageNumber="138">sensu</emphasis>
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<bibRefCitation id="EFAAF0A2FF92FF9B3255FE69FD7817BF" author="BROCHU C. A." box="[589,749,438,465]" pageId="19" pageNumber="138" pagination="357 - 397" refId="ref20017" refString="BROCHU C. A. 2003. - Phylogenetic approaches toward crocodylian history. Annual Review of Earth and Planetary Sciences 31: 357 - 397. https: // doi. org / 10.1146 / annurev. earth. 31.100901.141308" type="journal article" year="2003">Brochu (2003)</bibRefCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF92FF9B316AFE29FD8F147E" box="[370,538,502,528]" pageId="19" pageNumber="138" type="description">
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<paragraph id="8B848D53FF92FF9B316AFE29FD8F147E" blockId="19.[370,538,502,528]" box="[370,538,502,528]" pageId="19" pageNumber="138">Eusuchia indet.</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF92FF9B309CFDEFFD7A14F8" pageId="19" pageNumber="138" type="materials_examined">
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<paragraph id="8B848D53FF92FF9B309CFDEFFD7A14F8" blockId="19.[130,775,559,663]" pageId="19" pageNumber="138">
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EXAMINED MATERIAL. —
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<materialsCitation id="3B53870EFF92FF9B3174FDF0FD7A14F8" ID-GBIF-Occurrence="3053068306" pageId="19" pageNumber="138">
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<collectionCode id="ED2A1596FF92FF9B3174FDF0FE301429" box="[364,421,559,583]" pageId="19" pageNumber="138">FSAC</collectionCode>
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Bouj-410, could be the (?) ninth cervical vertebra;
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<collectionCode id="ED2A1596FF92FF9B30E0FD95FEA7140C" box="[248,306,586,610]" pageId="19" pageNumber="138">FSAC</collectionCode>
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Bouj-1b, anterior? cervical vertebra; 400, posterior dorsal vertebra; 1a, first caudal vertebra; 124, caudal vertebra; 96, two fragments of large dorsal osteoderms; 94, fragment of osteoderm.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF92FF9B309CFD69FD6712E0" pageId="19" pageNumber="138" type="description">
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<paragraph id="8B848D53FF92FF9B309CFD69FE8014A0" blockId="19.[130,776,694,1167]" box="[132,277,694,720]" pageId="19" pageNumber="138">DESCRIPTION</paragraph>
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<paragraph id="8B848D53FF92FF9B309CFD09FD6712E0" blockId="19.[130,776,694,1167]" pageId="19" pageNumber="138">
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Numerous postcranial remains have been found in Gueran.These include five procoelous vertebrae and several fragments of osteoderms.FSAC Bouj-410 is a posterior cervical vertebra (
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<figureCitation id="130091D6FF92FF9B32B6FCCAFD691541" box="[686,764,789,816]" captionStart="FIG" captionStartId="20.[132,143,1407,1424]" captionTargetBox="[132,1455,215,1365]" captionTargetId="figure-294@20.[698,1267,789,1366]" captionTargetPageId="20" captionText="FIG. 7. — Crocodyliformes remains from Gueran: A-J, Eusuchia indet.: A, FSAC BOUJ-410, last cervical (ninth) vertebra in left lateral view; B, FSAC BOUJ- 1b, anterior cervical vertebra in left lateral view; C, FSAC BOUJ-400, posterior dorsal vertebra in left lateral view; D, FSAC BOUJ-1a, first caudal vertebra in left lateral view; E, FSAC BOUJ-124, caudal vertebra in left lateral view; F, FSAC BOUJ-94, osteoderm in dorsal view; G, FSAC BOUJ-96, posterior portion of a dorsal osteoderm; H, FSAC BOUJ-96, lateral portion of a dorsal osteoderm in dorsal views; I, J, Crocodyliformes indet.: I, FSAC BOUJ-355, mid portion of a dentary in dorsal view; J, FSAC BOUJ-406, posterior fragment of a left mandibular ramus in dorsal view; K-P, Gavialoidea indet.: K, FSAC BOUJ-405, portion of a left dentary in dorsal view; L, FSAC BOUJ-402, mid portion of a right maxilla in ventral view; M, N, FSAC BOUJ-404, FSAC BOUJ-403 and FSAC BOUJ-401, anterior (M) and mid portion (N) of left maxilla in ventral views; O, P, FSAC BOUJ-407, posterior portion of a left maxilla in lateral (O) and ventral (P) views. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4605979" httpUri="https://zenodo.org/record/4605979/files/figure.png" pageId="19" pageNumber="138">Fig.7A</figureCitation>
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). It bears a long hypapophysis, and the location of the diapophysis and parapophysis suggests that it could be the ninth cervical vertebra. Bouj-1b is a more anterior cervical (
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<figureCitation id="130091D6FF92FF9B3229FCAAFDEB15E1" box="[561,638,885,911]" captionStart="FIG" captionStartId="20.[132,143,1407,1424]" captionTargetBox="[132,1455,215,1365]" captionTargetId="figure-294@20.[698,1267,789,1366]" captionTargetPageId="20" captionText="FIG. 7. — Crocodyliformes remains from Gueran: A-J, Eusuchia indet.: A, FSAC BOUJ-410, last cervical (ninth) vertebra in left lateral view; B, FSAC BOUJ- 1b, anterior cervical vertebra in left lateral view; C, FSAC BOUJ-400, posterior dorsal vertebra in left lateral view; D, FSAC BOUJ-1a, first caudal vertebra in left lateral view; E, FSAC BOUJ-124, caudal vertebra in left lateral view; F, FSAC BOUJ-94, osteoderm in dorsal view; G, FSAC BOUJ-96, posterior portion of a dorsal osteoderm; H, FSAC BOUJ-96, lateral portion of a dorsal osteoderm in dorsal views; I, J, Crocodyliformes indet.: I, FSAC BOUJ-355, mid portion of a dentary in dorsal view; J, FSAC BOUJ-406, posterior fragment of a left mandibular ramus in dorsal view; K-P, Gavialoidea indet.: K, FSAC BOUJ-405, portion of a left dentary in dorsal view; L, FSAC BOUJ-402, mid portion of a right maxilla in ventral view; M, N, FSAC BOUJ-404, FSAC BOUJ-403 and FSAC BOUJ-401, anterior (M) and mid portion (N) of left maxilla in ventral views; O, P, FSAC BOUJ-407, posterior portion of a left maxilla in lateral (O) and ventral (P) views. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4605979" httpUri="https://zenodo.org/record/4605979/files/figure.png" pageId="19" pageNumber="138">Fig. 7B</figureCitation>
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), but it is not possible to determine its exact location in the vertebral column. FSAC Bouj-400 is an isolated procoelous centrum lacking most of the neural arch (
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<figureCitation id="130091D6FF92FF9B3155FC0BFE0B1581" box="[333,414,980,1007]" captionStart="FIG" captionStartId="20.[132,143,1407,1424]" captionTargetBox="[132,1455,215,1365]" captionTargetId="figure-294@20.[698,1267,789,1366]" captionTargetPageId="20" captionText="FIG. 7. — Crocodyliformes remains from Gueran: A-J, Eusuchia indet.: A, FSAC BOUJ-410, last cervical (ninth) vertebra in left lateral view; B, FSAC BOUJ- 1b, anterior cervical vertebra in left lateral view; C, FSAC BOUJ-400, posterior dorsal vertebra in left lateral view; D, FSAC BOUJ-1a, first caudal vertebra in left lateral view; E, FSAC BOUJ-124, caudal vertebra in left lateral view; F, FSAC BOUJ-94, osteoderm in dorsal view; G, FSAC BOUJ-96, posterior portion of a dorsal osteoderm; H, FSAC BOUJ-96, lateral portion of a dorsal osteoderm in dorsal views; I, J, Crocodyliformes indet.: I, FSAC BOUJ-355, mid portion of a dentary in dorsal view; J, FSAC BOUJ-406, posterior fragment of a left mandibular ramus in dorsal view; K-P, Gavialoidea indet.: K, FSAC BOUJ-405, portion of a left dentary in dorsal view; L, FSAC BOUJ-402, mid portion of a right maxilla in ventral view; M, N, FSAC BOUJ-404, FSAC BOUJ-403 and FSAC BOUJ-401, anterior (M) and mid portion (N) of left maxilla in ventral views; O, P, FSAC BOUJ-407, posterior portion of a left maxilla in lateral (O) and ventral (P) views. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4605979" httpUri="https://zenodo.org/record/4605979/files/figure.png" pageId="19" pageNumber="138">Fig. 7C</figureCitation>
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). The transverse process is high on the centrum, which indicates that it is a dorsal vertebra. A first caudal vertebra with a biconvex centrum is preserved (
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<figureCitation id="130091D6FF92FF9B32B1FBCBFD6F1241" box="[681,762,1044,1071]" captionStart="FIG" captionStartId="20.[132,143,1407,1424]" captionTargetBox="[132,1455,215,1365]" captionTargetId="figure-294@20.[698,1267,789,1366]" captionTargetPageId="20" captionText="FIG. 7. — Crocodyliformes remains from Gueran: A-J, Eusuchia indet.: A, FSAC BOUJ-410, last cervical (ninth) vertebra in left lateral view; B, FSAC BOUJ- 1b, anterior cervical vertebra in left lateral view; C, FSAC BOUJ-400, posterior dorsal vertebra in left lateral view; D, FSAC BOUJ-1a, first caudal vertebra in left lateral view; E, FSAC BOUJ-124, caudal vertebra in left lateral view; F, FSAC BOUJ-94, osteoderm in dorsal view; G, FSAC BOUJ-96, posterior portion of a dorsal osteoderm; H, FSAC BOUJ-96, lateral portion of a dorsal osteoderm in dorsal views; I, J, Crocodyliformes indet.: I, FSAC BOUJ-355, mid portion of a dentary in dorsal view; J, FSAC BOUJ-406, posterior fragment of a left mandibular ramus in dorsal view; K-P, Gavialoidea indet.: K, FSAC BOUJ-405, portion of a left dentary in dorsal view; L, FSAC BOUJ-402, mid portion of a right maxilla in ventral view; M, N, FSAC BOUJ-404, FSAC BOUJ-403 and FSAC BOUJ-401, anterior (M) and mid portion (N) of left maxilla in ventral views; O, P, FSAC BOUJ-407, posterior portion of a left maxilla in lateral (O) and ventral (P) views. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4605979" httpUri="https://zenodo.org/record/4605979/files/figure.png" pageId="19" pageNumber="138">Fig. 7D</figureCitation>
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). The osteoderm fragments have their dorsal surfaces densely ornamented with deep pits (
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<figureCitation id="130091D6FF92FF9B3175FB8BFE311201" box="[365,420,1108,1135]" captionStart="FIG" captionStartId="20.[132,143,1407,1424]" captionTargetBox="[132,1455,215,1365]" captionTargetId="figure-294@20.[698,1267,789,1366]" captionTargetPageId="20" captionText="FIG. 7. — Crocodyliformes remains from Gueran: A-J, Eusuchia indet.: A, FSAC BOUJ-410, last cervical (ninth) vertebra in left lateral view; B, FSAC BOUJ- 1b, anterior cervical vertebra in left lateral view; C, FSAC BOUJ-400, posterior dorsal vertebra in left lateral view; D, FSAC BOUJ-1a, first caudal vertebra in left lateral view; E, FSAC BOUJ-124, caudal vertebra in left lateral view; F, FSAC BOUJ-94, osteoderm in dorsal view; G, FSAC BOUJ-96, posterior portion of a dorsal osteoderm; H, FSAC BOUJ-96, lateral portion of a dorsal osteoderm in dorsal views; I, J, Crocodyliformes indet.: I, FSAC BOUJ-355, mid portion of a dentary in dorsal view; J, FSAC BOUJ-406, posterior fragment of a left mandibular ramus in dorsal view; K-P, Gavialoidea indet.: K, FSAC BOUJ-405, portion of a left dentary in dorsal view; L, FSAC BOUJ-402, mid portion of a right maxilla in ventral view; M, N, FSAC BOUJ-404, FSAC BOUJ-403 and FSAC BOUJ-401, anterior (M) and mid portion (N) of left maxilla in ventral views; O, P, FSAC BOUJ-407, posterior portion of a left maxilla in lateral (O) and ventral (P) views. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4605979" httpUri="https://zenodo.org/record/4605979/files/figure.png" pageId="19" pageNumber="138">Fig.7</figureCitation>
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F-H). Bouj-96 is a fragment of large and thick osteoderm with a smooth anterior articular surface.
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</paragraph>
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</subSubSection>
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</treatment>
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</subSection>
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</document>
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