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<document id="EBA6189484D797AF7CC5D6257CB84557" ID="10.11646/zootaxa.4231.4.11SLASH10147" ID-CLB-Dataset="36552" ID-DOI="10.11646/zootaxa.4231.4.11" ID-GBIF-Dataset="335b9ae0-f347-4812-abb8-fe290a0b7630" ID-ISSN="1175-5326" ID-Zenodo-Dep="292650" ID-ZooBank="6001304A-3DAA-4F35-A017-75B146AE4814" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487062712267" checkinUser="plazi" docAuthor="Bramuzzo, Simone &amp; Nel, André" docDate="2017" docId="038AD85EFFA9865D9DD79E7AFA86F94A" docLanguage="en" docName="zootaxa.4231.4.11SLASH10147.pdf" docOrigin="Zootaxa 4231 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microphorites erikai Bramuzzo &amp; Nel, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="593" masterDocId="FFB3A026FFA9865E9D409B42FFB4FFC9" masterDocTitle="Youngest representative of the extinct genus Microphorites in the Eocene amber of France (Diptera: Dolichopodidae: Microphorinae)" masterLastPageNumber="594" masterPageNumber="590" pageNumber="590" updateTime="1698436887991" updateUser="ExternalLinkService">
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<mods:title id="E1C4BC7510C4F06F6B780A2C32120599">Youngest representative of the extinct genus Microphorites in the Eocene amber of France (Diptera: Dolichopodidae: Microphorinae)</mods:title>
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<mods:namePart id="61AC59E6FA1C4B8DE1A58164129AF545">Bramuzzo, Simone</mods:namePart>
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<mods:namePart id="8CB996324985DDDFDA379860931CB750">Nel, André</mods:namePart>
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<mods:date id="AC8B87D57999B7BA3656992F46CCBEF9">2017</mods:date>
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<treatment id="038AD85EFFA9865D9DD79E7AFA86F94A" ID-DOI="http://doi.org/10.5281/zenodo.6051221" ID-GBIF-Taxon="127695361" ID-Zenodo-Dep="6051221" LSID="urn:lsid:plazi:treatment:038AD85EFFA9865D9DD79E7AFA86F94A" httpUri="http://treatment.plazi.org/id/038AD85EFFA9865D9DD79E7AFA86F94A" lastPageId="3" lastPageNumber="593" pageId="0" pageNumber="590">
<subSubSection id="C3393AC3FFA9865E9DD79E7AFDC0FA9B" box="[151,628,1336,1362]" pageId="0" pageNumber="590" type="nomenclature">
<paragraph id="8B9C6948FFA9865E9DD79E7AFDC0FA9B" blockId="0.[151,1436,1336,2030]" box="[151,628,1336,1362]" pageId="0" pageNumber="590">
<emphasis id="B957B55AFFA9865E9DD79E7AFE40FA9B" bold="true" box="[151,500,1336,1362]" pageId="0" pageNumber="590">
<taxonomicName id="4C2312CBFFA9865E9DD79E7AFE3BFA9B" ID-CoL="9HM3S" box="[151,399,1336,1362]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="590" phylum="Arthropoda" rank="species" species="erikai" status="sp. nov.">
<emphasis id="B957B55AFFA9865E9DD79E7AFE3BFA9B" bold="true" box="[151,399,1336,1362]" italics="true" pageId="0" pageNumber="590">Microphorites erikai</emphasis>
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<taxonomicNameLabel id="A2640821FFA9865E9CD69E7AFE40FA9B" box="[406,500,1336,1362]" pageId="0" pageNumber="590" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131875CDFFA9865E9F439E7AFDD8FA9B" box="[515,620,1336,1362]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 4" captionStartId-0="1.[151,244,1969,1991]" captionStartId-1="2.[151,244,1191,1213]" captionStartId-2="3.[151,244,1064,1086]" captionTargetBox-0="[151,1437,789,1945]" captionTargetBox-1="[164,1424,191,1170]" captionTargetBox-2="[150,1436,191,1043]" captionTargetId-0="figure@1.[151,1437,789,1945]" captionTargetId-1="figure@2.[164,1424,191,1170]" captionTargetId-2="figure@3.[150,1436,191,1043]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Microphorites erikai sp. nov., holotype PA 979, photograph of habitus. Scale bar = 1 mm." captionText-1="FIGURE 2. Microphorites erikai sp. nov., holotype PA 979, photograph of head. Scale bar = 0.5 mm." captionText-2="FIGURE 4. Microphorites erikai sp. nov., holotype PA 979, photograph of wing base, arrow indicate series of setae on posterior margin. Scale bar = 0.5 mm." httpUri-0="https://zenodo.org/record/292651/files/figure.png" httpUri-1="https://zenodo.org/record/292652/files/figure.png" httpUri-2="https://zenodo.org/record/292653/files/figure.png" pageId="0" pageNumber="590">
Figs 1
<docIssue id="CF4016A7FFA9865E9F1B9E7AFDD8FA9B" box="[603,620,1336,1362]" pageId="0" pageNumber="590">4</docIssue>
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)
</paragraph>
</subSubSection>
<subSubSection id="C3393AC3FFA9865E9DD79E19FD18F9EB" pageId="0" pageNumber="590" type="diagnosis">
<paragraph id="8B9C6948FFA9865E9DD79E19FD18F9EB" blockId="0.[151,1436,1336,2030]" pageId="0" pageNumber="590">
<emphasis id="B957B55AFFA9865E9DD79E19FEB8FABB" bold="true" box="[151,268,1371,1394]" pageId="0" pageNumber="590">Diagnosis.</emphasis>
Head large, very long antenna, with prolonged postpedicel, broader at base, and tapered distally; postpedicel 1.2 times longer than basal and apical articles of arista-like stylus; thorax higher than longer, strongly arched; scutellum with a pair of long apical setae plus two pairs of shorter setae; no long prescutellar setae; one long seta in notopleuron; wing with costa ending at junction with R4+5; Sc weakened towards costa; distal half of subcostal cell darkened; dark pterostigma present at fusion of first radial vein and costa; crossvein dm-cu arched into cell dm; anal lobe of wing not developed, with long setae at base of anal margin.
</paragraph>
</subSubSection>
<subSubSection id="C3393AC3FFA9865F9D879D6FFD05FE96" lastPageId="1" lastPageNumber="591" pageId="0" pageNumber="590" type="description">
<paragraph id="8B9C6948FFA9865E9D879D6FFD7CF8DD" blockId="0.[151,1436,1336,2030]" pageId="0" pageNumber="590">
<emphasis id="B957B55AFFA9865E9D879D6FFE07F98D" bold="true" box="[199,435,1581,1604]" pageId="0" pageNumber="590">Description. Female.</emphasis>
Head rather large (
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long,
<quantity id="4CDBC4ADFFA9865E9E6E9D6FFC3BF98D" box="[814,911,1581,1604]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="0" pageNumber="590" unit="mm" value="0.74">0.74 mm</quantity>
high), eyes bare, dichoptic; facets of equal size; ocellar triangle large, three ocelli well developed; anterior pair of ocellar setae long, followed by pair of minute postocellar setae; antennae situated near middle of head; antenna long (length
<quantity id="4CDBC4ADFFA9865E9EAD9D31FBE6F943" box="[1005,1106,1651,1674]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="0" pageNumber="590" unit="mm" value="0.58">0.58 mm</quantity>
); scape and pedicel as long as wide, with ring of strong setae present on pedicel; postpedicel elongate,
<quantity id="4CDBC4ADFFA9865E9EDC9DD7FC4FF965" box="[924,1019,1685,1708]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="0" pageNumber="590" unit="mm" value="0.28">0.28 mm</quantity>
long, broader at base, tapering distally and densely covered by rather long microtrichia; arista-like stylus
<quantity id="4CDBC4ADFFA9865E9EC29DFAFC51F906" box="[898,997,1720,1743]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="0" pageNumber="590" unit="mm" value="0.22">0.22 mm</quantity>
long, slightly shorter than postpedicel, composed of two articles, basal article very small and inconspicuous; stylus covered in short microtrichia; labellum and maxillary palpus visible, with setae; face not visible.
</paragraph>
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Thorax higher than longer (length
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, height
<quantity id="4CDBC4ADFFA9865E9E699C62FC26F8FE" box="[809,914,1824,1847]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="0" pageNumber="590" unit="mm" value="0.94">0.94 mm</quantity>
), strongly arched; acrostichals tri- to biserial posteriorly, dorsocentrals uniserial, without two very long prescutellar setae; one long notopleural seta. Halter long (length
<quantity id="4CDBC4ADFFA9865E9DB09C27FEE3F8B5" box="[240,343,1893,1916]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="0" pageNumber="590" unit="mm" value="0.32">0.32 mm</quantity>
) and robust. Wing hyaline,
<quantity id="4CDBC4ADFFA9865E9FE29C27FCBCF8B5" box="[674,776,1893,1916]" metricMagnitude="-3" metricUnit="m" metricValue="2.11" pageId="0" pageNumber="590" unit="mm" value="2.11">2.11 mm</quantity>
long,
<quantity id="4CDBC4ADFFA9865E9E199C27FC0BF8B5" box="[857,959,1893,1916]" metricMagnitude="-4" metricUnit="m" metricValue="9.7" pageId="0" pageNumber="590" unit="mm" value="0.97">0.97 mm</quantity>
wide, entirely covered with microtrichia. Pterostigma (Pt) long, occupying basal 1/3 of costal section between R1 and R2+3; alula not visible; costa (C) uniformly covered with smaller setae; subcosta (Sc) closely approximated with R1, with distal half very faint, ending at costal margin, veins R2+3 and R4+5 diverging in basal half, parallel in distal half, both simple; origin of radial veins basal, at level of humeral crossvein (h); crossvein r-m in basal fourth of wing; discal medial cell (dm) anteriorly tapered, emitting three veins, i.e. M1, M2 and CuA1 ending in wing margin; crossvein bm-cu complete; basal part of M2 straight, basal medial cell (bm) more or less triangular, anal (cup) cell rounded apically.
</paragraph>
<paragraph id="8B9C6948FFA8865F9D879A40FCE5FED0" blockId="1.[151,1436,152,735]" box="[199,849,258,281]" pageId="1" pageNumber="591">Legs long and slender, simple, covered uniformly with setae.</paragraph>
<paragraph id="8B9C6948FFA8865F9D879A67FD05FE96" blockId="1.[151,1436,152,735]" pageId="1" pageNumber="591">
Abdomen cylindrical,
<quantity id="4CDBC4ADFFA8865F9CF79A67FDBDFEF5" box="[439,521,293,316]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="591" unit="mm" value="1.2">1.2 mm</quantity>
long, with long setae, other details not visible; female terminalia telescopic, relaxed, covered with setae, without acanthophorite spines.
</paragraph>
</subSubSection>
<subSubSection id="C3393AC3FFA8865F9D879A28FD30FDC5" pageId="1" pageNumber="591" type="materials_examined">
<paragraph id="8B9C6948FFA8865F9D879A28FB22FE48" blockId="1.[151,1436,152,735]" box="[199,1174,362,385]" pageId="1" pageNumber="591">
<materialsCitation id="3B4B6315FFA8865F9D879A28FB22FE48" ID-GBIF-Occurrence="1434647862" box="[199,1174,362,385]" collectorName="Farm Le Quesnoy &amp; Chevriere &amp; Creil &amp; Oise" country="France" location="France" pageId="1" pageNumber="591" specimenCount="1" typeStatus="holotype">
<emphasis id="B957B55AFFA8865F9D879A28FEE8FE48" bold="true" box="[199,348,362,385]" pageId="1" pageNumber="591">
<typeStatus id="5498D7EAFFA8865F9D879A28FF4BFE48" box="[199,255,362,385]" pageId="1" pageNumber="591">Type</typeStatus>
locality.
</emphasis>
<collectorName id="26D60C9EFFA8865F9C249A28FD92FE48" box="[356,550,362,385]" pageId="1" pageNumber="591">Farm Le Quesnoy</collectorName>
,
<collectorName id="26D60C9EFFA8865F9F709A28FD28FE48" box="[560,668,362,385]" pageId="1" pageNumber="591">Chevrière</collectorName>
, near
<collectorName id="26D60C9EFFA8865F9F9B9A28FCA7FE48" box="[731,787,362,385]" pageId="1" pageNumber="591">Creil</collectorName>
,
<collectorName id="26D60C9EFFA8865F9E5E9A28FCFBFE48" box="[798,847,362,385]" pageId="1" pageNumber="591">Oise</collectorName>
department (northern
<collectingCountry id="F33429D8FFA8865F997D9A28FB3FFE48" box="[1085,1163,362,385]" name="France" pageId="1" pageNumber="591">France</collectingCountry>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9C6948FFA8865F9D879ACFFF68FE0E" blockId="1.[151,1436,152,735]" pageId="1" pageNumber="591">
<emphasis id="B957B55AFFA8865F9D879ACFFED7FE6D" bold="true" box="[199,355,397,420]" pageId="1" pageNumber="591">Type horizon.</emphasis>
Lowermost Eocene, Sparnacian (ca. - 53 Ma), level MP7 of the mammal fauna of Dormaal (
<bibRefCitation id="EFB214B9FFA8865F987A9ACFFF65FE0E" author="Nel" pageId="1" pageNumber="591" refString="Nel, A., de Ploeg, G., Dejax, J., Dutheil, D., de Franceschi, D., Gheerbrant, E., Godinot, M., Hervet, S., Menier, J-J., Auge, M., Bignot, G., Cavagnetto, C., Duffaud, S., Gaudant, J., Hua S., Jossang, A., de Lapparent de Broin, F., Pozzi, J. - P., Paicheler, J. - C., Bouchet, F. &amp; Rage, J. - C. (1999) Un gisement sparnacien exceptionnel a plantes, arthropodes et vertebres (Eocene basal, MP 7): Le Quesnoy (Oise, France). Comptes Rendus de l'Academie des Sciences, Sciences de la terre et des planetes, 329, 65 - 72. https: // doi. org / 10.1016 / s 1251 - 8050 (99) 80229 - 8" type="journal article" year="1999">Nel et al. 1999</bibRefCitation>
).
</paragraph>
<paragraph id="8B9C6948FFA8865F9D879A90FD30FDC5" blockId="1.[151,1436,152,735]" pageId="1" pageNumber="591">
<materialsCitation id="3B4B6315FFA8865F9D879A90FD30FDC5" ID-GBIF-Occurrence="1434647861" collectionCode="MNHN" country="France" location="Paris" pageId="1" pageNumber="591" specimenCount="1" typeStatus="holotype">
<emphasis id="B957B55AFFA8865F9D879A90FE11FE20" bold="true" box="[199,421,466,489]" pageId="1" pageNumber="591">Material examined.</emphasis>
<typeStatus id="5498D7EAFFA8865F9CEA9A90FDB9FE20" box="[426,525,466,489]" pageId="1" pageNumber="591" type="holotype">Holotype</typeStatus>
PA 979, stored in the Paleontological Department,
<collectionCode id="ED32F18DFFA8865F996D9A90FB35FE20" box="[1069,1153,466,489]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="591">MNHN</collectionCode>
,
<location id="8EFC3F93FFA8865F99CB9A90FB77FE20" LSID="urn:lsid:plazi:treatment:038AD85EFFA9865D9DD79E7AFA86F94A:8EFC3F93FFA8865F99CB9A90FB77FE20" box="[1163,1219,466,489]" country="France" name="Paris" pageId="1" pageNumber="591">Paris</location>
,
<collectingCountry id="F33429D8FFA8865F998D9A90FAADFE20" box="[1229,1305,466,489]" name="France" pageId="1" pageNumber="591">France</collectingCountry>
; in the same piece of amber with a large wasp and a spider.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3393AC3FFA8865F9D87995AFC09FDE6" box="[199,957,536,559]" pageId="1" pageNumber="591" type="etymology">
<paragraph id="8B9C6948FFA8865F9D87995AFC09FDE6" blockId="1.[151,1436,152,735]" box="[199,957,536,559]" pageId="1" pageNumber="591">
<emphasis id="B957B55AFFA8865F9D87995AFEF3FDE6" bold="true" box="[199,327,536,559]" pageId="1" pageNumber="591">Etymology.</emphasis>
Named after Erika Degasperi, girlfriend of the first author.
</paragraph>
</subSubSection>
<subSubSection id="C3393AC3FFA8865D9D879978FA86F94A" lastPageId="3" lastPageNumber="593" pageId="1" pageNumber="591" type="discussion">
<paragraph id="8B9C6948FFA8865F9D879978FB0AFD16" blockId="1.[151,1436,152,735]" pageId="1" pageNumber="591">
<emphasis id="B957B55AFFA8865F9D879978FEF2FD98" bold="true" box="[199,326,570,593]" pageId="1" pageNumber="591">Discussion.</emphasis>
The generic diagnosis by
<bibRefCitation id="EFB214B9FFA8865F9F249978FC16FD98" author="Grimaldi" box="[612,930,570,593]" pageId="1" pageNumber="591" refString="Grimaldi, D. A. &amp; Cumming, J. M. (1999) Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History, 239, 1 - 124." type="journal article" year="1999">Grimaldi &amp; Cumming (1999)</bibRefCitation>
based on
<bibRefCitation id="EFB214B9FFA8865F99519978FB04FD98" author="Hennig" box="[1041,1200,570,593]" pageId="1" pageNumber="591" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (&quot; Microphorinae &quot;) aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig (1971)</bibRefCitation>
, with supplements by
<bibRefCitation id="EFB214B9FFA8865F9DD7991FFEE6FDBD" box="[151,338,604,628]" pageId="1" pageNumber="591" refString="Nel, A., Perrichot, V., Daugeron, C. &amp; Neraudeau, D. (2004) A new Microphorites in the Lower Cretaceous amber of the southwest of France (Diptera: Dolichopodidae, &quot; Microphorinae &quot;). Annales de la Societe Entomologique de France, New Series, 40, 23 - 29." type="journal article">
Nel
<emphasis id="B957B55AFFA8865F9D87991FFF43FDBD" box="[199,247,604,628]" italics="true" pageId="1" pageNumber="591">et al</emphasis>
. (2004)
</bibRefCitation>
, is followed. Our fossil has the typical characters for
<taxonomicName id="4C2312CBFFA8865F9EE7991EFB8AFDBD" box="[935,1086,604,628]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="genus">
<emphasis id="B957B55AFFA8865F9EE7991EFB8AFDBD" box="[935,1086,604,628]" italics="true" pageId="1" pageNumber="591">Microphorites</emphasis>
</taxonomicName>
, e.g., antennae located near the middle of anterior margins of the eyes; arista 2-articled; anal margin of wing not making a strong curvature; three radial veins; crossvein r-m in basal fourth of wing; crossvein bm-cu complete; cell dm present; two medial veins and vein CuA1 complete; absence of acanthophorite spines on tergite 10 of the female; female abdomen telescopic.
</paragraph>
<caption id="DF5C39C0FFA8865F9DD79CF3FBC6F80F" box="[151,1138,1969,1991]" httpUri="https://zenodo.org/record/292651/files/figure.png" pageId="1" pageNumber="591" targetBox="[151,1437,789,1945]" targetPageId="1">
<paragraph id="8B9C6948FFA8865F9DD79CF3FBC6F80F" blockId="1.[151,1138,1969,1991]" box="[151,1138,1969,1991]" pageId="1" pageNumber="591">
<emphasis id="B957B55AFFA8865F9DD79CF3FEB9F80E" bold="true" box="[151,269,1969,1991]" pageId="1" pageNumber="591">FIGURE 1.</emphasis>
<taxonomicName id="4C2312CBFFA8865F9C529CF3FE6CF80E" box="[274,472,1969,1991]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="species" species="erikai" status="sp. nov.">
<emphasis id="B957B55AFFA8865F9C529CF3FE6CF80E" box="[274,472,1969,1991]" italics="true" pageId="1" pageNumber="591">Microphorites erikai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2640821FFA8865F9C9E9CF3FD9EF80E" box="[478,554,1969,1991]" pageId="1" pageNumber="591" rank="species">
<emphasis id="B957B55AFFA8865F9C9E9CF3FD9EF80E" bold="true" box="[478,554,1969,1991]" pageId="1" pageNumber="591">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype PA 979, photograph of habitus. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="DF5C39C0FFAB865C9DD79FE5FBDFFB74" box="[151,1131,1191,1213]" httpUri="https://zenodo.org/record/292652/files/figure.png" pageId="2" pageNumber="592" targetBox="[164,1424,191,1170]" targetPageId="2">
<paragraph id="8B9C6948FFAB865C9DD79FE5FBDFFB74" blockId="2.[151,1131,1191,1213]" box="[151,1131,1191,1213]" pageId="2" pageNumber="592">
<emphasis id="B957B55AFFAB865C9DD79FE5FEB8FB74" bold="true" box="[151,268,1191,1213]" pageId="2" pageNumber="592">FIGURE 2.</emphasis>
<taxonomicName id="4C2312CBFFAB865C9C529FE5FE6CFB74" box="[274,472,1191,1213]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="592" phylum="Arthropoda" rank="species" species="erikai" status="sp. nov.">
<emphasis id="B957B55AFFAB865C9C529FE5FE6CFB74" box="[274,472,1191,1213]" italics="true" pageId="2" pageNumber="592">Microphorites erikai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2640821FFAB865C9C9E9FE5FD9EFB74" box="[478,554,1191,1213]" pageId="2" pageNumber="592" rank="species">
<emphasis id="B957B55AFFAB865C9C9E9FE5FD9EFB74" bold="true" box="[478,554,1191,1213]" pageId="2" pageNumber="592">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype PA 979, photograph of head. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B9C6948FFAB865C9DD79CFAFAA5F822" blockId="2.[151,1436,1976,2031]" pageId="2" pageNumber="592">
<emphasis id="B957B55AFFAB865C9DD79CFAFEB8F807" bold="true" box="[151,268,1976,1998]" pageId="2" pageNumber="592">FIGURE 3.</emphasis>
<taxonomicName id="4C2312CBFFAB865C9C529CFAFE6CF807" box="[274,472,1976,1998]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="592" phylum="Arthropoda" rank="species" species="erikai" status="sp. nov.">
<emphasis id="B957B55AFFAB865C9C529CFAFE6CF807" box="[274,472,1976,1998]" italics="true" pageId="2" pageNumber="592">Microphorites erikai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2640821FFAB865C9C9D9CFAFD9EF807" box="[477,554,1976,1998]" pageId="2" pageNumber="592" rank="species">
<emphasis id="B957B55AFFAB865C9C9D9CFAFD9EF807" bold="true" box="[477,554,1976,1998]" pageId="2" pageNumber="592">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="5498D7EAFFAB865C9F769CFAFD3EF804" box="[566,650,1976,1997]" pageId="2" pageNumber="592" type="holotype">holotype</typeStatus>
PA 979, photograph of wing. Abbreviations: h, humeral vein; Pt, pterostigma; Sc, subcostal vein; R1, R2+3, R4+5, branches of radius; M1, M2, branches of media; CuA, anterior cubital. Scale bar =
<quantity id="4CDBC4ADFFAB865C99819C94FABAF822" box="[1217,1294,2006,2027]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="592" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
<caption id="DF5C39C0FFAA865D9DD79F6AFE1DFB92" httpUri="https://zenodo.org/record/292653/files/figure.png" pageId="3" pageNumber="593" targetBox="[150,1436,191,1043]" targetPageId="3">
<paragraph id="8B9C6948FFAA865D9DD79F6AFE1DFB92" blockId="3.[151,1435,1064,1115]" pageId="3" pageNumber="593">
<emphasis id="B957B55AFFAA865D9DD79F6AFEBBFBF7" bold="true" box="[151,271,1064,1086]" pageId="3" pageNumber="593">FIGURE 4.</emphasis>
<taxonomicName id="4C2312CBFFAA865D9C579F6AFE54FBF7" box="[279,480,1064,1086]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="erikai" status="sp. nov.">
<emphasis id="B957B55AFFAA865D9C579F6AFE54FBF7" box="[279,480,1064,1086]" italics="true" pageId="3" pageNumber="593">Microphorites erikai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2640821FFAA865D9CA89F6AFD83FBF7" box="[488,567,1064,1086]" pageId="3" pageNumber="593" rank="species">
<emphasis id="B957B55AFFAA865D9CA89F6AFD83FBF7" bold="true" box="[488,567,1064,1086]" pageId="3" pageNumber="593">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype PA 979, photograph of wing base, arrow indicate series of setae on posterior margin. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B9C6948FFAA865D9D879FC4FDE8FAE1" blockId="3.[151,1436,1157,1667]" pageId="3" pageNumber="593">
Among the fossils currently attributed to the genus
<taxonomicName id="4C2312CBFFAA865D9FBD9FC7FB92FB54" authority="Macquart, 1827" authorityName="Macquart" authorityYear="1827" box="[765,1062,1157,1181]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="B957B55AFFAA865D9FBD9FC7FCDBFB54" box="[765,879,1157,1181]" italics="true" pageId="3" pageNumber="593">Microphor</emphasis>
Macquart, 1827
</taxonomicName>
, our fossil differs from the poorly known
<taxonomicName id="4C2312CBFFAA865D9DA99FEAFD16FB09" authority="Handlirsch, 1910" authorityName="Handlirsch" authorityYear="1910" box="[233,674,1192,1216]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="defunctus">
<emphasis id="B957B55AFFAA865D9DA99FEAFE7FFB09" box="[233,459,1192,1216]" italics="true" pageId="3" pageNumber="593">Microphor defunctus</emphasis>
(
<bibRefCitation id="EFB214B9FFAA865D9C9C9FEBFD2EFB09" author="Handlirsch" box="[476,666,1193,1216]" pageId="3" pageNumber="593" refString="Handlirsch, A. (1910) Canadian Fossil Insects. Insects from the Tertiary lake deposits of the southern interior of British Columbia, collected by Mr. Lawrence M. Lambe, in 1906. Contributions to Canadian Palaeontology, Volume 2. Canada Department of Mines, Geological Survey Branch, Memoir, 12 - P, 93 - 129." type="journal article" year="1910">Handlirsch, 1910</bibRefCitation>
)
</taxonomicName>
in the smaller wings (
<quantity id="4CDBC4ADFFAA865D9EDC9FEBFC4AFB09" box="[924,1022,1193,1216]" metricMagnitude="-3" metricUnit="m" metricValue="2.11" pageId="3" pageNumber="593" unit="mm" value="2.11">2.11 mm</quantity>
long instead of 4.0 mm) (
<bibRefCitation id="EFB214B9FFAA865D98659FEBFF64FB2A" author="Handlirsch" pageId="3" pageNumber="593" refString="Handlirsch, A. (1910) Canadian Fossil Insects. Insects from the Tertiary lake deposits of the southern interior of British Columbia, collected by Mr. Lawrence M. Lambe, in 1906. Contributions to Canadian Palaeontology, Volume 2. Canada Department of Mines, Geological Survey Branch, Memoir, 12 - P, 93 - 129." type="journal article" year="1910">Handlirsch 1910</bibRefCitation>
).
<taxonomicName id="4C2312CBFFAA865D9DA69F89FDCAFB2A" authority="Meunier, 1902" authorityName="Meunier" authorityYear="1902" box="[230,638,1227,1251]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="eocenica">
<emphasis id="B957B55AFFAA865D9DA69F89FE77FB2A" box="[230,451,1227,1251]" italics="true" pageId="3" pageNumber="593">Microphor eocenica</emphasis>
(
<bibRefCitation id="EFB214B9FFAA865D9C959F8EFDCCFB2A" author="Meunier" box="[469,632,1228,1251]" pageId="3" pageNumber="593" refString="Meunier, F. (1902) Etude de quelques Dipteres de l'ambre. Annales de Science Naturelle, Zoologie, Series 8, 16, 395 - 406." type="journal article" year="1902">Meunier, 1902</bibRefCitation>
)
</taxonomicName>
has a short postpedicel and wing length
<quantity id="4CDBC4ADFFAA865D99129F8EFB0CFB2A" box="[1106,1208,1228,1251]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="593" unit="mm" value="0.75">0.75 mm</quantity>
, unlike in our fossil (
<bibRefCitation id="EFB214B9FFAA865D9DDE9FACFE8EFACC" author="Meunier" box="[158,314,1262,1285]" pageId="3" pageNumber="593" refString="Meunier, F. (1902) Etude de quelques Dipteres de l'ambre. Annales de Science Naturelle, Zoologie, Series 8, 16, 395 - 406." type="journal article" year="1902">Meunier 1902</bibRefCitation>
: 402; 1908).
<taxonomicName id="4C2312CBFFAA865D9C869FAFFCFCFACC" authority="Meunier, 1908" authorityName="Meunier" authorityYear="1908" box="[454,840,1261,1285]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="rusticus">
<emphasis id="B957B55AFFAA865D9C869FAFFD20FACC" box="[454,660,1261,1285]" italics="true" pageId="3" pageNumber="593">Microphor rusticus</emphasis>
(
<bibRefCitation id="EFB214B9FFAA865D9FE19FACFCF5FACC" author="Meunier" box="[673,833,1262,1285]" pageId="3" pageNumber="593" refString="Meunier, F. (1908) Monographie des Empidae de l'ambre de la Baltique. Annales de Science Naturelle, Zoologie, Series 9, 7, 81 - 135." type="journal article" year="1908">Meunier, 1908</bibRefCitation>
)
</taxonomicName>
has an elongate postpedicel, but not pear-shaped unlike in our fossil (
<bibRefCitation id="EFB214B9FFAA865D9C649E53FDFBFAE1" author="Meunier" box="[292,591,1297,1320]" pageId="3" pageNumber="593" refString="Meunier, F. (1908) Monographie des Empidae de l'ambre de la Baltique. Annales de Science Naturelle, Zoologie, Series 9, 7, 81 - 135." type="journal article" year="1908">Meunier 1908: pl. 6, fig. 12</bibRefCitation>
).
</paragraph>
<paragraph id="8B9C6948FFAA865D9D879E76FB72FA31" blockId="3.[151,1436,1157,1667]" pageId="3" pageNumber="593">
Within the genus
<taxonomicName id="4C2312CBFFAA865D9CC89E71FDABFA82" box="[392,543,1331,1355]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="B957B55AFFAA865D9CC89E71FDABFA82" box="[392,543,1331,1355]" italics="true" pageId="3" pageNumber="593">Microphorites</emphasis>
</taxonomicName>
, the new fossil differs from all species except
<taxonomicName id="4C2312CBFFAA865D99629E76FB02FA82" box="[1058,1206,1332,1355]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B957B55AFFAA865D99629E76FB02FA82" box="[1058,1206,1332,1355]" italics="true" pageId="3" pageNumber="593">M. moravicus</emphasis>
</taxonomicName>
by the presence of a pterostigma. It differs from
<taxonomicName id="4C2312CBFFAA865D9C829E14FDE1FAA4" box="[450,597,1366,1389]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B957B55AFFAA865D9C829E14FDE1FAA4" box="[450,597,1366,1389]" italics="true" pageId="3" pageNumber="593">M. moravicus</emphasis>
</taxonomicName>
in few characters, viz. in the shape of the scape and pedicel. In
<taxonomicName id="4C2312CBFFAA865D98499E14FA28FAA4" box="[1289,1436,1366,1389]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B957B55AFFAA865D98499E14FA28FAA4" box="[1289,1436,1366,1389]" italics="true" pageId="3" pageNumber="593">M. moravicus</emphasis>
</taxonomicName>
the scape and pedicel are longer than broad, whereas in
<taxonomicName id="4C2312CBFFAA865D9E479E3BFCD8FA59" box="[775,876,1400,1424]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="erikai" status="sp. nov.">
<emphasis id="B957B55AFFAA865D9E479E3BFCD8FA59" box="[775,876,1400,1424]" italics="true" pageId="3" pageNumber="593">M. erikai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2640821FFAA865D9E359E3BFC7AFA59" box="[885,974,1401,1424]" pageId="3" pageNumber="593" rank="species">
<emphasis id="B957B55AFFAA865D9E359E3BFC7AFA59" bold="true" box="[885,974,1401,1424]" pageId="3" pageNumber="593">sp. nov.</emphasis>
</taxonomicNameLabel>
they are broader than long. However, the distinctive shape of the scape and pedicel in
<taxonomicName id="4C2312CBFFAA865D9F309EDEFCB6FA7A" box="[624,770,1436,1459]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B957B55AFFAA865D9F309EDEFCB6FA7A" box="[624,770,1436,1459]" italics="true" pageId="3" pageNumber="593">M. moravicus</emphasis>
</taxonomicName>
could be due to taphonomic deformation in amber. The thorax of
<taxonomicName id="4C2312CBFFAA865D9DF59EFCFEACFA1C" box="[181,280,1469,1493]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="erikai">
<emphasis id="B957B55AFFAA865D9DF59EFCFEACFA1C" box="[181,280,1469,1493]" italics="true" pageId="3" pageNumber="593">M. erikai</emphasis>
</taxonomicName>
is higher than longer, unlike
<taxonomicName id="4C2312CBFFAA865D9F179EFCFD5EFA1C" box="[599,746,1470,1493]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B957B55AFFAA865D9F179EFCFD5EFA1C" box="[599,746,1470,1493]" italics="true" pageId="3" pageNumber="593">M. moravicus</emphasis>
</taxonomicName>
. The head of
<taxonomicName id="4C2312CBFFAA865D9EC39EFCFC51FA1C" box="[899,997,1469,1493]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="erikai">
<emphasis id="B957B55AFFAA865D9EC39EFCFC51FA1C" box="[899,997,1469,1493]" italics="true" pageId="3" pageNumber="593">M. erikai</emphasis>
</taxonomicName>
is larger than that of
<taxonomicName id="4C2312CBFFAA865D99909EFCFAD7FA1C" box="[1232,1379,1470,1493]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B957B55AFFAA865D99909EFCFAD7FA1C" box="[1232,1379,1470,1493]" italics="true" pageId="3" pageNumber="593">M. moravicus</emphasis>
</taxonomicName>
. Our fossil also has long setae at the base of the anal margin of the wing, apparently unlike
<taxonomicName id="4C2312CBFFAA865D996C9EA3FB0BFA31" box="[1068,1215,1505,1528]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B957B55AFFAA865D996C9EA3FB0BFA31" box="[1068,1215,1505,1528]" italics="true" pageId="3" pageNumber="593">M. moravicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C6948FFAA865D9D879D46FA86F94A" blockId="3.[151,1436,1157,1667]" pageId="3" pageNumber="593">
There is some uncertainty concerning the age of the amber in which the
<typeStatus id="5498D7EAFFAA865D99569D46FBF0F9D2" box="[1046,1092,1540,1563]" pageId="3" pageNumber="593">type</typeStatus>
specimen of
<taxonomicName id="4C2312CBFFAA865D99A59D46FAC9F9D2" box="[1253,1405,1540,1563]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B957B55AFFAA865D99A59D46FAC9F9D2" box="[1253,1405,1540,1563]" italics="true" pageId="3" pageNumber="593">M. moravicus</emphasis>
</taxonomicName>
is embedded. The sediment from which it was found is Paleogene in age, but the amber itself is similar to some Late Cretaceous ambers (
<bibRefCitation id="EFB214B9FFAA865D9C3E9D0BFDFFF9A9" author="Tkoc" box="[382,587,1608,1632]" pageId="3" pageNumber="593" refString="Tkoc, M., Nel, A. &amp; Prokop, J. (2016) Discovery of a new species of the Cretaceous genus Microphorites Hennig, 1971 (Diptera: Dolichopodidae s. lat.) in Paleogene amber from eastern Moravia (Czech Republic). Insect Systematics &amp; Evolution, 47, 181 - 193." type="journal article" year="2016">
Tkoč
<emphasis id="B957B55AFFAA865D9C859D0BFE48F9A9" box="[453,508,1608,1632]" italics="true" pageId="3" pageNumber="593">et al</emphasis>
. 2016
</bibRefCitation>
). With the present discovery, we can confirm the presence of the genus
<taxonomicName id="4C2312CBFFAA865D9DD79D29FE9AF94A" box="[151,302,1643,1667]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="B957B55AFFAA865D9DD79D29FE9AF94A" box="[151,302,1643,1667]" italics="true" pageId="3" pageNumber="593">Microphorites</emphasis>
</taxonomicName>
in the Paleogene and consequently this taxon is present through the Cretaceous-Cenozoic crisis.
</paragraph>
</subSubSection>
</treatment>
</document>