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<document id="456564885D88BA21F6CC35020E6E3F2B" ID-CLB-Dataset="217329" ID-DOI="10.37520/aemnp.2022.017" ID-GBIF-Dataset="795cf556-a4a9-477b-b55d-cb80ed8a0f2d" ID-ISSN="1804-6487" ID-Zenodo-Dep="7399702" ID-ZooBank="28D55112-98B1-49A5-B382-58B1B068570B" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1670240514228" checkinUser="felipe" docAuthor="Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas &amp; Solodovnikov, Alexey" docDate="2022" docId="038987A0FFDF4B31B3BDA57817D6FA8A" docLanguage="en" docName="ActaEntMusNatPra.62.1.225-299.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2022.017" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Quedius sundukovi Smetana 2003" docType="treatment" docVersion="8" lastPageNumber="265" masterDocId="FFB0FFD8FFFA4B19B016A005151AFF9F" masterDocTitle="Revision of Quedius sensu stricto (Coleoptera: Staphylinidae)" masterLastPageNumber="299" masterPageNumber="225" pageNumber="262" updateTime="1699412962807" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="DAC63CD561F2C5B6470CEE678813AF94">Revision of Quedius sensu stricto (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="EA397DDD75BC855574451F2840E8F410">Brunke, Adam</mods:namePart>
<mods:affiliation id="AD59CCDF130446E2DD4C0AE9BAB50BD9">Agriculture and Agri-Food Canada, 960 Carling Avenue, Ottawa, Ontario, K</mods:affiliation>
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<mods:affiliation id="969B528F0D5D2DAAEB0B9D700A95298B">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark; e-mails: akhansen @ snm. ku. dk; asolodovnikov @ snm. ku. dk &amp; Zoological Institute, Russian Academy of Science, Universitetskaja nab.</mods:affiliation>
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<heading id="D0D781DAFFDF4B3CB3BDA5781036FA0A" box="[939,1324,1405,1429]" centered="true" fontSize="10" level="2" pageId="37" pageNumber="262" reason="3">
<taxonomicName id="4C204D35FFDF4B3CB3BDA5781036FA0A" ID-CoL="4R2Z9" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[939,1324,1405,1429]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="262" phylum="Arthropoda" rank="species" species="sundukovi">
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<emphasis id="B954EAA4FFDF4B3CB3BDA578119AFA0A" bold="true" box="[939,1152,1405,1429]" italics="true" pageId="37" pageNumber="262">Quedius sundukovi</emphasis>
<bibRefCitation id="EFB14B47FFDF4B3CB491A57B1036FA0A" author="SMETANA A." box="[1159,1324,1406,1429]" pageId="37" pageNumber="262" pagination="189 - 193" refId="ref92269" refString="SMETANA A. 2003: Quedius (Quedius) sundukovi (Coleoptera, Staphylinidae, Staphylinini, Quediina), an interesting new species from the Russian Far East. Elytra 31: 189 - 193." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
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<paragraph id="8B9F36B6FFDF4B3CB3F8A59911F0FA30" blockId="37.[828,1436,1436,1516]" box="[1006,1258,1436,1455]" pageId="37" pageNumber="262">
(
<figureCitation id="131B2A33FFDF4B3CB3E2A5991130FA30" box="[1012,1066,1436,1455]" captionStart="Fig" captionStartId="1.[152,183,2006,2025]" captionTargetBox="[222,1381,1063,1978]" captionTargetId="figure-474@1.[207,1384,1060,1988]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen &amp; Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups." figureDoi="http://doi.org/10.5281/zenodo.7399704" httpUri="https://zenodo.org/record/7399704/files/figure.png" pageId="37" pageNumber="262">Figs 1</figureCitation>
,
<figureCitation id="131B2A33FFDF4B3CB423A599115BFA30" box="[1077,1089,1436,1455]" captionStart="Fig" captionStartId="12.[151,182,2008,2027]" captionTargetBox="[182,1406,199,1981]" captionTargetId="figure-13@12.[182,1407,189,1989]" captionTargetPageId="12" captionText="Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP &lt;0.90, UFB &lt;95 and SH-aLRT &lt;80." figureDoi="http://doi.org/10.5281/zenodo.7399710" httpUri="https://zenodo.org/record/7399710/files/figure.png" pageId="37" pageNumber="262">4</figureCitation>
,
<figureCitation id="131B2A33FFDF4B3CB45CA599114CFA30" box="[1098,1110,1436,1455]" captionStart="Fig" captionStartId="17.[151,182,2023,2042]" captionTargetBox="[180,1386,189,1955]" captionTargetId="figure-13@17.[177,1411,186,1980]" captionTargetPageId="17" captionText="Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961." figureDoi="http://doi.org/10.5281/zenodo.7399720" httpUri="https://zenodo.org/record/7399720/files/figure.png" pageId="37" pageNumber="262">7</figureCitation>
,
<figureCitation id="131B2A33FFDF4B3CB449A599116CFA30" box="[1119,1142,1436,1455]" captionStart="Fig" captionStartId="18.[151,182,2027,2046]" captionTargetBox="[189,1393,195,2007]" captionTargetId="figure-13@18.[174,1413,185,2007]" captionTargetPageId="18" captionText="Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003." figureDoi="http://doi.org/10.5281/zenodo.7399722" httpUri="https://zenodo.org/record/7399722/files/figure.png" pageId="37" pageNumber="262">8F</figureCitation>
,
<figureCitation id="131B2A33FFDF4B3CB469A59911BEFA30" box="[1151,1188,1436,1455]" captionStart="Fig" captionStartId="31.[151,182,2004,2023]" captionTargetBox="[207,1368,203,1953]" captionTargetId="figure-12@31.[191,1396,189,1962]" captionTargetPageId="31" captionText="Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view." figureDoi="http://doi.org/10.5281/zenodo.7399732" httpUri="https://zenodo.org/record/7399732/files/figure.png" pageId="37" pageNumber="262">13B</figureCitation>
,
<figureCitation id="131B2A33FFDF4B3CB4BBA59911DEFA30" box="[1197,1220,1436,1455]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="37" pageNumber="262">15</figureCitation>
,
<figureCitation id="131B2A33FFDF4B3CB4DBA59911F9FA30" box="[1229,1251,1436,1455]" captionStart="Fig" captionStartId="59.[151,182,2028,2047]" captionTargetBox="[190,1392,189,2003]" captionTargetId="figure-13@59.[190,1393,189,2003]" captionTargetPageId="59" captionText="Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399748" httpUri="https://zenodo.org/record/7399748/files/figure.png" pageId="37" pageNumber="262">20</figureCitation>
)
</paragraph>
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<treatmentCitation id="0A8110A7FFDF4B3CB32AA5BB11B4FA4E" author="SMETANA A." box="[828,1198,1470,1489]" page="193" pageId="37" pageNumber="262" year="2003">
<taxonomicName id="4C204D35FFDF4B3CB32AA5BB11B4FA4E" ID-CoL="4R2Z9" authority="Smetana 2003: 189 - 193" authorityName="Smetana" authorityPageNumber="189 - 193" authorityYear="2003" box="[828,1198,1470,1489]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="262" phylum="Arthropoda" rank="species" species="sundukovi">
<emphasis id="B954EAA4FFDF4B3CB32AA5BB16C5FA4E" box="[828,991,1470,1489]" italics="true" pageId="37" pageNumber="262">Quedius sundukovi</emphasis>
<bibRefCitation id="EFB14B47FFDF4B3CB3F5A5BB11B4FA4E" author="SMETANA A." box="[995,1198,1470,1489]" pageId="37" pageNumber="262" pagination="189 - 193" refId="ref92269" refString="SMETANA A. 2003: Quedius (Quedius) sundukovi (Coleoptera, Staphylinidae, Staphylinini, Quediina), an interesting new species from the Russian Far East. Elytra 31: 189 - 193." type="journal article" year="2003">Smetana 2003: 189-193</bibRefCitation>
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[
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locality: Badshalskyi Khrebet mountains]
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<emphasis id="B954EAA4FFDF4B3CB32AA60616BEF989" bold="true" box="[828,932,1539,1558]" pageId="37" pageNumber="262">References.</emphasis>
Sආൾඍൺඇൺ &amp; SHൺඏඋංඇ (2018): 834 (distribution); Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2018a):130–131 (distribution); (2019):50 (characters and distribution).
</paragraph>
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Material examined.
<collectingCountry id="F3377626FFDF4B3CB3F8A667119DF9EA" box="[1006,1159,1634,1653]" name="Kazakhstan" pageId="37" pageNumber="262">KAZAKHSTAN</collectingCountry>
:
</emphasis>
SW
<collectingRegion id="49E4F854FFDF4B3CB4A6A66711C6F9EA" box="[1200,1244,1634,1653]" country="Russia" name="Altay" pageId="37" pageNumber="262">Altai</collectingRegion>
, East of Narymskij Mt. Ridge, upper course of Ozernaja River,
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,
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,
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, supalpine zone,
<date id="FF9E1076FFDF4B3CB3D4A6921136F935" box="[962,1068,1687,1706]" pageId="37" pageNumber="262" value="1997-07-18">18.VII.1997</date>
, leg.R.Yu.Dudko &amp; V.K. Zinchenko (
<collectionCode id="ED31AE73FFDF4B3CB57AA692108EF935" box="[1388,1428,1687,1706]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="37" pageNumber="262" type="Museum">ZIN</collectionCode>
).
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<collectingCountry id="F3377626FFDF4B3CB32AA6B71697F95A" box="[828,909,1714,1733]" name="Russia" pageId="37" pageNumber="262">RUSSIA</collectingCountry>
: JൾඐංඌH Aඎඍඈඇඈආඈඎඌ RൾG.:
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Kuldur vill.,
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,
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, pitfall trap, stream,
<quantity id="4CD89B53FFDF4B3CB3CBA6C81115F97F" box="[989,1039,1741,1760]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="37" pageNumber="262" unit="m" value="480.0">480m</quantity>
,
<date id="FF9E1076FFDF4B3CB401A6C811BBF97F" box="[1047,1185,1741,1760]" pageId="37" pageNumber="262" value="2017-07-14" valueMax="2017-07-15" valueMin="2017-07-14">14.-15.VII.2017</date>
, leg.A.Bergmann (1 J cSch).
<emphasis id="B954EAA4FFDF4B3CB32AA6E216DAF965" bold="true" box="[828,960,1767,1786]" pageId="37" pageNumber="262">Eൺඌඍ YൺKඎඍංൺ:</emphasis>
Suntar-Khayata Mts, Valley of Tyry river valley, Kidyrki river near Khalya, [62.35, 138.46],
<quantity id="4CD89B53FFDF4B3CB462A70711CFF88A" box="[1140,1237,1794,1813]" metricMagnitude="2" metricUnit="m" metricValue="8.75" metricValueMax="9.5" metricValueMin="8.0" pageId="37" pageNumber="262" unit="m" value="875.0" valueMax="950.0" valueMin="800.0">800-950 m</quantity>
,
<date id="FF9E1076FFDF4B3CB4C9A7071068F88A" box="[1247,1394,1794,1813]" pageId="37" pageNumber="262" value="1991-08-14" valueMax="1991-08-19" valueMin="1991-08-14">14.-19.VIII.1991</date>
, leg. Alexeev (
<collectionCode id="ED31AE73FFDF4B3CB399A71816CFF8AF" box="[911,981,1821,1840]" pageId="37" pageNumber="262">NHMD</collectionCode>
).
<emphasis id="B954EAA4FFDF4B3CB3F5A7181184F8AF" bold="true" box="[995,1182,1821,1840]" pageId="37" pageNumber="262">KHൺൻൺඋඈඏඌK Tൾඋඋ.:</emphasis>
Badshalskyi chr., Omot lake, [50.54, 134.26], tundra,
<quantity id="4CD89B53FFDF4B3CB413A732116DF8D5" box="[1029,1143,1847,1866]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.05" metricValueMin="1.85" pageId="37" pageNumber="262" unit="m" value="1950.0" valueMax="2050.0" valueMin="1850.0">1850-2050 m</quantity>
,
<date id="FF9E1076FFDF4B3CB469A73211E5F8D5" box="[1151,1279,1847,1866]" pageId="37" pageNumber="262" value="1997-07-07" valueMax="1997-07-19" valueMin="1997-07-07">7.-19.VII.1997</date>
, leg A.Plutenko (2 JJ
<specimenCount id="9D26FD3FFFDF4B3CB349A7571690F8FB" box="[863,906,1867,1893]" pageId="37" pageNumber="262" type="female">2♀♀</specimenCount>
cSch); Levaja Bureya riv., estuary Don riv., [51.68, 134.51],
<date id="FF9E1076FFDF4B3CB56DA75716BCF81F" pageId="37" pageNumber="262" value="2006-07-24" valueMax="2006-07-25" valueMin="2006-07-24">24.- 25.VII.2006</date>
, leg. U. Valainis (
<collectionCode id="ED31AE73FFDF4B3CB42BA768117FF81F" box="[1085,1125,1901,1920]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="37" pageNumber="262" type="Museum">ZIN</collectionCode>
); Levaja Bureya riv., the left bank of the Imganakh river near the mouth,
<geoCoordinate id="EE145071FFDF4B3CB475A78211B0F805" box="[1123,1194,1927,1946]" degrees="51.7355" direction="north" orientation="latitude" pageId="37" pageNumber="262" precision="5" value="51.7355">51.7355</geoCoordinate>
,
<geoCoordinate id="EE145071FFDF4B3CB4A7A78211E2F805" box="[1201,1272,1927,1946]" degrees="134.572" direction="east" orientation="longitude" pageId="37" pageNumber="262" precision="55" value="134.572">134.572</geoCoordinate>
,
<quantity id="4CD89B53FFDF4B3CB516A7821046F805" box="[1280,1372,1927,1946]" metricMagnitude="2" metricUnit="m" metricValue="6.6" metricValueMax="6.7" metricValueMin="6.5" pageId="37" pageNumber="262" unit="m" value="660.0" valueMax="670.0" valueMin="650.0">650-670 m</quantity>
, sloppy rocky hills with burrow pits, litter and mosses of
<taxonomicName id="4C204D35FFDF4B3CB4CAA7A71079F82A" box="[1244,1379,1954,1973]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="37" pageNumber="262" phylum="Tracheophyta" rank="species" species="ajanensis">
<emphasis id="B954EAA4FFDF4B3CB4CAA7A71079F82A" box="[1244,1379,1954,1973]" italics="true" pageId="37" pageNumber="262">Picea ajanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C204D35FFDF4B3CB57BA7A71684F84F" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="37" pageNumber="262" phylum="Tracheophyta" rank="species" species="nephrolepis">
<emphasis id="B954EAA4FFDF4B3CB57BA7A71684F84F" italics="true" pageId="37" pageNumber="262">Abies nephrolepis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C204D35FFDF4B3CB3B1A7B81107F84F" box="[935,1053,1981,2000]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="37" pageNumber="262" phylum="Tracheophyta" rank="species" species="gmelinii">
<emphasis id="B954EAA4FFDF4B3CB3B1A7B81107F84F" box="[935,1053,1981,2000]" italics="true" pageId="37" pageNumber="262">Larix gmelinii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C204D35FFDF4B3CB433A7B811EFF84F" box="[1061,1269,1981,2000]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="37" pageNumber="262" phylum="Tracheophyta" rank="species" species="dauricum">
<emphasis id="B954EAA4FFDF4B3CB433A7B811EFF84F" box="[1061,1269,1981,2000]" italics="true" pageId="37" pageNumber="262">Rhododendron dauricum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C204D35FFDF4B3CB4E8A7B8108DF84F" box="[1278,1431,1981,2000]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="37" pageNumber="262" phylum="Tracheophyta" rank="species" species="platyphylla">
<emphasis id="B954EAA4FFDF4B3CB4E8A7B8108DF84F" box="[1278,1431,1981,2000]" italics="true" pageId="37" pageNumber="262">Betula platyphylla</emphasis>
</taxonomicName>
,
<taxonomicName id="4C204D35FFDF4B3CB32AA7D2111FF875" box="[828,1029,2007,2026]" class="Bryopsida" family="Hylocomiaceae" genus="Hylocomium" kingdom="Plantae" order="Hypnales" pageId="37" pageNumber="262" phylum="Bryophyta" rank="species" species="splendens">
<emphasis id="B954EAA4FFDF4B3CB32AA7D2111FF875" box="[828,1029,2007,2026]" italics="true" pageId="37" pageNumber="262">Hylocomium splendens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C204D35FFDF4B3CB404A7D211D6F875" authorityName="Mitten" authorityYear="1869" box="[1042,1228,2007,2026]" class="Bryopsida" family="Hylocomiaceae" genus="Pleurozium" kingdom="Plantae" order="Hypnales" pageId="37" pageNumber="262" phylum="Bryophyta" rank="species" species="schreberi">
<emphasis id="B954EAA4FFDF4B3CB404A7D211D6F875" box="[1042,1228,2007,2026]" italics="true" pageId="37" pageNumber="262">Pleurozium schreberi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C204D35FFDF4B3CB4CFA7D216A6F79A" authority=", Dircranum" authorityName="Dircranum" class="Insecta" family="Ptiliidae" genus="Ptilium" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="262" phylum="Arthropoda" rank="species" species="crista-castrensis">
<emphasis id="B954EAA4FFDF4B3CB4CFA7D21649F79A" italics="true" pageId="37" pageNumber="262">Ptilium crista-castrensis</emphasis>
,
<emphasis id="B954EAA4FFDF4B3CB348A7F716A6F79A" box="[862,956,2034,2053]" italics="true" pageId="37" pageNumber="262">Dircranum</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C204D35FFDF4B3CB3F1A7F71168F79A" box="[999,1138,2034,2053]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichum" kingdom="Plantae" order="Polytrichales" pageId="37" pageNumber="262" phylum="Bryophyta" rank="species" species="undetermined">
<emphasis id="B954EAA4FFDF4B3CB3F1A7F7114AF79A" box="[999,1104,2034,2053]" italics="true" pageId="37" pageNumber="262">Polytrichum</emphasis>
sp.
</taxonomicName>
,
<emphasis id="B954EAA4FFDF4B3CB46DA7F7105DF79A" box="[1147,1351,2034,2053]" italics="true" pageId="37" pageNumber="262">Sphagnum girgensohnii</emphasis>
,
<emphasis id="B954EAA4FFDF4B3CB545A7F71063F79A" box="[1363,1401,2034,2053]" italics="true" pageId="37" pageNumber="262">Sph.</emphasis>
sp.,
<date id="FF9E1076FFDF4B3CB32AA8081687F7BF" box="[828,925,2061,2080]" pageId="37" pageNumber="262" value="2011-06-26">26.VI.2011</date>
, leg.A.B. Ryvkin (
<collectionCode id="ED31AE73FFDF4B3CB42FA808117AF7BF" box="[1081,1120,2061,2080]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="37" pageNumber="262" type="Museum">ZIN</collectionCode>
);Verkhnebureinsky natural park, Ust--Urgal project gauging station Ust-Niman Topolevnik,
<geoCoordinate id="EE145071FFDD4B3EB276A0BB17BDFF4E" box="[608,679,190,209]" degrees="51.3995" direction="north" orientation="latitude" pageId="39" pageNumber="264" precision="5" value="51.3995">51.3995</geoCoordinate>
,
<geoCoordinate id="EE145071FFDD4B3EB2B8A0BB17EFFF4E" box="[686,757,190,209]" degrees="132.733" direction="east" orientation="longitude" pageId="39" pageNumber="264" precision="55" value="132.733">132.733</geoCoordinate>
,
<date id="FF9E1076FFDD4B3EB081A0DD15F5FF74" box="[151,239,216,235]" pageId="39" pageNumber="264" value="2009-09-01">1.IX.2009</date>
, leg. L.A. Trilikauskas (
<collectionCode id="ED31AE73FFDD4B3EB1D2A0DD1710FF74" box="[452,522,216,235]" pageId="39" pageNumber="264">NHMD</collectionCode>
); Verkhnebureinsky natural park, Ust-Urgal project gauging station Ust-Niman Topolevnik,
<geoCoordinate id="EE145071FFDD4B3EB2BBA0F617EEFE99" box="[685,756,243,262]" degrees="51.3995" direction="north" orientation="latitude" pageId="39" pageNumber="264" precision="5" value="51.3995">51.3995</geoCoordinate>
,
<geoCoordinate id="EE145071FFDD4B3EB081A10B15C4FEBE" box="[151,222,270,289]" degrees="132.733" direction="east" orientation="longitude" pageId="39" pageNumber="264" precision="55" value="132.733">132.733</geoCoordinate>
,
<quantity id="4CD89B53FFDD4B3EB0FEA10B143BFEBE" box="[232,289,270,289]" metricMagnitude="2" metricUnit="m" metricValue="3.15" pageId="39" pageNumber="264" unit="m" value="315.0">315 m</quantity>
,
<emphasis id="B954EAA4FFDD4B3EB13DA10B1798FEBE" box="[299,642,270,289]" italics="true" pageId="39" pageNumber="264">
<taxonomicName id="4C204D35FFDD4B3EB13DA10B1484FEBE" authority=", Alnus" authorityName="Alnus" box="[299,414,270,289]" class="Magnoliopsida" family="Rosaceae" genus="Padus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="39" pageNumber="264" phylum="Tracheophyta" rank="genus">Padus, Alnus</taxonomicName>
,
<taxonomicName id="4C204D35FFDD4B3EB1B1A10B14CEFEBE" box="[423,468,270,289]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="39" pageNumber="264" phylum="Tracheophyta" rank="genus">Salix</taxonomicName>
,
<taxonomicName id="4C204D35FFDD4B3EB1C8A10B1798FEBE" authority=", Plagiomium" authorityName="Plagiomium" box="[478,642,270,289]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="39" pageNumber="264" phylum="Tracheophyta" rank="genus">Picea, Plagiomium</taxonomicName>
</emphasis>
,
<date id="FF9E1076FFDD4B3EB298A10B17EEFEBE" box="[654,756,270,289]" pageId="39" pageNumber="264" value="2009-08-04">4.VIII.2009</date>
, leg. A.B. Ryvkin (
<collectionCode id="ED31AE73FFDD4B3EB121A12D1445FEA4" box="[311,351,296,315]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="39" pageNumber="264" type="Museum">ZIN</collectionCode>
).
<emphasis id="B954EAA4FFDD4B3EB179A12D170EFEA4" bold="true" box="[367,532,296,315]" pageId="39" pageNumber="264">SൺKHൺඅංඇ Iඌඅൺඇൽ:</emphasis>
<quantity id="4CD89B53FFDD4B3EB20DA12D1749FEA4" box="[539,595,296,315]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="39" pageNumber="264" unit="km" value="13.0">13 km</quantity>
N from Korsakov,
<quantity id="4CD89B53FFDD4B3EB081A14615A7FEC9" box="[151,189,323,342]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="264" unit="km" value="2.0">2km</quantity>
N from Solovyovka, [46.74, 142.73],
<emphasis id="B954EAA4FFDD4B3EB1EDA146177BFEC9" box="[507,609,323,342]" italics="true" pageId="39" pageNumber="264">
<taxonomicName id="4C204D35FFDD4B3EB1EDA1461739FEC9" box="[507,547,323,342]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="39" pageNumber="264" phylum="Tracheophyta" rank="genus">Acer</taxonomicName>
,
<taxonomicName id="4C204D35FFDD4B3EB23CA146177BFEC9" box="[554,609,323,342]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="39" pageNumber="264" phylum="Tracheophyta" rank="genus">Betula</taxonomicName>
</emphasis>
,
<date id="FF9E1076FFDD4B3EB27DA14617CBFEC9" box="[619,721,323,342]" pageId="39" pageNumber="264" value="1992-08-05">5.VIII.1992</date>
, leg. V. Gusarov, (
<collectionCode id="ED31AE73FFDD4B3EB113A15B142FFEEE" box="[261,309,350,369]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="39" pageNumber="264">CNC</collectionCode>
);
<quantity id="4CD89B53FFDD4B3EB155A15B1462FEEE" box="[323,376,350,369]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="39" pageNumber="264" unit="km" value="13.0">13 km</quantity>
N from Korsakov,
<quantity id="4CD89B53FFDD4B3EB20DA15B175FFEEE" box="[539,581,350,369]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="264" unit="km" value="2.0">2 km</quantity>
N from Solovyovka, [46.74, 142.73],
<emphasis id="B954EAA4FFDD4B3EB109A17D171FFE14" box="[287,517,376,395]" italics="true" pageId="39" pageNumber="264">
<taxonomicName id="4C204D35FFDD4B3EB109A17D144AFE14" box="[287,336,376,395]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="39" pageNumber="264" phylum="Tracheophyta" rank="genus">Abies</taxonomicName>
,
<taxonomicName id="4C204D35FFDD4B3EB140A17D14D2FE14" authority=", Ledum" authorityName="Ledum" box="[342,456,376,395]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="39" pageNumber="264" phylum="Tracheophyta" rank="genus">Larix, Ledum</taxonomicName>
,
<taxonomicName id="4C204D35FFDD4B3EB1D9A17D171FFE14" box="[463,517,376,395]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="39" pageNumber="264" phylum="Tracheophyta" rank="genus">Betula</taxonomicName>
</emphasis>
,
<date id="FF9E1076FFDD4B3EB218A17D1769FE14" box="[526,627,376,395]" pageId="39" pageNumber="264" value="1992-08-06">6.VIII.1992</date>
, leg.V.Gusarov (
<collectionCode id="ED31AE73FFDD4B3EB08BA19615D7FE39" box="[157,205,403,422]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="39" pageNumber="264">CNC</collectionCode>
).
<emphasis id="B954EAA4FFDD4B3EB0CDA196148AFE39" bold="true" box="[219,400,403,422]" pageId="39" pageNumber="264">ZൺൻൺඒKൺඅඌKඒ Kඋൺං:</emphasis>
Stanovoy Highlands, W part of Kodar, top Chara River,
<quantity id="4CD89B53FFDD4B3EB11FA1AB1425FE5E" box="[265,319,430,449]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="39" pageNumber="264" unit="km" value="50.0">50 km</quantity>
WSW River Novaya Chara, [56.65, 117.54],
<quantity id="4CD89B53FFDD4B3EB2D0A1AB15C3FE44" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="39" pageNumber="264" unit="m" value="1850.0" valueMax="2000.0" valueMin="1700.0">1700- 2000 m</quantity>
,
<date id="FF9E1076FFDD4B3EB0F7A1CD1451FE44" box="[225,331,456,475]" pageId="39" pageNumber="264" value="1995-07-27">27.VII.1995</date>
, leg.A. &amp; R. Dudko, D. Lomakin (
<collectionCode id="ED31AE73FFDD4B3EB267A1CD1783FE44" box="[625,665,456,475]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="39" pageNumber="264" type="Museum">ZIN</collectionCode>
); Stanovoy Highlands, top Chara River, Lake Leprindo,
<geoCoordinate id="EE145071FFDD4B3EB20CA1E6174DFE69" box="[538,599,483,502]" degrees="56.636" direction="north" orientation="latitude" pageId="39" pageNumber="264" precision="55" value="56.636">56.636</geoCoordinate>
,
<geoCoordinate id="EE145071FFDD4B3EB277A1E617B2FE69" box="[609,680,483,502]" degrees="117.537" direction="east" orientation="longitude" pageId="39" pageNumber="264" precision="55" value="117.537">117.537</geoCoordinate>
,
<quantity id="4CD89B53FFDD4B3EB2A7A1E617EFFE69" box="[689,757,483,502]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="39" pageNumber="264" unit="m" value="1000.0">1000 m</quantity>
,
<date id="FF9E1076FFDD4B3EB081A1FB141BFD8E" box="[151,257,510,529]" pageId="39" pageNumber="264" value="1995-07-23">23.VII.1995</date>
, leg. A. &amp; R. Dudko, D. Lomakin (
<collectionCode id="ED31AE73FFDD4B3EB238A1FB174CFD8E" box="[558,598,510,529]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="39" pageNumber="264" type="Museum">ZIN</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF5F663EFFDC4B3FB081A7B814E5F7BF" ID-DOI="http://doi.org/10.5281/zenodo.7399734" ID-Zenodo-Dep="7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="38" pageNumber="263" startId="38.[151,182,1981,2000]" targetBox="[199,1380,186,1956]" targetPageId="38">
<paragraph id="8B9F36B6FFDC4B3FB081A7B814E5F7BF" blockId="38.[151,1437,1981,2080]" pageId="38" pageNumber="263">
Fig. 14. Aedeagus of
<taxonomicName id="4C204D35FFDC4B3FB145A7B81480F84F" authorityName="Stephens" authorityYear="1829" box="[339,410,1981,2000]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="263" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B954EAA4FFDC4B3FB145A7B81480F84F" box="[339,410,1981,2000]" italics="true" pageId="38" pageNumber="263">Quedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757DFFFDC4B3FB1B7A7B814CAF84F" box="[417,464,1981,2000]" pageId="38" pageNumber="263" sensu="stricto">s. str.</taxonomicNameLabel>
A –
<taxonomicName id="4C204D35FFDC4B3FB1ECA7B81614F84F" authority="Kiesenwetter, 1847" authorityName="Kiesenwetter" authorityYear="1847" box="[506,782,1981,2000]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="263" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B954EAA4FFDC4B3FB1ECA7B81744F84F" box="[506,606,1981,2000]" italics="true" pageId="38" pageNumber="263">Q. unicolor</emphasis>
<bibRefCitation id="EFB14B47FFDC4B3FB273A7B81614F84F" author="KIESENWETTER H. &amp; MARKEL F." box="[613,782,1981,2000]" pageId="38" pageNumber="263" pagination="73 - 87" refId="ref85769" refString="KIESENWETTER H. ඏඈඇ &amp; MARKEL F. 1847: Eine entomologische Excursion im Riesengebirge im Juli 1846 (Fortsetzung). Entomologische Zeitung (Stettin) 8: 73 - 87." type="journal article" year="1847">Kiesenwetter, 1847</bibRefCitation>
</taxonomicName>
; B –
<taxonomicName id="4C204D35FFDC4B3FB328A7B81135F84F" authority="Coiffait, 1964" authorityName=", Coiffait" authorityYear="1964" box="[830,1071,1981,2000]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="263" phylum="Arthropoda" rank="species" species="levasseuri">
<emphasis id="B954EAA4FFDC4B3FB328A7B816AAF84F" box="[830,944,1981,2000]" italics="true" pageId="38" pageNumber="263">Q. levasseuri</emphasis>
<bibRefCitation id="EFB14B47FFDC4B3FB3A1A7B81135F84F" author="COIFFAIT H." box="[951,1071,1981,2000]" pageId="38" pageNumber="263" pagination="293 - 296" refId="ref81616" refString="COIFFAIT H. 1964: Deux nouveaux Quedius s. str. de la region mediterraneenne occidentale. Revue Francaise d'Entomologie 31: 293 - 296." type="journal article" year="1964">Coiffait, 1964</bibRefCitation>
</taxonomicName>
(syn. of
<taxonomicName id="4C204D35FFDC4B3FB468A7B8104BF84F" authority="Lucas, 1846" authorityName="Lucas" authorityYear="1846" box="[1150,1361,1981,2000]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="263" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFDC4B3FB468A7B811C4F84F" box="[1150,1246,1981,2000]" italics="true" pageId="38" pageNumber="263">Q. pallipes</emphasis>
<bibRefCitation id="EFB14B47FFDC4B3FB4F2A7B8104BF84F" author="LUCAS P. H." box="[1252,1361,1981,2000]" pageId="38" pageNumber="263" refId="ref87139" refString="LUCAS P. H. 1846 - 1849: Histoire naturelle des animaux articules. Deuxieme partie. Insectes. In: Exploration scientifique de l'Algerie pendant les annees 1840, 1841, 1842 publiee par ordre du Gouvernement et avec le concours d'une Commission Academique. Sciences physiques. Zoologie II. A. Bertrand, Paris, 590 pp." type="book" year="1846">Lucas, 1846</bibRefCitation>
</taxonomicName>
); C –
<taxonomicName id="4C204D35FFDC4B3FB591A7B81493F874" authority="Bernhauer, 1898" authorityName="Bernhauer" authorityYear="1898" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="263" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFDC4B3FB591A7B815E8F874" italics="true" pageId="38" pageNumber="263">Q. hispanicus</emphasis>
<bibRefCitation id="EFB14B47FFDC4B3FB0EFA7DD1493F874" author="BERNHAUER M." box="[249,393,2008,2027]" pageId="38" pageNumber="263" pagination="338 - 341" refId="ref80314" refString="BERNHAUER M. 1898: Neue Staphyliniden aus Europa. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 48: 338 - 341." type="journal article" year="1898">Bernhauer, 1898</bibRefCitation>
</taxonomicName>
; D –
<taxonomicName id="4C204D35FFDC4B3FB1ACA7DD1790F874" authority="Lucas, 1846" authorityName="Lucas" authorityYear="1846" box="[442,650,2008,2027]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="263" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFDC4B3FB1ACA7DD1703F874" box="[442,537,2008,2027]" italics="true" pageId="38" pageNumber="263">Q. pallipes</emphasis>
<bibRefCitation id="EFB14B47FFDC4B3FB209A7DD1790F874" author="LUCAS P. H." box="[543,650,2008,2027]" pageId="38" pageNumber="263" refId="ref87139" refString="LUCAS P. H. 1846 - 1849: Histoire naturelle des animaux articules. Deuxieme partie. Insectes. In: Exploration scientifique de l'Algerie pendant les annees 1840, 1841, 1842 publiee par ordre du Gouvernement et avec le concours d'une Commission Academique. Sciences physiques. Zoologie II. A. Bertrand, Paris, 590 pp." type="book" year="1846">Lucas, 1846</bibRefCitation>
</taxonomicName>
; E –
<taxonomicName id="4C204D35FFDC4B3FB2AFA7DD16CFF874" authority="(Gravenhorst, 1802)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[697,981,2008,2027]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="263" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B954EAA4FFDC4B3FB2AFA7DD1605F874" box="[697,799,2008,2027]" italics="true" pageId="38" pageNumber="263">Q. laticollis</emphasis>
(
<bibRefCitation id="EFB14B47FFDC4B3FB33CA7DD16D5F874" author="GRAVENHORST J. L. C." box="[810,975,2008,2027]" pageId="38" pageNumber="263" refId="ref83803" refString="GRAVENHORST J. L. C. 1802: Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera, familiae et species distribuit. C. Reichard, Brunsuigae, 206 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
)
</taxonomicName>
; F –
<taxonomicName id="4C204D35FFDC4B3FB414A7DD11CCF874" authority="Casey, 1915" authorityName="Casey" authorityYear="1915" box="[1026,1238,2008,2027]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="263" phylum="Arthropoda" rank="species" species="strenuus">
<emphasis id="B954EAA4FFDC4B3FB414A7DD117FF874" box="[1026,1125,2008,2027]" italics="true" pageId="38" pageNumber="263">Q. strenuus</emphasis>
<bibRefCitation id="EFB14B47FFDC4B3FB47DA7DD11CCF874" author="CASEY T. L." box="[1131,1238,2008,2027]" pageId="38" pageNumber="263" pagination="395 - 450" refId="ref81365" refString="CASEY T. L. 1915: Studies in some staphylinid genera of North America. Memoirs on the Coleoptera 6: 395 - 450." type="journal article" year="1915">Casey, 1915</bibRefCitation>
</taxonomicName>
. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.
</paragraph>
</caption>
<subSubSection id="C33A653DFFDD4B3EB081A22C114EFD2A" pageId="39" pageNumber="264" type="description">
<paragraph id="8B9F36B6FFDD4B3EB081A22C15FAFC5F" blockId="39.[151,760,548,1408]" pageId="39" pageNumber="264">
<emphasis id="B954EAA4FFDD4B3EB081A22C145AFDDF" bold="true" box="[151,320,553,576]" pageId="39" pageNumber="264">Redescription.</emphasis>
Measurements JJ (n = 5): HW = 1.40– 1.69 (1.53); HL = 1.22–1.40 (1.29); HL/HW 0.79–0.89 (0.85); PW = 1.82–2.22 (2.00); PL = 1.71–2.07 (1.81); PL/PW 0.80–0.96 (0.91); EW = 1.80–2.09 (1.88); EL = 1.44–1.71 (1.60); EL/EW 0.80–0.89 (0.85); EL/PL 0.83–0.95 (0.89); PW/HW 1.46–1.67 (1.55); forebody length 4.38–5.18 (4.70).
<specimenCount id="9D26FD3FFFDD4B3EB1BBA2E714C9FC9E" box="[429,467,738,769]" pageId="39" pageNumber="264" type="female">♀♀</specimenCount>
(n = 5): HW = 1.58–1.73 (1.67); HL = 1.33–1.42 (1.38); HL/HW 0.78–0.87 (0.83); PW = 1.93–2.24 (2.04); PL = 1.80–2.11 (1.88); PL/PW 0.90–0.94 (0.92); EW = 1.89–2.09 (1.96); EL = 1.69–1.78 (1.72); EL/EW 0.83–0.91 (0.88); EL/PL 0.82–0.96 (0.92); PW/HW 1.39–1.58 (1.48); forebody length 4.82–5.27 (4.99).
</paragraph>
<paragraph id="8B9F36B6FFDD4B3EB0A3A3CC17E2FC7F" blockId="39.[151,760,548,1408]" box="[181,760,969,992]" pageId="39" pageNumber="264">
Small sized species; body dark brown to black (
<figureCitation id="131B2A33FFDD4B3EB28BA3CC17F6FC7F" box="[669,748,969,992]" captionStart="Fig" captionStartId="18.[151,182,2027,2046]" captionTargetBox="[189,1393,195,2007]" captionTargetId="figure-13@18.[174,1413,185,2007]" captionTargetPageId="18" captionText="Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003." figureDoi="http://doi.org/10.5281/zenodo.7399722" httpUri="https://zenodo.org/record/7399722/files/figure.png" pageId="39" pageNumber="264">Fig. 8F</figureCitation>
).
</paragraph>
<paragraph id="8B9F36B6FFDD4B3EB0A3A3EC141AFB5F" blockId="39.[151,760,548,1408]" pageId="39" pageNumber="264">
Head black, distinctly transverse, with eyes medium sized (EyL/TL = 1.73–2.00 (1.83)); microsculpture of fine transverse waves; no additional punctures between anterior frontal punctures (cf.
<figureCitation id="131B2A33FFDD4B3EB1CFA44C1730FBFF" box="[473,554,1097,1120]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="39" pageNumber="264">Fig. 6F</figureCitation>
); antennae dark internally becoming continuously lighter, all antennomeres slightly elongate; palpi pale with apical palpomere slightly darkened.
</paragraph>
<paragraph id="8B9F36B6FFDD4B3EB0A3A4CC1140FE8A" blockId="39.[151,760,548,1408]" lastBlockId="39.[828,1437,190,1397]" pageId="39" pageNumber="264">Thorax: pronotum black, wider than long, wider than head, with microsculpture of transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond level of first puncture of dorsal row; scutellum sparsely punctured and pubescent; elytra most often fully darkened but occasional dark reddish brown, short uniformly pubescent, clearly shortened wider than long, clearly shorter than pronotum; legs dark with tarsi lighter.</paragraph>
<paragraph id="8B9F36B6FFDD4B3EB34CA11B16A1FEEA" blockId="39.[828,1437,190,1397]" pageId="39" pageNumber="264">Abdomen black, tergites sparsely and uniformly punctured, without palisade fringe of tergite 7, without clear iridescence.</paragraph>
<paragraph id="8B9F36B6FFDD4B3EB34CA178114EFD2A" blockId="39.[828,1437,190,1397]" pageId="39" pageNumber="264">
<emphasis id="B954EAA4FFDD4B3EB34CA1781683FE0A" bold="true" box="[858,921,381,405]" italics="true" pageId="39" pageNumber="264">Male.</emphasis>
Aedeagus (
<figureCitation id="131B2A33FFDD4B3EB40EA17B1161FE0A" box="[1048,1147,382,405]" captionStart="Fig" captionStartId="31.[151,182,2004,2023]" captionTargetBox="[207,1368,203,1953]" captionTargetId="figure-12@31.[191,1396,189,1962]" captionTargetPageId="31" captionText="Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view." figureDoi="http://doi.org/10.5281/zenodo.7399732" httpUri="https://zenodo.org/record/7399732/files/figure.png" pageId="39" pageNumber="264">Fig. 13B</figureCitation>
): paramere lanceolate with slight medial attenuation and extending into a slight expansion broadest below apex, apex slightly asymmetric, reaching just beyond apex of median lobe, with sensory peg setae forming two long rows fusing together towards apex (
<figureCitation id="131B2A33FFDD4B3EB36DA21B16D2FDAA" box="[891,968,542,565]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="39" pageNumber="264">Fig. 15</figureCitation>
); median lobe broad with gentle constriction to a point at apex, on parameral side with two small teeth pointing slightly basad, positioned at level near basal level of peg setae band of paramere; internal sac without a continuation of C-sclerite.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFDD4B3EB32AA2BB1087FC4A" pageId="39" pageNumber="264" type="diagnosis">
<paragraph id="8B9F36B6FFDD4B3EB32AA2BB1087FC4A" blockId="39.[828,1437,190,1397]" pageId="39" pageNumber="264">
<emphasis id="B954EAA4FFDD4B3EB32AA2BB112BFD4A" bold="true" box="[828,1073,702,725]" pageId="39" pageNumber="264">Differential diagnosis.</emphasis>
<taxonomicName id="4C204D35FFDD4B3EB423A2BB11E5FD4A" authorityName="Smetana" authorityYear="2003" box="[1077,1279,702,725]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="264" phylum="Arthropoda" rank="species" species="sundukovi">
<emphasis id="B954EAA4FFDD4B3EB423A2BB11E5FD4A" box="[1077,1279,702,725]" italics="true" pageId="39" pageNumber="264">Quedius sundukovi</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C204D35FFDD4B3EB341A2DB16CBFD6A" authorityName="Kiesenwetter" authorityYear="1847" box="[855,977,734,757]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="264" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B954EAA4FFDD4B3EB341A2DB16CBFD6A" box="[855,977,734,757]" italics="true" pageId="39" pageNumber="264">Q. unicolor</emphasis>
</taxonomicName>
,
<taxonomicName id="4C204D35FFDD4B3EB3C9A2DB1164FD6A" authorityName="Korge" authorityYear="1961" box="[991,1150,734,757]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="264" phylum="Arthropoda" rank="species" species="subunicolor">
<emphasis id="B954EAA4FFDD4B3EB3C9A2DB1164FD6A" box="[991,1150,734,757]" italics="true" pageId="39" pageNumber="264">Q. subunicolor</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C204D35FFDD4B3EB4AEA2DB104EFD6A" box="[1208,1364,734,757]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="264" phylum="Arthropoda" rank="species" species="molochinus">
<emphasis id="B954EAA4FFDD4B3EB4AEA2DB104EFD6A" box="[1208,1364,734,757]" italics="true" pageId="39" pageNumber="264">Q. molochinus</emphasis>
</taxonomicName>
. It can easily be distinguished from all these species by its smaller size and brachypterous habitus with shortened elytra and absent palisade fringe on tergite VII. If in doubt, the aedeagus can be checked for characters listed in the diagnosis. It is also similar to Siberian species
<taxonomicName id="4C204D35FFDD4B3EB48DA37B16E6FC2A" authority="Sahlberg, 1880" authorityName="Sahlberg" authorityYear="1880" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="264" phylum="Arthropoda" rank="species" species="jenisseensis" subGenus="Raphirus">
<emphasis id="B954EAA4FFDD4B3EB48DA37B11AFFC0A" box="[1179,1205,894,917]" italics="true" pageId="39" pageNumber="264">Q.</emphasis>
(
<emphasis id="B954EAA4FFDD4B3EB4D4A37B103EFC0A" box="[1218,1316,894,917]" italics="true" pageId="39" pageNumber="264">Raphirus</emphasis>
)
<emphasis id="B954EAA4FFDD4B3EB524A37B1642FC2A" italics="true" pageId="39" pageNumber="264">jenisseensis</emphasis>
Sahlberg, 1880
</taxonomicName>
, from which it is easily distinguished by the entire labrum (medially incisioned in
<taxonomicName id="4C204D35FFDD4B3EB4F9A3BB1097FC4A" authorityName="Sahlberg" authorityYear="1880" box="[1263,1421,958,981]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="264" phylum="Arthropoda" rank="species" species="jenisseensis">
<emphasis id="B954EAA4FFDD4B3EB4F9A3BB1097FC4A" box="[1263,1421,958,981]" italics="true" pageId="39" pageNumber="264">Q. jenisseensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFDD4B31B32AA3DB17EEFC8A" lastPageId="40" lastPageNumber="265" pageId="39" pageNumber="264" type="discussion">
<paragraph id="8B9F36B6FFDD4B31B32AA3DB17EEFC8A" blockId="39.[828,1437,190,1397]" lastBlockId="40.[151,761,190,1301]" lastPageId="40" lastPageNumber="265" pageId="39" pageNumber="264">
<emphasis id="B954EAA4FFDD4B3EB32AA3DB16A4FC6A" bold="true" box="[828,958,990,1013]" pageId="39" pageNumber="264">Comments.</emphasis>
<taxonomicName id="4C204D35FFDD4B3EB3D5A3DB1194FC6A" authorityName="Smetana" authorityYear="2003" box="[963,1166,990,1013]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="264" phylum="Arthropoda" rank="species" species="sundukovi">
<emphasis id="B954EAA4FFDD4B3EB3D5A3DB1194FC6A" box="[963,1166,990,1013]" italics="true" pageId="39" pageNumber="264">Quedius sundukovi</emphasis>
</taxonomicName>
was described by Sආൾඍൺ-ඇൺ (2003) based on
<specimenCount id="9D26FD3FFFDD4B3EB40CA3FB11B6FB8A" box="[1050,1196,1022,1045]" pageId="39" pageNumber="264" type="generic">13 specimens</specimenCount>
from the Badshalskyi Khrebet mountains in
<collectingRegion id="49E4F854FFDD4B3EB43FA41B11C5FBAA" box="[1065,1247,1054,1077]" country="Russia" name="Khabarovsk" pageId="39" pageNumber="264">Khabarovsk Krai</collectingRegion>
in the Far East of
<collectingCountry id="F3377626FFDD4B3EB32AA43B169DFBCA" box="[828,903,1086,1109]" name="Russia" pageId="39" pageNumber="264">Russia</collectingCountry>
. Sආൾඍൺඇൺ &amp; SHൺඏඋංඇ (2018) reported additional specimens from Vitimskiy Nature Reserve and the Udokan Plateau extending the species range significantly westwards in
<collectingCountry id="F3377626FFDD4B3EB34EA49B1685FB2A" box="[856,927,1182,1205]" name="Russia" pageId="39" pageNumber="264">Russia</collectingCountry>
to Transbaikalia. Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2018a) reported a number of records of this species from the
<collectingRegion id="49E4F854FFDD4B3EB372A4DB1683FB6A" box="[868,921,1246,1269]" country="Russia" name="Altay" pageId="39" pageNumber="264">Altai</collectingRegion>
Mountains in
<collectingCountry id="F3377626FFDD4B3EB42CA4DB11A0FB6A" box="[1082,1210,1246,1269]" name="Kazakhstan" pageId="39" pageNumber="264">Kazakhstan</collectingCountry>
. Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈ-ൽඈඏඇංĸඈඏ (2019) reported the occurence of this species in Buryat Republic in Transbaikalia and in the Russian Far East regions of Amur Oblast and Sakhalin Island (
<figureCitation id="131B2A33FFDD4B3EB567A53B1640FAEA" captionStart="Fig" captionStartId="59.[151,182,2028,2047]" captionTargetBox="[190,1392,189,2003]" captionTargetId="figure-13@59.[190,1393,189,2003]" captionTargetPageId="59" captionText="Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399748" httpUri="https://zenodo.org/record/7399748/files/figure.png" pageId="39" pageNumber="264">Fig. 20</figureCitation>
). Here we add detailed information on many of these records along with COI barcodes from specimens across the range (
<figureCitation id="131B2A33FFD24B31B111A0DB1452FF6A" box="[263,328,222,245]" captionStart="Fig" captionStartId="12.[151,182,2008,2027]" captionTargetBox="[182,1406,199,1981]" captionTargetId="figure-13@12.[182,1407,189,1989]" captionTargetPageId="12" captionText="Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP &lt;0.90, UFB &lt;95 and SH-aLRT &lt;80." figureDoi="http://doi.org/10.5281/zenodo.7399710" httpUri="https://zenodo.org/record/7399710/files/figure.png" pageId="40" pageNumber="265">Fig. 4</figureCitation>
). The barcodes showed high variation as we recovered five OTUs (four BINs) with as high as 7.7% divergence between the OTUs (
<tableCitation id="C6A2030DFFD24B31B1F9A11B175EFEAA" box="[495,580,286,309]" captionStart="Table 2" captionStartId="13.[151,201,190,209]" captionTargetPageId="13" captionText="Table 2. Genetic distance between clusters of CO1 barcodes.n = number of specimens barcoded. Number of OTUs found by species delimiter plugin in Geneious. Intraspecific distance within cluster. Interspecific distance is distance to nearest other cluster." httpUri="http://table.plazi.org/id/DF5F663EFFF74B14B081A0BB1713FEBE" pageId="40" pageNumber="265" tableUuid="DF5F663EFFF74B14B081A0BB1713FEBE">Table 2</tableCitation>
). Morphological studies of the specimens also revealed some variation in size – but not in the shape – of the paramere (
<figureCitation id="131B2A33FFD24B31B262A15B17D8FEEA" box="[628,706,350,373]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="40" pageNumber="265">Fig. 15</figureCitation>
). We found no differences to correspond to any of the molecular clusters, which also did not cluster geographically. Lack of a clear congruence between the molecular and morphological variation, as well as no hiatus between different morphological variants, suggests that they all represent a single species. The species is known across a large area with complex montane orography and it is flightless, with the palisade fringe of abdominal segment VII absent and clearly shortened elytra.Apparently, the high degree of the intraspecific variation can be related to the flightlessness of the species leading to low dispersal abilities of members of different populations and accelerated rates of divergence in the barcode region (Mංඍඍൾඋൻඈൾർĸ &amp; Aൽൺආඈඐංർඓ 2013).
</paragraph>
</subSubSection>
<caption id="DF5F663EFFDD4B3EB08EA7F415EDF780" ID-DOI="http://doi.org/10.5281/zenodo.7399736" ID-Zenodo-Dep="7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="39" pageNumber="264" startId="39.[152,183,2033,2052]" targetBox="[284,1300,1425,2027]" targetPageId="39">
<paragraph id="8B9F36B6FFDD4B3EB08EA7F415EDF780" blockId="39.[151,1436,2033,2079]" pageId="39" pageNumber="264">
Fig. 15. Variation of the shape of the paramere of
<taxonomicName id="4C204D35FFDD4B3EB253A7F4166AF79B" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[581,880,2033,2052]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="264" phylum="Arthropoda" rank="species" species="sundukovi">
<emphasis id="B954EAA4FFDD4B3EB253A7F417F3F79B" box="[581,745,2033,2052]" italics="true" pageId="39" pageNumber="264">Quedius sundukovi</emphasis>
<bibRefCitation id="EFB14B47FFDD4B3EB2F8A7F4166AF79B" author="SMETANA A." box="[750,880,2033,2052]" pageId="39" pageNumber="264" pagination="189 - 193" refId="ref92269" refString="SMETANA A. 2003: Quedius (Quedius) sundukovi (Coleoptera, Staphylinidae, Staphylinini, Quediina), an interesting new species from the Russian Far East. Elytra 31: 189 - 193." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
</taxonomicName>
. Locality and BOLD BIN number are indicated under respective specimens.
</paragraph>
</caption>
<subSubSection id="C33A653DFFD24B31B081A31B17E3FC6A" pageId="40" pageNumber="265" type="description">
<paragraph id="8B9F36B6FFD24B31B081A31B17E3FC6A" blockId="40.[151,761,190,1301]" pageId="40" pageNumber="265">
<emphasis id="B954EAA4FFD24B31B081A31B1402FCAA" bold="true" box="[151,280,798,821]" pageId="40" pageNumber="265">Bionomics.</emphasis>
Little is known about the specific habitat requirement of
<taxonomicName id="4C204D35FFD24B31B12CA33B14DCFCCA" authorityName="Smetana" authorityYear="2003" box="[314,454,830,853]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="265" phylum="Arthropoda" rank="species" species="sundukovi">
<emphasis id="B954EAA4FFD24B31B12CA33B14DCFCCA" box="[314,454,830,853]" italics="true" pageId="40" pageNumber="265">Q. sundukovi</emphasis>
</taxonomicName>
. Based on the examined material, the species appear to occur in various leaf litter, both coniferous and deciduous, and is commonly associated with talus debris and moss, often near streams or rivers. Specimens are found within a wide range of elevations from
<quantity id="4CD89B53FFD24B31B0C6A3DB1473FC6A" box="[208,361,990,1013]" metricMagnitude="3" metricUnit="m" metricValue="1.3074999999999999" metricValueMax="2.3" metricValueMin="0.315" pageId="40" pageNumber="265" unit="m" value="1307.5" valueMax="2300.0" valueMin="315.0">315 to 2300 m</quantity>
, at higher elevation at lower latitudes.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFD24B31B081A3FB17D6FA8A" pageId="40" pageNumber="265" type="distribution">
<paragraph id="8B9F36B6FFD24B31B081A3FB17D6FA8A" blockId="40.[151,761,190,1301]" pageId="40" pageNumber="265">
<emphasis id="B954EAA4FFD24B31B081A3FB1433FB8A" bold="true" box="[151,297,1022,1045]" pageId="40" pageNumber="265">Distribution.</emphasis>
<taxonomicName id="4C204D35FFD24B31B13BA3FB14EDFB8A" authorityName="Smetana" authorityYear="2003" box="[301,503,1022,1045]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="265" phylum="Arthropoda" rank="species" species="sundukovi">
<emphasis id="B954EAA4FFD24B31B13BA3FB14EDFB8A" box="[301,503,1022,1045]" italics="true" pageId="40" pageNumber="265">Quedius sundukovi</emphasis>
</taxonomicName>
is currently known from relatively few specimens scattered across a large range comprising the Altai Mountains, Transbaikalia, Bureya and the Suntar-Khayata Range (
<figureCitation id="131B2A33FFD24B31B1F4A45B1735FBEA" box="[482,559,1118,1141]" captionStart="Fig" captionStartId="59.[151,182,2028,2047]" captionTargetBox="[190,1392,189,2003]" captionTargetId="figure-13@59.[190,1393,189,2003]" captionTargetPageId="59" captionText="Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399748" httpUri="https://zenodo.org/record/7399748/files/figure.png" pageId="40" pageNumber="265">Fig. 20</figureCitation>
). It is also found on Sakhalin Island.Although currently known from only a few localities, the species distribution is most likely continuous. The range maybe poorly known because it spans through an area with very little recent and historical entomological activity, especially for leaf-litter dwelling organisms.
</paragraph>
</subSubSection>
</treatment>
</document>
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