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<document id="9250B080607C7E22DF3872BCFFE4EFB0" ID-DOI="10.11646/zootaxa.3670.2.11" ID-GBIF-Dataset="bc516484-c1c5-4ee3-8912-7ff89f906a8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="219876" ID-ZooBank="67402B31-9D14-4116-8752-2927A1084240" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460633918393" checkinUser="plazi" docAuthor="Reboleira, Ana Sofia P. S., Zaragoza, Juan A., Gonçalves, Fernando &amp; Oromí, Pedro" docDate="2013" docId="015C614EF403FF905C94F88C036CFE7E" docLanguage="en" docName="zt03670p299.pdf" docOrigin="Zootaxa 3670 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Roncocreagris gepesi Zaragoza &amp; Reboleira, sp. nov." docType="treatment" docUuid="1F3B1AE9-F53A-4159-9ED3-42751F47BF29" docUuidSource="ZooBank" docVersion="9" lastPageNumber="294" masterDocId="FD651936F40BFF9B5C03FFB30766FFAE" masterDocTitle="On hypogean Roncocreagris (Arachnida: Pseudoscorpiones: Neobisiidae) from Portugal, with descriptions of three new species" masterLastPageNumber="299" masterPageNumber="283" pageNumber="291" updateTime="1698334986200" updateUser="plazi">
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<mods:title id="6300E5B485A15E0548767099F0FAAE95">On hypogean Roncocreagris (Arachnida: Pseudoscorpiones: Neobisiidae) from Portugal, with descriptions of three new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="BDF8F4BC4770CE6B01601032F8485010">Reboleira, Ana Sofia P. S.</mods:namePart>
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<mods:namePart id="7B24B04B3743AC304B616F09226ED1E9">Zaragoza, Juan A.</mods:namePart>
</mods:name>
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<mods:namePart id="B90FCECCB341054BE42A79BCF55C34F6">Gonçalves, Fernando</mods:namePart>
</mods:name>
<mods:name id="D9091DF2616D0DED1B4F4B4D7E8C4A7D" type="personal">
<mods:role id="56122061681C02AFB79E92F129F70DEB">
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<mods:namePart id="BB449663A82399F358A8303ABD36EE97">Oromí, Pedro</mods:namePart>
</mods:name>
<mods:typeOfResource id="BD5B9FFFCC4031B23C1985657A146A7E">text</mods:typeOfResource>
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<mods:title id="B2636C01A7E33F01F4BBD3EF239ECF3E">Zootaxa</mods:title>
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<mods:part id="DF790BA28437C1A8317708F9E6452AAB">
<mods:date id="987CC30DDC5283F07601EE61371C1957">2013</mods:date>
<mods:detail id="2D5713F18354C1023B51592D483FDB54" type="volume">
<mods:number id="993F53749D64195017F216FA33FD51ED">3670</mods:number>
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<mods:identifier id="3D20635BDCEAF213D9191839A8157976" type="DOI">10.11646/zootaxa.3670.2.11</mods:identifier>
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<mods:identifier id="5AA283447377BBE2EE0AA142D9EAD2A9" type="ISSN">1175-5326</mods:identifier>
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<treatment id="015C614EF403FF905C94F88C036CFE7E" ID-DOI="http://doi.org/10.5281/zenodo.6146634" ID-GBIF-Taxon="119570119" ID-Zenodo-Dep="6146634" LSID="urn:lsid:zoobank.org:act:1F3B1AE9-F53A-4159-9ED3-42751F47BF29" httpUri="http://treatment.plazi.org/id/015C614EF403FF905C94F88C036CFE7E" lastPageId="11" lastPageNumber="294" pageId="8" pageNumber="291">
<subSubSection id="C1EF83D3F403FF935C94F88C0641F8D7" pageId="8" pageNumber="291" type="nomenclature">
<paragraph id="894AD058F403FF935C94F88C047BF8F7" blockId="8.[151,797,1855,1913]" box="[151,797,1855,1881]" pageId="8" pageNumber="291">
<heading id="D2026734F403FF935C94F88C047BF8F7" bold="true" box="[151,797,1855,1881]" fontSize="11" level="1" pageId="8" pageNumber="291" reason="1">
<emphasis id="BB810C4AF403FF935C94F88C047BF8F7" bold="true" box="[151,797,1855,1881]" pageId="8" pageNumber="291">
<taxonomicName id="4EF5ABDBF403FF935C94F88C05DEF8F7" ID-CoL="4TB4B" authority="Zaragoza &amp; Reboleira" authorityName="Zaragoza &amp; Reboleira" box="[151,696,1855,1881]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="291" phylum="Arthropoda" rank="species" species="gepesi" status="sp. nov.">
<emphasis id="BB810C4AF403FF935C94F88C06FFF8F7" bold="true" box="[151,409,1855,1881]" italics="true" pageId="8" pageNumber="291">Roncocreagris gepesi</emphasis>
Zaragoza &amp; Reboleira
</taxonomicName>
<taxonomicNameLabel id="A0B2B131F403FF935EBCF88C047BF8F7" box="[703,797,1855,1881]" pageId="8" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="894AD058F403FF935C94F8D20641F8D7" blockId="8.[151,797,1855,1913]" box="[151,295,1889,1913]" pageId="8" pageNumber="291">
(
<figureCitation id="11CECCDDF403FF935C9CF8D20679F8D7" box="[159,287,1889,1913]" captionStart="FIGURES 23 – 32" captionStartId="9.[151,269,1878,1902]" captionTargetBox="[170,1415,213,1845]" captionTargetId="figure@9.[151,1436,194,1856]" captionTargetPageId="9" captionText="FIGURES 23 – 32. Roncocreagris gepesi sp. nov., female holotype. 23. carapace; 24. anterior margin of carapace; 25. anterior and medial processes of left coxa I; 26. left chelicera; 27. partial view of fingers of left chelicera, showing galea; 28. right pedipalp minus chela, dorsal view; 29. right chela, dorsal view; 30. right chela, lateral view; 31. claw of left leg IV; 32. subterminal seta of left leg IV. Scale bars (in mm): 0.05 (Figs 25, 31, 32), 0.10 (Figs 24, 26, 27), 0.30 (Fig. 23), 0.50 (Figs 28 – 30)." httpUri="https://zenodo.org/record/219881/files/figure.png" pageId="8" pageNumber="291">Figs 23–32</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F403FF935C94F81903ACF848" pageId="8" pageNumber="291" type="materials_examined">
<paragraph id="894AD058F403FF935C94F81903ACF848" blockId="8.[151,1436,1961,2023]" pageId="8" pageNumber="291">
<emphasis id="BB810C4AF403FF935C94F8190620F86C" bold="true" box="[151,326,1961,1986]" pageId="8" pageNumber="291">
<typeStatus id="564E6EFAF403FF935C94F81907B4F86C" box="[151,210,1962,1986]" pageId="8" pageNumber="291">Type</typeStatus>
material.
</emphasis>
<typeStatus id="564E6EFAF403FF935D4DF81A06D0F86F" box="[334,438,1961,1985]" pageId="8" pageNumber="291" type="holotype">Holotype</typeStatus>
Ƥ,
<collectingCountry id="F1E290C8F403FF935DE1F81A0522F86F" box="[482,580,1961,1985]" name="Portugal" pageId="8" pageNumber="291">Portugal</collectingCountry>
, Sicó Massif, district of Coimbra, Penela, Gruta do Algarinho/Dueça Cave system (39°59ʹ4.89&quot;N, 8°23ʹ5.86&quot;W;
<specimenCount id="9FF31BD1F403FF935E3EF87D05E9F848" box="[573,655,1998,2022]" pageId="8" pageNumber="291" type="generic">
<quantity id="4E0D7DBDF403FF935E3EF87D05EDF848" box="[573,651,1998,2022]" metricMagnitude="2" metricUnit="m" metricValue="2.08" pageId="8" pageNumber="291" unit="m" value="208.0">208 m</quantity>
.
</specimenCount>
a.s.l.),
<date id="FD4BF698F403FF935ED5F87D0432F848" box="[726,852,1998,2022]" pageId="8" pageNumber="291" value="2009-04-25">25.IV.2009</date>
, lgt. A.S.P.S. Reboleira (
<collectionCode id="EFE4489DF403FF935869F87D03D9F848" box="[1130,1215,1998,2022]" pageId="8" pageNumber="291">DEUA</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DD8A80D0F402FF925C94F8E5063BF843" httpUri="https://zenodo.org/record/219881/files/figure.png" pageId="9" pageNumber="292" targetBox="[170,1415,213,1845]" targetPageId="9">
<paragraph id="894AD058F402FF925C94F8E5063BF843" blockId="9.[151,1436,1878,2029]" pageId="9" pageNumber="292">
<emphasis id="BB810C4AF402FF925C94F8E50606F8C0" bold="true" box="[151,352,1878,1902]" pageId="9" pageNumber="292">FIGURES 23–32.</emphasis>
<taxonomicName id="4EF5ABDBF402FF925D6BF8E50536F8C3" box="[360,592,1878,1901]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="292" phylum="Arthropoda" rank="species" species="gepesi" status="sp. nov.">
<emphasis id="BB810C4AF402FF925D6BF8E50536F8C3" box="[360,592,1878,1901]" italics="true" pageId="9" pageNumber="292">Roncocreagris gepesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0B2B131F402FF925E5AF8E505D7F8C0" box="[601,689,1878,1902]" pageId="9" pageNumber="292" rank="species">
<emphasis id="BB810C4AF402FF925E5AF8E505D7F8C0" bold="true" box="[601,689,1878,1902]" pageId="9" pageNumber="292">sp. nov.</emphasis>
</taxonomicNameLabel>
, female holotype.
<emphasis id="BB810C4AF402FF925F7DF8E504F9F8C0" bold="true" box="[894,927,1878,1902]" pageId="9" pageNumber="292">23.</emphasis>
carapace;
<emphasis id="BB810C4AF402FF925816F8E50350F8C0" bold="true" box="[1045,1078,1878,1902]" pageId="9" pageNumber="292">24.</emphasis>
anterior margin of carapace;
<emphasis id="BB810C4AF402FF925978F8E502FAF8C0" bold="true" box="[1403,1436,1878,1902]" pageId="9" pageNumber="292">25.</emphasis>
anterior and medial processes of left coxa I;
<emphasis id="BB810C4AF402FF925E6DF8C505F6F820" bold="true" box="[622,656,1910,1934]" pageId="9" pageNumber="292">26.</emphasis>
left chelicera;
<emphasis id="BB810C4AF402FF925F2DF8C50429F820" bold="true" box="[814,847,1910,1934]" pageId="9" pageNumber="292">27.</emphasis>
partial view of fingers of left chelicera, showing galea;
<emphasis id="BB810C4AF402FF925C94F82507DEF800" bold="true" box="[151,184,1942,1966]" pageId="9" pageNumber="292">28.</emphasis>
right pedipalp minus chela, dorsal view;
<emphasis id="BB810C4AF402FF925E72F82505F5F800" bold="true" box="[625,659,1942,1966]" pageId="9" pageNumber="292">29.</emphasis>
right chela, dorsal view;
<emphasis id="BB810C4AF402FF925FA1F82504A5F800" bold="true" box="[930,963,1942,1966]" pageId="9" pageNumber="292">30.</emphasis>
right chela, lateral view;
<emphasis id="BB810C4AF402FF9258D0F8250392F800" bold="true" box="[1235,1268,1942,1966]" pageId="9" pageNumber="292">31.</emphasis>
claw of left leg IV;
<emphasis id="BB810C4AF402FF925CC3F8050787F860" bold="true" box="[192,225,1974,1998]" pageId="9" pageNumber="292">32.</emphasis>
subterminal seta of left leg IV. Scale bars (in mm): 0.05 (Figs 25, 31, 32), 0.10 (Figs 24, 26, 27), 0.30 (Fig. 23), 0.50 (Figs 28–30).
</paragraph>
</caption>
<subSubSection id="C1EF83D3F401FF915CC4FF2B05A6FF59" pageId="10" pageNumber="293" type="etymology">
<paragraph id="894AD058F401FF915CC4FF2B05A6FF59" blockId="10.[151,1437,151,2029]" pageId="10" pageNumber="293">
<emphasis id="BB810C4AF401FF915CC4FF2B0628FF1E" bold="true" box="[199,334,152,176]" pageId="10" pageNumber="293">Etymology.</emphasis>
This species is dedicated to the speleological association Grupo Protecção Sicó (GPS), for its contribution to the knowledge and protection of caves and karst of Sicó region, and as recognition to its members for their effort and support during our field work.
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F401FF915CC4FEB003C7FE24" pageId="10" pageNumber="293" type="diagnosis">
<paragraph id="894AD058F401FF915CC4FEB003C7FE24" blockId="10.[151,1437,151,2029]" pageId="10" pageNumber="293">
<emphasis id="BB810C4AF401FF915CC4FEB00627FEB2" bold="true" box="[199,321,259,284]" pageId="10" pageNumber="293">Diagnosis.</emphasis>
No eyes or eye-spots. Strong troglomorphic adaptations. Carapace with 21 setae, 7 on the posterior margin. Tergite
<date id="FD4BF698F401FF915D4BFE9B0600FE91" box="[328,358,295,319]" pageId="10" pageNumber="293">I 6</date>
setae. Female cheliceral galea long, with five short, apical rami. Pedipalp: femur ratio about 6.6, femur longer than movable finger; chela+ ratio about 5.9; chelal hand widest at middle, ratio movable finger/hand+ 1.4; trichobothium
<emphasis id="BB810C4AF401FF915D6DFEC706EFFE25" box="[366,393,372,395]" italics="true" pageId="10" pageNumber="293">ist</emphasis>
close to middle of finger, ratio
<emphasis id="BB810C4AF401FF915EE9FEC0044FFE25" box="[746,809,371,395]" italics="true" pageId="10" pageNumber="293">ib–ist</emphasis>
/
<emphasis id="BB810C4AF401FF915F33FEC7040EFE25" box="[816,872,371,395]" italics="true" pageId="10" pageNumber="293">ist–it</emphasis>
1.3, ratio
<emphasis id="BB810C4AF401FF915FD8FEC00345FE25" box="[987,1059,371,395]" italics="true" pageId="10" pageNumber="293">isb–ist</emphasis>
/
<emphasis id="BB810C4AF401FF915828FEC00316FE25" box="[1067,1136,371,395]" italics="true" pageId="10" pageNumber="293">ib–isb</emphasis>
1.8.
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F401FF915CC4FE250592F8F4" pageId="10" pageNumber="293" type="description">
<paragraph id="894AD058F401FF915CC4FE25032FFE7C" blockId="10.[151,1437,151,2029]" pageId="10" pageNumber="293">
<emphasis id="BB810C4AF401FF915CC4FE25063FFE01" bold="true" box="[199,345,406,431]" pageId="10" pageNumber="293">Description.</emphasis>
Large pseudoscorpion. Opisthosoma elongate, advanced troglomorphic adaptations. Opisthosoma and legs yellowish. Carapace, chelicerae and pedipalps reddish brown.
</paragraph>
<paragraph id="894AD058F401FF915CC4FE6C04BEFD91" blockId="10.[151,1437,151,2029]" pageId="10" pageNumber="293">
Carapace distinctly longer than broad (
<figureCitation id="11CECCDDF401FF915E9AFE6C0595FE59" box="[665,755,479,503]" captionStart="FIGURES 23 – 32" captionStartId="9.[151,269,1878,1902]" captionTargetBox="[170,1415,213,1845]" captionTargetId="figure@9.[151,1436,194,1856]" captionTargetPageId="9" captionText="FIGURES 23 – 32. Roncocreagris gepesi sp. nov., female holotype. 23. carapace; 24. anterior margin of carapace; 25. anterior and medial processes of left coxa I; 26. left chelicera; 27. partial view of fingers of left chelicera, showing galea; 28. right pedipalp minus chela, dorsal view; 29. right chela, dorsal view; 30. right chela, lateral view; 31. claw of left leg IV; 32. subterminal seta of left leg IV. Scale bars (in mm): 0.05 (Figs 25, 31, 32), 0.10 (Figs 24, 26, 27), 0.30 (Fig. 23), 0.50 (Figs 28 – 30)." httpUri="https://zenodo.org/record/219881/files/figure.png" pageId="10" pageNumber="293">Fig. 23</figureCitation>
). Without eyes or eye-spots. Anterior margin moderately prominent medially, epistome blunt and short (
<figureCitation id="11CECCDDF401FF915EF3FDB10434FDB4" box="[752,850,514,538]" captionStart="FIGURES 23 – 32" captionStartId="9.[151,269,1878,1902]" captionTargetBox="[170,1415,213,1845]" captionTargetId="figure@9.[151,1436,194,1856]" captionTargetPageId="9" captionText="FIGURES 23 – 32. Roncocreagris gepesi sp. nov., female holotype. 23. carapace; 24. anterior margin of carapace; 25. anterior and medial processes of left coxa I; 26. left chelicera; 27. partial view of fingers of left chelicera, showing galea; 28. right pedipalp minus chela, dorsal view; 29. right chela, dorsal view; 30. right chela, lateral view; 31. claw of left leg IV; 32. subterminal seta of left leg IV. Scale bars (in mm): 0.05 (Figs 25, 31, 32), 0.10 (Figs 24, 26, 27), 0.30 (Fig. 23), 0.50 (Figs 28 – 30)." httpUri="https://zenodo.org/record/219881/files/figure.png" pageId="10" pageNumber="293">Fig. 24</figureCitation>
). Chaetotaxy: 21 setae, formula 4:4:6:7. Four microlyrifissures in ocular zone, two between median and posterior zones.
</paragraph>
<paragraph id="894AD058F401FF915CC4FDF90570FD04" blockId="10.[151,1437,151,2029]" pageId="10" pageNumber="293">
Coxal area. Manducatory process with 3 setae. Anterior process of coxa I with simple tooth shape, long and apically pointed; medial process straight with some small tubercles (
<figureCitation id="11CECCDDF401FF915FA2FDDC0491FD29" box="[929,1015,623,647]" captionStart="FIGURES 23 – 32" captionStartId="9.[151,269,1878,1902]" captionTargetBox="[170,1415,213,1845]" captionTargetId="figure@9.[151,1436,194,1856]" captionTargetPageId="9" captionText="FIGURES 23 – 32. Roncocreagris gepesi sp. nov., female holotype. 23. carapace; 24. anterior margin of carapace; 25. anterior and medial processes of left coxa I; 26. left chelicera; 27. partial view of fingers of left chelicera, showing galea; 28. right pedipalp minus chela, dorsal view; 29. right chela, dorsal view; 30. right chela, lateral view; 31. claw of left leg IV; 32. subterminal seta of left leg IV. Scale bars (in mm): 0.05 (Figs 25, 31, 32), 0.10 (Figs 24, 26, 27), 0.30 (Fig. 23), 0.50 (Figs 28 – 30)." httpUri="https://zenodo.org/record/219881/files/figure.png" pageId="10" pageNumber="293">Fig. 25</figureCitation>
). Pedipalpal coxa with 7 setae, pedal coxa I with 6, II: 5–6, III: 3, IV: 5.
</paragraph>
<paragraph id="894AD058F401FF915CC4FD040323FC94" blockId="10.[151,1437,151,2029]" pageId="10" pageNumber="293">
Tergal chaetotaxy I–X: 6:6:7:10:9:10:10:10:9:9, most hairs lacking. Genital area with 5 thin setae on sternite II. Sternal chaetotaxy III–X: 9:7:12:12:12:9:9:8; sternites VI and VII each with 2 discal setae (included in sternal formula); discal setae glandular, with visible duct. Segment
<date id="FD4BF698F401FF915F3EFCB30416FCB9" box="[829,880,767,791]" pageId="10" pageNumber="293">XI 9</date>
setae. Chaetotaxy of stigmata of sternites III and IV with 3 microsetae on each side. Anal cone with two dorsal and two ventral setae.
</paragraph>
<paragraph id="894AD058F401FF915CC4FCF40334FC41" blockId="10.[151,1437,151,2029]" pageId="10" pageNumber="293">
Chelicerae (
<figureCitation id="11CECCDDF401FF915D4DFCF406B5FCF1" box="[334,467,839,863]" captionStart="FIGURES 23 – 32" captionStartId="9.[151,269,1878,1902]" captionTargetBox="[170,1415,213,1845]" captionTargetId="figure@9.[151,1436,194,1856]" captionTargetPageId="9" captionText="FIGURES 23 – 32. Roncocreagris gepesi sp. nov., female holotype. 23. carapace; 24. anterior margin of carapace; 25. anterior and medial processes of left coxa I; 26. left chelicera; 27. partial view of fingers of left chelicera, showing galea; 28. right pedipalp minus chela, dorsal view; 29. right chela, dorsal view; 30. right chela, lateral view; 31. claw of left leg IV; 32. subterminal seta of left leg IV. Scale bars (in mm): 0.05 (Figs 25, 31, 32), 0.10 (Figs 24, 26, 27), 0.30 (Fig. 23), 0.50 (Figs 28 – 30)." httpUri="https://zenodo.org/record/219881/files/figure.png" pageId="10" pageNumber="293">Figs 26–27</figureCitation>
). Palm with 6 setae, subgaleal seta 0.68 from base of movable finger. Galea
<quantity id="4E0D7DBDF401FF915934FCF402FAFCF1" box="[1335,1436,839,863]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="293" unit="mm" value="0.04">0.04 mm</quantity>
long, with five short, apical rami. Fixed finger with 5 small, blunt distal denticles and 14 other small, medium teeth; movable finger with 13 teeth, one large and blunt subdistal tooth, the others medium or small, dental row ending just proximad of subgaleal seta. Rallum with 8 blades, all unilaterally pinnate on anterior face, basal blade about half length of others. Serrula exterior with 32 blades, serrula interior 27 blades.
</paragraph>
<paragraph id="894AD058F401FF915CC4FC490690FA47" blockId="10.[151,1437,151,2029]" pageId="10" pageNumber="293">
Pedipalps (
<figureCitation id="11CECCDDF401FF915D45FC4906ABFBBC" box="[326,461,1018,1042]" captionStart="FIGURES 23 – 32" captionStartId="9.[151,269,1878,1902]" captionTargetBox="[170,1415,213,1845]" captionTargetId="figure@9.[151,1436,194,1856]" captionTargetPageId="9" captionText="FIGURES 23 – 32. Roncocreagris gepesi sp. nov., female holotype. 23. carapace; 24. anterior margin of carapace; 25. anterior and medial processes of left coxa I; 26. left chelicera; 27. partial view of fingers of left chelicera, showing galea; 28. right pedipalp minus chela, dorsal view; 29. right chela, dorsal view; 30. right chela, lateral view; 31. claw of left leg IV; 32. subterminal seta of left leg IV. Scale bars (in mm): 0.05 (Figs 25, 31, 32), 0.10 (Figs 24, 26, 27), 0.30 (Fig. 23), 0.50 (Figs 28 – 30)." httpUri="https://zenodo.org/record/219881/files/figure.png" pageId="10" pageNumber="293">Figs 28–30</figureCitation>
). Trochanter, femur, distal third of patella and hand at base of the fingers distinctly granulated, more pronounced on paraxial faces. Lyrifissures as in
<figureCitation id="11CECCDDF401FF915F9BFBAC037AFB99" box="[920,1052,1055,1079]" captionStart="FIGURES 23 – 32" captionStartId="9.[151,269,1878,1902]" captionTargetBox="[170,1415,213,1845]" captionTargetId="figure@9.[151,1436,194,1856]" captionTargetPageId="9" captionText="FIGURES 23 – 32. Roncocreagris gepesi sp. nov., female holotype. 23. carapace; 24. anterior margin of carapace; 25. anterior and medial processes of left coxa I; 26. left chelicera; 27. partial view of fingers of left chelicera, showing galea; 28. right pedipalp minus chela, dorsal view; 29. right chela, dorsal view; 30. right chela, lateral view; 31. claw of left leg IV; 32. subterminal seta of left leg IV. Scale bars (in mm): 0.05 (Figs 25, 31, 32), 0.10 (Figs 24, 26, 27), 0.30 (Fig. 23), 0.50 (Figs 28 – 30)." httpUri="https://zenodo.org/record/219881/files/figure.png" pageId="10" pageNumber="293">Figs 28–30</figureCitation>
. Antiaxial face of trochanter and femur with one tiny tubercle at middle or slightly more distad; femur bearing one distal glandular pore. Patella with one micropore at base of pedicel. Chelal hand elliptic in dorsal view, maximum width at middle; three dorsal micropores on pedicel; two glandular pores on antiaxial face close to finger base. Fixed finger with 98 teeth, most of them apically cusped, dental row almost reaching level of trichobothrium
<emphasis id="BB810C4AF401FF91580AFB1C0348FB69" box="[1033,1070,1199,1223]" italics="true" pageId="10" pageNumber="293">esb</emphasis>
;
<emphasis id="BB810C4AF401FF91583CFB1C038CFB69" box="[1087,1258,1199,1223]" italics="true" pageId="10" pageNumber="293">nodus ramosus</emphasis>
short, reaching level of 5th distal tooth; trichobothrium
<emphasis id="BB810C4AF401FF915E48FB64053DFB40" box="[587,603,1239,1262]" italics="true" pageId="10" pageNumber="293">it</emphasis>
proximad of
<emphasis id="BB810C4AF401FF915EF2FB640463FB40" box="[753,773,1239,1262]" italics="true" pageId="10" pageNumber="293">et</emphasis>
, closer to
<emphasis id="BB810C4AF401FF915F7BFB6404F1FB40" box="[888,919,1239,1262]" italics="true" pageId="10" pageNumber="293">est</emphasis>
than to
<emphasis id="BB810C4AF401FF915FF3FB640362FB40" box="[1008,1028,1239,1262]" italics="true" pageId="10" pageNumber="293">et</emphasis>
; trichobothrium
<emphasis id="BB810C4AF401FF9158C2FB6403BAFB40" box="[1217,1244,1239,1262]" italics="true" pageId="10" pageNumber="293">ist</emphasis>
slightly closer to base of finger than to apex; distance between trichobothria
<emphasis id="BB810C4AF401FF915F36FB4A042DFABF" box="[821,843,1273,1297]" italics="true" pageId="10" pageNumber="293">ib</emphasis>
and
<emphasis id="BB810C4AF401FF915F80FB4904F8FABF" box="[899,926,1274,1297]" italics="true" pageId="10" pageNumber="293">ist</emphasis>
1.3 times longer than that between
<emphasis id="BB810C4AF401FF915931FB49022AFABF" box="[1330,1356,1274,1297]" italics="true" pageId="10" pageNumber="293">ist</emphasis>
and
<emphasis id="BB810C4AF401FF915987FB4902F5FABF" box="[1412,1427,1274,1297]" italics="true" pageId="10" pageNumber="293">it</emphasis>
; distance between trichobothria
<emphasis id="BB810C4AF401FF915E0AFAAE054CFA9B" box="[521,554,1309,1333]" italics="true" pageId="10" pageNumber="293">isb</emphasis>
and
<emphasis id="BB810C4AF401FF915E6EFAAC05EEFA98" box="[621,648,1311,1334]" italics="true" pageId="10" pageNumber="293">ist</emphasis>
1.7 times longer than that between
<emphasis id="BB810C4AF401FF915845FAAE0301FA9B" box="[1094,1127,1309,1333]" italics="true" pageId="10" pageNumber="293">isb</emphasis>
and
<emphasis id="BB810C4AF401FF9158A9FAAE03A6FA9B" box="[1194,1216,1309,1333]" italics="true" pageId="10" pageNumber="293">ib</emphasis>
; distance between trichobothria
<emphasis id="BB810C4AF401FF915D2DFAF10624FAF7" box="[302,322,1346,1369]" italics="true" pageId="10" pageNumber="293">et</emphasis>
and
<emphasis id="BB810C4AF401FF915D7BFAF106EEFAF7" box="[376,392,1346,1369]" italics="true" pageId="10" pageNumber="293">it</emphasis>
1.2 times longer that between
<emphasis id="BB810C4AF401FF915EDDFAF10588FAF7" box="[734,750,1346,1369]" italics="true" pageId="10" pageNumber="293">it</emphasis>
and
<emphasis id="BB810C4AF401FF915F20FAF10424FAF7" box="[803,834,1346,1369]" italics="true" pageId="10" pageNumber="293">est</emphasis>
. Chelal microsetae pattern:
<emphasis id="BB810C4AF401FF91587FFAF103C7FAF7" box="[1148,1185,1346,1369]" italics="true" pageId="10" pageNumber="293">Em</emphasis>
3 curved setae,
<emphasis id="BB810C4AF401FF915956FAF102E7FAF7" box="[1365,1409,1346,1369]" italics="true" pageId="10" pageNumber="293">Mm</emphasis>
2,
<emphasis id="BB810C4AF401FF915C94FAD407D2FAD0" box="[151,180,1383,1406]" italics="true" pageId="10" pageNumber="293">Im</emphasis>
1. Movable finger with 103 teeth, most of them apically cusped, dental row ending distad of trichobothrium
<emphasis id="BB810C4AF401FF915984FAD502F3FAD0" box="[1415,1429,1382,1406]" italics="true" pageId="10" pageNumber="293">b</emphasis>
, shorter than that of fixed finger. Distance between trichobothria s
<emphasis id="BB810C4AF401FF915F6CFA3A041BFA0F" box="[879,893,1417,1441]" italics="true" pageId="10" pageNumber="293">b</emphasis>
and
<emphasis id="BB810C4AF401FF915FB0FA3904A0FA0F" box="[947,966,1418,1441]" italics="true" pageId="10" pageNumber="293">st</emphasis>
1.2 times longer than that between
<emphasis id="BB810C4AF401FF915957FA3A020BFA0F" box="[1364,1389,1417,1441]" italics="true" pageId="10" pageNumber="293">sb</emphasis>
and
<emphasis id="BB810C4AF401FF915C94FA1D07C3FA68" box="[151,165,1454,1478]" italics="true" pageId="10" pageNumber="293">b</emphasis>
. One sensillum near the tip of both fingers; diploid sensillum
<emphasis id="BB810C4AF401FF915F8AFA1C04C2FA68" box="[905,932,1455,1478]" italics="true" pageId="10" pageNumber="293">pc</emphasis>
not raised, close to dental margin, slightly proximad of trichobothrium
<emphasis id="BB810C4AF401FF915DD5FA620689FA47" box="[470,495,1489,1513]" italics="true" pageId="10" pageNumber="293">sb</emphasis>
.
</paragraph>
<paragraph id="894AD058F401FF915CC4FA4503AAF99F" blockId="10.[151,1437,151,2029]" pageId="10" pageNumber="293">
Legs. Claws of legs I and IV with a tiny dorsal tooth proximad to middle (
<figureCitation id="11CECCDDF401FF915FFFFA460328F9A3" box="[1020,1102,1525,1549]" captionStart="FIGURES 23 – 32" captionStartId="9.[151,269,1878,1902]" captionTargetBox="[170,1415,213,1845]" captionTargetId="figure@9.[151,1436,194,1856]" captionTargetPageId="9" captionText="FIGURES 23 – 32. Roncocreagris gepesi sp. nov., female holotype. 23. carapace; 24. anterior margin of carapace; 25. anterior and medial processes of left coxa I; 26. left chelicera; 27. partial view of fingers of left chelicera, showing galea; 28. right pedipalp minus chela, dorsal view; 29. right chela, dorsal view; 30. right chela, lateral view; 31. claw of left leg IV; 32. subterminal seta of left leg IV. Scale bars (in mm): 0.05 (Figs 25, 31, 32), 0.10 (Figs 24, 26, 27), 0.30 (Fig. 23), 0.50 (Figs 28 – 30)." httpUri="https://zenodo.org/record/219881/files/figure.png" pageId="10" pageNumber="293">Fig. 31</figureCitation>
), subterminal setae with three rami, trident shaped (
<figureCitation id="11CECCDDF401FF915D86F9AA06BFF99F" box="[389,473,1561,1585]" captionStart="FIGURES 23 – 32" captionStartId="9.[151,269,1878,1902]" captionTargetBox="[170,1415,213,1845]" captionTargetId="figure@9.[151,1436,194,1856]" captionTargetPageId="9" captionText="FIGURES 23 – 32. Roncocreagris gepesi sp. nov., female holotype. 23. carapace; 24. anterior margin of carapace; 25. anterior and medial processes of left coxa I; 26. left chelicera; 27. partial view of fingers of left chelicera, showing galea; 28. right pedipalp minus chela, dorsal view; 29. right chela, dorsal view; 30. right chela, lateral view; 31. claw of left leg IV; 32. subterminal seta of left leg IV. Scale bars (in mm): 0.05 (Figs 25, 31, 32), 0.10 (Figs 24, 26, 27), 0.30 (Fig. 23), 0.50 (Figs 28 – 30)." httpUri="https://zenodo.org/record/219881/files/figure.png" pageId="10" pageNumber="293">Fig. 32</figureCitation>
). Leg IV tibia TS ratio 0.48, basitarsus TS 0.13, telotarsus TS 0.27.
</paragraph>
<paragraph id="894AD058F401FF915CC4F98D0592F8F4" blockId="10.[151,1437,151,2029]" pageId="10" pageNumber="293">
<emphasis id="BB810C4AF401FF915CC4F98D0565F9F8" bold="true" box="[199,515,1597,1622]" pageId="10" pageNumber="293">Measurements and ratios.</emphasis>
Body 2.19. Carapace 0.95/0.65 (1.5). Chelicera: palm 0.57/0.27 (2.1), movable finger 0.36. Pedipalp: trochanter 0.79/0.21 (3.9); femur 1.46/0.22 (6.6); patella 1.41/0.26 (5.5), pedicel 0.36, club 1.05/0.26 (4.1), ratio club/pedicel: 2.92; chela+ 2.39/0.41 (5.9), chela - 2.21/0.41 (5.5); hand+ 1.03 (2.5), hand - 0.85 (2.1); movable finger 1.39; ratio finger/hand+ 1.4; ratio chela+/carapace 2.5; femur/carapace 1.5; femur/finger 1.1; femur/patella 1.0; patella/ hand+ 1.4. Leg I: femur 0.73/0.14 (5.4); patella 0.51/0.12 (4.4); tibia 0.68/.08 (8.7); basitarsus 0.33/0.07 (5.0); telotarsus 0.45/0.06 (8.2); ratio femur/patella 1.4; telotarsus/basitarsus 1.4. Leg IV: femur+patella 1.12/0.25 (4.5); tibia 1.12/0.13 (8.4); basitarsus 0.41/0.1 (4.2); telotarsus 0.58/0.08 (7.4); ratio femur+patella/tibia 1.0; ratio telotarsus/basitarsus 1.4.
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F401FF905CC4F8D504C4FEB2" lastPageId="11" lastPageNumber="294" pageId="10" pageNumber="293" type="discussion">
<paragraph id="894AD058F401FF905CC4F8D504C4FEB2" blockId="10.[151,1437,151,2029]" lastBlockId="11.[151,1436,151,464]" lastPageId="11" lastPageNumber="294" pageId="10" pageNumber="293">
<emphasis id="BB810C4AF401FF915CC4F8D5065DF8D0" bold="true" box="[199,315,1894,1918]" pageId="10" pageNumber="293">Remarks.</emphasis>
The new species is easily distinguishable from
<taxonomicName id="4EF5ABDBF401FF915F5BF8D404ADF8D0" box="[856,971,1894,1918]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="293" phylum="Arthropoda" rank="species" species="borgesi" status="sp. nov.">
<emphasis id="BB810C4AF401FF915F5BF8D404ADF8D0" box="[856,971,1894,1918]" italics="true" pageId="10" pageNumber="293">R. borgesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0B2B131F401FF915FD0F8D5034BF8D0" box="[979,1069,1894,1918]" pageId="10" pageNumber="293" rank="species">
<emphasis id="BB810C4AF401FF915FD0F8D5034BF8D0" bold="true" box="[979,1069,1894,1918]" pageId="10" pageNumber="293">sp. nov.</emphasis>
</taxonomicNameLabel>
by the shape of the chelal hand, which is elliptic and widest at middle in
<taxonomicName id="4EF5ABDBF401FF915E5BF83805DBF80C" box="[600,701,1931,1954]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="293" phylum="Arthropoda" rank="species" species="gepesi">
<emphasis id="BB810C4AF401FF915E5BF83805DBF80C" box="[600,701,1931,1954]" italics="true" pageId="10" pageNumber="293">R. gepesi</emphasis>
</taxonomicName>
versus oval and widest proximad of middle in
<taxonomicName id="4EF5ABDBF401FF9158C4F838025EF80C" box="[1223,1336,1930,1954]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="293" phylum="Arthropoda" rank="species" species="borgesi">
<emphasis id="BB810C4AF401FF9158C4F838025EF80C" box="[1223,1336,1930,1954]" italics="true" pageId="10" pageNumber="293">R. borgesi</emphasis>
</taxonomicName>
; also the pedipalps are larger and more slender in
<taxonomicName id="4EF5ABDBF401FF915E6BF81C05B6F868" box="[616,720,1967,1990]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="293" phylum="Arthropoda" rank="species" species="gepesi">
<emphasis id="BB810C4AF401FF915E6BF81C05B6F868" box="[616,720,1967,1990]" italics="true" pageId="10" pageNumber="293">R. gepesi</emphasis>
</taxonomicName>
. The new species can be differentiated from
<taxonomicName id="4EF5ABDBF401FF9158D0F81C021EF868" box="[1235,1400,1966,1990]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="293" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="BB810C4AF401FF9158D0F81C021EF868" box="[1235,1400,1966,1990]" italics="true" pageId="10" pageNumber="293">R. cavernicola</emphasis>
</taxonomicName>
by the chelal movable finger/hand+ ratio (1.4, versus 1.7) and by the chaetotaxy of the posterior margin of the carapace (7 setae, versus 4) and of tergite I (6 setae, versus 4).
<taxonomicName id="4EF5ABDBF400FF905EFEFF2A0496FF1E" box="[765,1008,153,176]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="294" phylum="Arthropoda" rank="species" species="gepesi">
<emphasis id="BB810C4AF400FF905EFEFF2A0496FF1E" box="[765,1008,153,176]" italics="true" pageId="11" pageNumber="294">Roncocreagris gepesi</emphasis>
</taxonomicName>
is more closely related to
<taxonomicName id="4EF5ABDBF400FF905922FF2A0621FF7A" authority="Zaragoza, 2002" authorityName="Zaragoza" authorityYear="2002" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="294" phylum="Arthropoda" rank="species" species="lucensis">
<emphasis id="BB810C4AF400FF905922FF2A02FDFF1E" box="[1313,1435,152,176]" italics="true" pageId="11" pageNumber="294">R. lucensis</emphasis>
<bibRefCitation id="ED64ADA9F400FF905C94FF0E0621FF7A" author="Zaragoza" box="[151,327,188,213]" pageId="11" pageNumber="294" refString="Zaragoza, J. A. (2002) Dos nuevos Roncocreagris cavernicolas de Galicia (Arachnida, Pseudoscorpiones, Neobisiidae). Revista Iberica de Aracnologia, 5, 91 - 98." type="journal article" year="2002">Zaragoza, 2002</bibRefCitation>
</taxonomicName>
(from Galicia,
<collectingCountry id="F1E290C8F400FF905DFAFF0F055BFF7A" box="[505,573,188,212]" name="Spain" pageId="11" pageNumber="294">Spain</collectingCountry>
), despite being
<quantity id="4E0D7DBDF400FF905EF2FF0F042EFF7A" box="[753,840,188,212]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="294" unit="km" value="400.0">400 km</quantity>
away, but in the latter the pedipalps are shorter and less slender than in
<emphasis id="BB810C4AF400FF905D72FF52068CFF56" box="[369,490,224,248]" italics="true" pageId="11" pageNumber="294">
<taxonomicName id="4EF5ABDBF400FF905D72FF520680FF56" box="[369,486,224,248]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="294" phylum="Arthropoda" rank="species" species="borgesi">R. borgesi</taxonomicName>
,
</emphasis>
and the distance between trichobothria
<emphasis id="BB810C4AF400FF905FA7FF5204DEFF56" box="[932,952,225,248]" italics="true" pageId="11" pageNumber="294">et</emphasis>
and
<emphasis id="BB810C4AF400FF905FEDFF520498FF56" box="[1006,1022,225,248]" italics="true" pageId="11" pageNumber="294">it</emphasis>
is 3.3 times longer than that between
<emphasis id="BB810C4AF400FF905C94FEB607C1FEB2" box="[151,167,261,284]" italics="true" pageId="11" pageNumber="294">it</emphasis>
and
<emphasis id="BB810C4AF400FF905CDEFEB6079AFEB2" box="[221,252,261,284]" italics="true" pageId="11" pageNumber="294">est</emphasis>
in the Spanish species, versus
<quantity id="4E0D7DBDF400FF905E55FEB705F0FEB2" box="[598,662,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="11" pageNumber="294" unit="in" value="1.2">1.2 in</quantity>
the Portuguese species.
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F400FF905CC4FE94036CFE7E" pageId="11" pageNumber="294" type="distribution">
<paragraph id="894AD058F400FF905CC4FE94036CFE7E" blockId="11.[151,1436,151,464]" pageId="11" pageNumber="294">
<emphasis id="BB810C4AF400FF905CC4FE940695FEEE" bold="true" box="[199,499,295,320]" pageId="11" pageNumber="294">Distribution and habitat.</emphasis>
<taxonomicName id="4EF5ABDBF400FF905DF8FE9A0588FEEE" box="[507,750,297,320]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="294" phylum="Arthropoda" rank="species" species="gepesi" status="sp. nov.">
<emphasis id="BB810C4AF400FF905DF8FE9A0588FEEE" box="[507,750,297,320]" italics="true" pageId="11" pageNumber="294">Roncocreagris gepesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0B2B131F400FF905EF4FE9B0437FEEE" box="[759,849,296,320]" pageId="11" pageNumber="294" rank="species">
<emphasis id="BB810C4AF400FF905EF4FE9B0437FEEE" bold="true" box="[759,849,296,320]" pageId="11" pageNumber="294">sp. nov.</emphasis>
</taxonomicNameLabel>
was collected in Algarinho, one of the caves of the Dueça system, opened in 1998 with the aid of a backhoe (
<bibRefCitation id="ED64ADA9F400FF905F4FFEFF0342FECA" author="Neves" box="[844,1060,332,356]" pageId="11" pageNumber="294" refString="Neves, J., Pessoa, M. &amp; Redinha, N. (2005) O Sistema Espeleologico do Dueca. Espeleodivulgacao, 6, 14 - 18." type="journal article" year="2005">
Neves
<emphasis id="BB810C4AF400FF905F9CFEFE04BAFECA" box="[927,988,332,356]" italics="true" pageId="11" pageNumber="294">et al.</emphasis>
2005
</bibRefCitation>
). This species was not found in Soprador do Carvalho, a cave located only
<quantity id="4E0D7DBDF400FF905EADFEDC059BFE29" box="[686,765,367,391]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="294" unit="m" value="200.0">200 m</quantity>
from the entrance of Algarinho Cave. The mean cave temperature is 14.9ºC at soil level. The new species shares its habitat with other hypogean species, such as the woodlouse
<taxonomicName id="4EF5ABDBF400FF905D14FE0B0572FE7E" box="[279,532,440,464]" class="Malacostraca" family="Porcellionidae" genus="Porcellio" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="294" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="BB810C4AF400FF905D14FE0B0572FE7E" box="[279,532,440,464]" italics="true" pageId="11" pageNumber="294">Porcellio cavernicolus</emphasis>
</taxonomicName>
and the dipluran
<taxonomicName id="4EF5ABDBF400FF905EDBFE0B0438FE7E" box="[728,862,440,464]" class="Entognatha" family="Campodeidae" genus="Podocampa" kingdom="Animalia" order="Diplura" pageId="11" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="BB810C4AF400FF905EDBFE0B0438FE7E" box="[728,862,440,464]" italics="true" pageId="11" pageNumber="294">Podocampa</emphasis>
</taxonomicName>
cf.
<emphasis id="BB810C4AF400FF905F8AFE0B036CFE7E" box="[905,1034,440,464]" italics="true" pageId="11" pageNumber="294">fragiloides.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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