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<document id="CB0F1E96A4B8883261A8514B2A19DCFA" ID-CLB-Dataset="30935" ID-DOI="10.11646/zootaxa.4375.2.1" ID-GBIF-Dataset="c8d905a3-01d4-4d5f-a904-fad72add2ae1" ID-ISSN="1175-5326" ID-Zenodo-Dep="1158445" ID-ZooBank="7A4595B8-9FDF-41A9-9C39-A8A4A83FF302" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1516789970485" checkinUser="plazi" docAuthor="Phan, Quoc Toan, Kompier, Tom, Karube, Haruki &amp; Hayashi, Fumio" docDate="2018" docId="006F87D3FFFCEC02B594F99EFB05D1DE" docLanguage="en" docName="zootaxa.4375.2.1.pdf" docOrigin="Zootaxa 4375 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euphaea pahyapi : Hamalainen 1985" docType="treatment" docVersion="7" lastPageNumber="177" masterDocId="FC56FFABFFE5EC18B503FF87FFBFD376" masterDocTitle="A synopsis of the Euphaeidae (Odonata: Zygoptera) of Vietnam, with descriptions of two new species of Euphaea" masterLastPageNumber="190" masterPageNumber="151" pageNumber="176" updateTime="1698516995768" updateUser="ExternalLinkService">
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<mods:title id="12A3312C9081943AE45891715974A69F">A synopsis of the Euphaeidae (Odonata: Zygoptera) of Vietnam, with descriptions of two new species of Euphaea</mods:title>
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<mods:namePart id="07EE7873C9367425795D717A1EAF6E8E">Phan, Quoc Toan</mods:namePart>
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<taxonomicName id="4FC64D46FFFCEC01B594F99EFDEAD545" ID-CoL="3CLZH" authority="Hamalainen, 1985" authorityName=": Hamalainen" authorityYear="1985" box="[151,597,1561,1587]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="176" phylum="Arthropoda" rank="species" species="pahyapi">
<emphasis id="BAB2EAD7FFFCEC01B594F99EFDEAD545" bold="true" box="[151,597,1561,1587]" pageId="25" pageNumber="176">
<emphasis id="BAB2EAD7FFFCEC01B594F99EFED4D545" bold="true" box="[151,363,1561,1587]" italics="true" pageId="25" pageNumber="176">Euphaea pahyapi</emphasis>
<bibRefCitation id="EC574B34FFFCEC01B470F99EFDEAD545" author="Hamalainen" box="[371,597,1561,1587]" pageId="25" pageNumber="176" refString="Hamalainen, M. (1985) Euphaea pahyapi sp. nov. from Peninsular Thailand (Zygoptera: Euphaeidae). Odonatologica, 14 (4), 341 - 344." type="journal article" year="1985">Hämäläinen, 1985</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="887936C5FFFCEC01B594F9BBFE2FD523" blockId="25.[151,597,1561,1621]" box="[151,400,1596,1621]" pageId="25" pageNumber="176">
(
<figureCitation id="10FD2A40FFFCEC01B59CF9BBFEBBD523" box="[159,260,1596,1621]" captionStart="FIGURE15" captionStartId="24.[151,250,929,951]" captionTargetBox="[186,1410,221,879]" captionTargetId="figure@24.[153,1433,193,907]" captionTargetPageId="24" captionText="FIGURE15.Schematic drawingsof abdominal hair tufts inlateral view ofmale Euphaeaspp.(15a) E.saola sp.nov.; (15b)E. guerini; (15c) E.hirta; (15d) E.masoni and E. pahyapi. IX show abdominal segments S110." httpUri="https://zenodo.org/record/3096107/files/figure.png" pageId="25" pageNumber="176">Figs 15d</figureCitation>
,
<figureCitation id="10FD2A40FFFCEC01B40CF9BBFE88D523" box="[271,311,1596,1621]" captionStart="FIGURE 18" captionStartId="26.[151,250,1894,1916]" captionTargetBox="[335,1233,745,1853]" captionTargetId="figure@26.[313,1273,735,1872]" captionTargetPageId="26" captionText="FIGURE 18. Euphaea spp. [18af] vesicle of male and [18gj] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc." httpUri="https://zenodo.org/record/3096117/files/figure.png" pageId="25" pageNumber="176">18f</figureCitation>
,
<figureCitation id="10FD2A40FFFCEC01B440F9BBFE36D523" box="[323,393,1596,1621]" captionStart="FIGURE 19" captionStartId="27.[151,250,1020,1042]" captionTargetBox="[168,1415,193,965]" captionTargetId="figure@27.[153,1433,193,998]" captionTargetPageId="27" captionText="FIGURE 19. Euphaea pahyapi. [19ad] ♂, Phu Quoc. (19a) head and thorax; (19b) head, antero-dorsal view; (19c) appendages, lateral view; (19d) wings." httpUri="https://zenodo.org/record/3096125/files/figure.png" pageId="25" pageNumber="176">19ad</figureCitation>
)
</paragraph>
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<paragraph id="887936C5FFFCEC01B594F903FF40D5CE" blockId="25.[151,1436,1668,1843]" pageId="25" pageNumber="176">
<taxonomicName id="4FC64D46FFFCEC01B594F903FDA2D5EC" ID-CoL="3CLZH" authority=": Hamalainen (1985)" authorityName=": Hamalainen" authorityYear="1985" box="[151,541,1668,1690]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="176" phylum="Arthropoda" rank="species" species="pahyapi">
<emphasis id="BAB2EAD7FFFCEC01B594F903FEF6D5EC" box="[151,329,1668,1690]" italics="true" pageId="25" pageNumber="176">Euphaea pahyapi</emphasis>
:
<bibRefCitation id="EC574B34FFFCEC01B454F903FDA2D5EC" author="Hamalainen" box="[343,541,1668,1690]" pageId="25" pageNumber="176" refString="Hamalainen, M. (1985) Euphaea pahyapi sp. nov. from Peninsular Thailand (Zygoptera: Euphaeidae). Odonatologica, 14 (4), 341 - 344." type="journal article" year="1985">Hämäläinen (1985)</bibRefCitation>
</taxonomicName>
, pp. 341344,
<figureCitation id="10FD2A40FFFCEC01B7B7F903FCB1D5EC" box="[692,782,1668,1690]" captionStart-0="FIGURE1" captionStart-1="FIGURE 2" captionStart-2="FIGURE3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1517,1539]" captionStartId-1="4.[151,250,1803,1825]" captionStartId-2="6.[151,250,990,1012]" captionStartId-3="7.[151,250,1647,1669]" captionTargetBox-0="[233,1358,627,1465]" captionTargetBox-1="[295,1288,877,1761]" captionTargetBox-2="[186,1420,213,944]" captionTargetBox-3="[318,1254,631,1600]" captionTargetId-0="figure@3.[217,1369,615,1495]" captionTargetId-1="figure@4.[281,1305,857,1781]" captionTargetId-2="figure@6.[153,1433,195,968]" captionTargetId-3="figure@7.[315,1272,622,1626]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE1. Anisopleurabipugio. [1ac] ♂,A Luoi and [1dh]♀, Hon Ba.(1a)head and thorax;(1b)appendages,dorsal view; (1c) appendages, lateral view; (1d) body; (1e) head, frontal view; (1f) prothorax,lateral view; (1g) prothorax, dorsal view; (1h) basal half of HW." captionText-1="FIGURE 2. Anisopleura qingyuanensis. [2ad] ♂ and [2ef] ♀, Tu Le. (2a) head and thorax; (2b) head, oblique-dorsal view; (2c) appendages, dorsal view; (2d) appendages, lateral view; (2e) head and thorax; (2f) head, oblique-dorsal view." captionText-2="FIGURE3. Anisopleurayunnanensis. [3ac] ♂,Yunnan and [3d]♀,Sapa.(3a) head and thorax; (3b) appendages, dorsalview; (3c) appendages, lateral view; (3d) head and thorax." captionText-3="FIGURE 4. Bayadera bidentata. [3ae] ♂, Tam Dao and [4fh] ♀. (4a) head and thorax; (4b) appendages, dorsal view; (4c) appendages, ventral view; (4d) appendages, lateral view; (4e) wings; (4f) head and thorax, Guangdong, China; (4g) head and thorax, Guangxi, China, and Vietnam; (4h) abdominal tip." httpUri-0="https://zenodo.org/record/3096039/files/figure.png" httpUri-1="https://zenodo.org/record/3096041/files/figure.png" httpUri-2="https://zenodo.org/record/3096045/files/figure.png" httpUri-3="https://zenodo.org/record/3096051/files/figure.png" pageId="25" pageNumber="176">Figs 14</figureCitation>
;
<bibRefCitation id="EC574B34FFFCEC01B619F903FB82D5EC" author="Kitagawa" box="[794,1085,1668,1690]" pageId="25" pageNumber="176" refString="Kitagawa, K. &amp; Katatani, N. (2003) Notes on the Odonata of Thailand, Part 2. The Odonata from Trang Prefecture in March 1998. Aeschna, 40, 13 - 23. [in Japanese]" type="journal article" year="2003">Kitagawa &amp; Katatani (2003)</bibRefCitation>
, p. 16,
<figureCitation id="10FD2A40FFFCEC01B18BF903FB79D5EC" box="[1160,1222,1668,1690]" captionStart="FIGURE 4" captionStartId="7.[151,250,1647,1669]" captionTargetBox="[318,1254,631,1600]" captionTargetId="figure@7.[315,1272,622,1626]" captionTargetPageId="7" captionText="FIGURE 4. Bayadera bidentata. [3ae] ♂, Tam Dao and [4fh] ♀. (4a) head and thorax; (4b) appendages, dorsal view; (4c) appendages, ventral view; (4d) appendages, lateral view; (4e) wings; (4f) head and thorax, Guangdong, China; (4g) head and thorax, Guangxi, China, and Vietnam; (4h) abdominal tip." httpUri="https://zenodo.org/record/3096051/files/figure.png" pageId="25" pageNumber="176">Fig. 4</figureCitation>
;
<bibRefCitation id="EC574B34FFFCEC01B1D2F903FA28D5EC" author="Hamalainen" box="[1233,1431,1668,1690]" pageId="25" pageNumber="176" refString="Hamalainen, M. (2017) The Caloptera damselflies of Thailand - Distribution maps by provinces (Odonata: Calopterygoidea). Faunistic studies in South-East Asian and Pacific Island Odonata, 19, 1 - 28." type="journal article" year="2017">Hämäläinen (2017)</bibRefCitation>
, p. 11.
</paragraph>
<paragraph id="887936C5FFFCEC01B594F946FC85D5A1" blockId="25.[151,1436,1668,1843]" box="[151,826,1729,1751]" pageId="25" pageNumber="176">
<taxonomicName id="4FC64D46FFFCEC01B594F946FDFAD5A1" ID-CoL="3CLZC" authority=": Asahina (1985)" authorityName=": Asahina" authorityYear="1985" box="[151,581,1729,1751]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="176" phylum="Arthropoda" rank="species" species="khaochongensis">
<emphasis id="BAB2EAD7FFFCEC01B594F946FE26D5A1" box="[151,409,1729,1751]" italics="true" pageId="25" pageNumber="176">Euphaea khaochongensis</emphasis>
:
<bibRefCitation id="EC574B34FFFCEC01B4A5F946FDFAD5A1" author="Asahina" box="[422,581,1729,1751]" pageId="25" pageNumber="176" refString="Asahina, S. (1985) A list of the Odonata recorded from Thailand. Part XI. Euphaeidae. Cho Cho, 8 (12), 18 - 38." type="journal article" year="1985">Asahina (1985)</bibRefCitation>
</taxonomicName>
, pp. 2729, Figs 2528.
</paragraph>
<paragraph id="887936C5FFFCEC01B594F967FD46D462" blockId="25.[151,1436,1668,1843]" pageId="25" pageNumber="176">
<taxonomicName id="4FC64D46FFFCEC01B594F967FE6DD580" ID-CoL="9WTZV" authority=": Bui (2007)" authorityName=": Bui" authorityYear="2007" box="[151,466,1760,1782]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="176" phylum="Arthropoda" rank="species" species="ochracea">
<emphasis id="BAB2EAD7FFFCEC01B594F967FEEAD580" box="[151,341,1760,1782]" italics="true" pageId="25" pageNumber="176">Euphaea ochracea</emphasis>
:
<bibRefCitation id="EC574B34FFFCEC01B461F967FE6DD580" author="Bui" box="[354,466,1760,1782]" pageId="25" pageNumber="176" refString="Bui, H. M. (2007) Danh l uc b ang hinh anh cac loai chu on chu on Phu Qu oc [Approximate translation: &quot; Photographic checklist of Odonata from Phu Quoc Island &quot;]. Wildlife At Risk Published as a book. Wildlife At Risk, Vietnam, 55 pp. (in Vietnamese). Available from: http: // www. wildlifeatrisk. org / upload / download / books. P 10. % 20 PQ-Dragonfly-book. pdf (accessed 26 November 2017)" type="book" year="2007">Bui (2007)</bibRefCitation>
</taxonomicName>
, pp. 1, 6 (photos) [Specimens from Phu Quoc];
<bibRefCitation id="EC574B34FFFCEC01B6B6F967FB2FD580" box="[949,1168,1760,1782]" pageId="25" pageNumber="176" refString="Kosterin, O. E., Chartier, G., Holden, J. &amp; Mey, F. S. (2012) New records of Odonata from Cambodia, based mostly on photographs. Cambodian Journal of Natural History, 2012 (2), 150 - 163." type="journal article">
Kosterin
<emphasis id="BAB2EAD7FFFCEC01B111F966FBFAD580" box="[1042,1093,1760,1782]" italics="true" pageId="25" pageNumber="176">et al.</emphasis>
(2012)
</bibRefCitation>
, p. 153,
<figureCitation id="10FD2A40FFFCEC01B1E6F967FAA0D580" box="[1253,1311,1760,1782]" captionStart="FIGURE 2" captionStartId="4.[151,250,1803,1825]" captionTargetBox="[295,1288,877,1761]" captionTargetId="figure@4.[281,1305,857,1781]" captionTargetPageId="4" captionText="FIGURE 2. Anisopleura qingyuanensis. [2ad] ♂ and [2ef] ♀, Tu Le. (2a) head and thorax; (2b) head, oblique-dorsal view; (2c) appendages, dorsal view; (2d) appendages, lateral view; (2e) head and thorax; (2f) head, oblique-dorsal view." httpUri="https://zenodo.org/record/3096041/files/figure.png" pageId="25" pageNumber="176">Fig. 2</figureCitation>
(Specimens from Kbal Chhay Cascades, Preah Sihanouk Province).
</paragraph>
<paragraph id="887936C5FFFCEC01B594F89AFDB7D445" blockId="25.[151,1436,1668,1843]" box="[151,520,1821,1843]" pageId="25" pageNumber="176">
<taxonomicName id="4FC64D46FFFCEC01B594F89AFF4ED445" ID-CoL="4G8X" authorityName="Selys" authorityYear="1840" box="[151,241,1821,1843]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="176" phylum="Arthropoda" rank="genus">
<emphasis id="BAB2EAD7FFFCEC01B594F89AFF4ED445" box="[151,241,1821,1843]" italics="true" pageId="25" pageNumber="176">Euphaea</emphasis>
</taxonomicName>
sp.:
<bibRefCitation id="EC574B34FFFCEC01B422F89AFE7BD445" author="Kosterin" box="[289,452,1821,1843]" pageId="25" pageNumber="176" refString="Kosterin, O. E. (2016) A survey of Odonata of Mondulkiri, the elevated eastern province of Cambodia, for ten days in June 2016. International Dragonfly Fund Report. Journal of the International Dragonfly Fund, 98, 1 - 85." type="journal article" year="2016">Kosterin (2016)</bibRefCitation>
, p. 19.
</paragraph>
</subSubSection>
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<paragraph id="887936C5FFFCEC01B594F8D8FD13D4EA" blockId="25.[151,1436,1887,2021]" pageId="25" pageNumber="176">
<materialsCitation id="38AE3C98FFFCEC01B594F8D8FADAD40E" ID-GBIF-Occurrence="1805098072" box="[151,1381,1887,1912]" collectingDate="2015-12-30" collectorName="T. Kompier" country="Vietnam" location="Phu Quoc Isl." municipality="Materials" pageId="25" pageNumber="176" specimenCount="1" specimenCount-male="1" stateProvince="Kien Giang">
<emphasis id="BAB2EAD7FFFCEC01B594F8D8FE33D40E" bold="true" box="[151,396,1887,1912]" pageId="25" pageNumber="176">
<collectingMunicipality id="681DACBFFFFCEC01B594F8D8FEB5D40E" box="[151,266,1887,1912]" pageId="25" pageNumber="176">Materials</collectingMunicipality>
examined.
</emphasis>
[
<collectingCountry id="F0D17655FFFCEC01B498F8D8FDBFD40E" box="[411,512,1887,1912]" name="Vietnam" pageId="25" pageNumber="176">Vietnam</collectingCountry>
]
<specimenCount id="9EC0FD4CFFFCEC01B70CF8E7FD8CD40E" box="[527,563,1888,1912]" pageId="25" pageNumber="176" type="male">1♂</specimenCount>
,
<location id="8D19601EFFFCEC01B742F8D8FD60D40E" LSID="urn:lsid:plazi:treatment:006F87D3FFFCEC02B594F99EFB05D1DE:8D19601EFFFCEC01B742F8D8FD60D40E" box="[577,735,1887,1912]" country="Vietnam" municipality="Materials" name="Phu Quoc Isl." pageId="25" pageNumber="176" stateProvince="Kien Giang">Phu Quoc Isl.</location>
,
<collectingRegion id="4A02F827FFFCEC01B7EEF8D8FCCED40E" box="[749,881,1887,1912]" country="Vietnam" name="Kien Giang" pageId="25" pageNumber="176">Kien Giang</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFFCEC01B6C1F8D8FBF2D40E" box="[962,1101,1887,1912]" pageId="25" pageNumber="176" value="2015-12-30">
<date id="FC781005FFFCEC01B6C1F8D8FBF2D40E" box="[962,1101,1887,1912]" pageId="25" pageNumber="176" value="2015-12-30">30.XII.2015</date>
</collectingDate>
,
<collectorName id="25335313FFFCEC01B159F8E7FB62D40E" box="[1114,1245,1887,1912]" pageId="25" pageNumber="176">T. Kompier</collectorName>
leg. (TKC)
</materialsCitation>
;
<materialsCitation id="38AE3C98FFFCEC01B071F8E7FD13D4EA" ID-GBIF-Occurrence="1805098079" collectingDate="2016-04-14" collectorName="T. Kompier" country="Vietnam" location="Phu Quoc Isl." municipality="Materials" pageId="25" pageNumber="176" specimenCount="2" specimenCount-male="2" stateProvince="Kien Giang">
<specimenCount id="9EC0FD4CFFFCEC01B071F8E7FA27D40E" box="[1394,1432,1888,1912]" pageId="25" pageNumber="176" type="male">2♂</specimenCount>
, same location and collector,
<collectingDate id="EC3CE9EDFFFCEC01B4D4F803FDECD4EB" box="[471,595,1924,1949]" pageId="25" pageNumber="176" value="2016-04-14">
<date id="FC781005FFFCEC01B4D4F803FDECD4EB" box="[471,595,1924,1949]" pageId="25" pageNumber="176" value="2016-04-14">14.IV.2016</date>
</collectingDate>
(TKC).
</materialsCitation>
</paragraph>
<paragraph id="887936C5FFFCEC01B5C4F82FFE0ED493" blockId="25.[151,1436,1887,2021]" pageId="25" pageNumber="176">
<materialsCitation id="38AE3C98FFFCEC01B5C4F82FFE0ED493" ID-GBIF-Occurrence="1805098085" collectingDate="2016-08-19" collectingDateMax="2016-08-20" collectingDateMin="2016-08-19" collectorName="T. Kompier." country="Vietnam" location="Phu Quoc Isl." pageId="25" pageNumber="176" specimenCount="1" stateProvince="Kien Giang">
<emphasis id="BAB2EAD7FFFCEC01B5C4F82FFCA6D4B7" bold="true" box="[199,793,1960,1985]" pageId="25" pageNumber="176">Other materials confirmed by field observations.</emphasis>
[
<collectingCountry id="F0D17655FFFCEC01B62AF82FFC31D4B7" box="[809,910,1960,1985]" name="Vietnam" pageId="25" pageNumber="176">Vietnam</collectingCountry>
] 6 exs,
<collectingDate id="EC3CE9EDFFFCEC01B6E9F82FFB15D4B7" box="[1002,1194,1960,1985]" pageId="25" pageNumber="176" value="2016-08-19" valueMax="2016-08-20" valueMin="2016-08-19">
<date id="FC781005FFFCEC01B6E9F82FFB15D4B7" box="[1002,1194,1960,1985]" pageId="25" pageNumber="176" value="2016-08-19" valueMax="2016-08-20" valueMin="2016-08-19">1920.VIII.2016</date>
</collectingDate>
,
<location id="8D19601EFFFCEC01B1B4F82FFAE8D4B7" LSID="urn:lsid:plazi:treatment:006F87D3FFFCEC02B594F99EFB05D1DE:8D19601EFFFCEC01B1B4F82FFAE8D4B7" box="[1207,1367,1960,1985]" country="Vietnam" name="Phu Quoc Isl." pageId="25" pageNumber="176" stateProvince="Kien Giang">Phu Quoc Isl.</location>
,
<collectingRegion id="4A02F827FFFCEC01B065F82FFF63D493" country="Vietnam" name="Kien Giang" pageId="25" pageNumber="176">Kien Giang</collectingRegion>
Prov.,
<collectorName id="25335313FFFCEC01B429F84AFE0ED493" box="[298,433,1996,2021]" pageId="25" pageNumber="176">T. Kompier.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="887936C5FFFFEC02B5C4FF10FD53D16E" blockId="26.[151,1436,151,680]" pageId="26" pageNumber="177">
<emphasis id="BAB2EAD7FFFFEC02B5C4FF10FEAED3C6" bold="true" box="[199,273,151,176]" pageId="26" pageNumber="177">Notes.</emphasis>
This species is newly recorded for
<collectingCountry id="F0D17655FFFFEC02B7B4FF10FCA6D3C6" box="[695,793,151,176]" name="Vietnam" pageId="26" pageNumber="177">Vietnam</collectingCountry>
, although
<bibRefCitation id="EC574B34FFFFEC02B697FF10FBADD3C6" author="Bui" box="[916,1042,151,176]" pageId="26" pageNumber="177" refString="Bui, H. M. (2007) Danh l uc b ang hinh anh cac loai chu on chu on Phu Qu oc [Approximate translation: &quot; Photographic checklist of Odonata from Phu Quoc Island &quot;]. Wildlife At Risk Published as a book. Wildlife At Risk, Vietnam, 55 pp. (in Vietnamese). Available from: http: // www. wildlifeatrisk. org / upload / download / books. P 10. % 20 PQ-Dragonfly-book. pdf (accessed 26 November 2017)" type="book" year="2007">Bui (2007)</bibRefCitation>
published the same taxon first as
<taxonomicName id="4FC64D46FFFFEC02B594FF3BFED5D3A3" authorityName=": Bui" authorityYear="2007" box="[151,362,188,213]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="ochracea">
<emphasis id="BAB2EAD7FFFFEC02B594FF3BFED5D3A3" box="[151,362,188,213]" italics="true" pageId="26" pageNumber="177">Euphaea ochracea</emphasis>
</taxonomicName>
.
<bibRefCitation id="EC574B34FFFFEC02B47BFF3BFDEED3A3" author="Hamalainen" box="[376,593,188,213]" pageId="26" pageNumber="177" refString="Hamalainen, M. (1985) Euphaea pahyapi sp. nov. from Peninsular Thailand (Zygoptera: Euphaeidae). Odonatologica, 14 (4), 341 - 344." type="journal article" year="1985">Hämäläinen (1985)</bibRefCitation>
described
<taxonomicName id="4FC64D46FFFFEC02B7CFFF3BFC2ED3A3" authorityName=": Hamalainen" authorityYear="1985" box="[716,913,188,213]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="pahyapi">
<emphasis id="BAB2EAD7FFFFEC02B7CFFF3BFC2ED3A3" box="[716,913,188,213]" italics="true" pageId="26" pageNumber="177">Euphaea pahyapi</emphasis>
</taxonomicName>
from the peninsular part of
<collectingCountry id="F0D17655FFFFEC02B1CDFF3BFA8DD3A3" box="[1230,1330,188,213]" name="Thailand" pageId="26" pageNumber="177">Thailand</collectingCountry>
based on male specimens.
<bibRefCitation id="EC574B34FFFFEC02B45BFF58FDB9D38E" author="Asahina" box="[344,518,223,248]" pageId="26" pageNumber="177" refString="Asahina, S. (1985) A list of the Odonata recorded from Thailand. Part XI. Euphaeidae. Cho Cho, 8 (12), 18 - 38." type="journal article" year="1985">Asahina (1985)</bibRefCitation>
also described
<taxonomicName id="4FC64D46FFFFEC02B7B7FF58FC6CD38E" authorityName=": Asahina" authorityYear="1985" box="[692,979,223,248]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="khaochongensis">
<emphasis id="BAB2EAD7FFFFEC02B7B7FF58FC6CD38E" box="[692,979,223,248]" italics="true" pageId="26" pageNumber="177">Euphaea khaochongensis</emphasis>
</taxonomicName>
from another province of the same area, but this was treated as a junior synonym of
<taxonomicName id="4FC64D46FFFFEC02B78AFE82FCBCD26B" authorityName=": Hamalainen" authorityYear="1985" box="[649,771,260,285]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="pahyapi">
<emphasis id="BAB2EAD7FFFFEC02B78AFE82FCBCD26B" box="[649,771,260,285]" italics="true" pageId="26" pageNumber="177">E. pahyapi</emphasis>
</taxonomicName>
by
<bibRefCitation id="EC574B34FFFFEC02B633FE83FB91D26A" box="[816,1070,260,285]" pageId="26" pageNumber="177" refString="Pinratana, A., Kiauta, B. &amp; Hamalainen, M. (1988) List of the odonata of Thailand and annotated bibliography. Brothers of St. Gabriel in Thailand, Bangkok, 21 pp., 20 col. pls. (unpaginated)" type="book">
Pinratana
<emphasis id="BAB2EAD7FFFFEC02B6A1FE82FC6CD26B" box="[930,979,260,285]" italics="true" pageId="26" pageNumber="177">et al</emphasis>
. (1988)
</bibRefCitation>
and in
<bibRefCitation id="EC574B34FFFFEC02B186FE83FF5BD236" author="Hamalainen" pageId="26" pageNumber="177" refString="Hamalainen, M. &amp; Pinratana, A. (1999) Atlas of the dragonflies of Thailand - Distribution maps by Provinces. Brothers of St. Gabriel in Thailand, Bangkok, 176 pp." type="book" year="1999">Hämäläinen &amp; Pinratana (1999)</bibRefCitation>
.
<taxonomicName id="4FC64D46FFFFEC02B5ECFEA0FE0BD236" authorityName=": Hamalainen" authorityYear="1985" box="[239,436,295,320]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="pahyapi">
<emphasis id="BAB2EAD7FFFFEC02B5ECFEA0FE0BD236" box="[239,436,295,320]" italics="true" pageId="26" pageNumber="177">Euphaea pahyapi</emphasis>
</taxonomicName>
can be distinguished from all Vietnamese
<taxonomicName id="4FC64D46FFFFEC02B68CFEA0FC4CD236" authorityName="Selys" authorityYear="1840" box="[911,1011,295,320]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="BAB2EAD7FFFFEC02B68CFEA0FC4CD236" box="[911,1011,295,320]" italics="true" pageId="26" pageNumber="177">Euphaea</emphasis>
</taxonomicName>
species by the unique completely and evenly amber-colored wings of the male (
<figureCitation id="10FD2A40FFFFEC02B770FECBFD69D213" box="[627,726,332,357]" captionStart="FIGURE 18" captionStartId="26.[151,250,1894,1916]" captionTargetBox="[335,1233,745,1853]" captionTargetId="figure@26.[313,1273,735,1872]" captionTargetPageId="26" captionText="FIGURE 18. Euphaea spp. [18af] vesicle of male and [18gj] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc." httpUri="https://zenodo.org/record/3096117/files/figure.png" pageId="26" pageNumber="177">Fig. 18d</figureCitation>
). Color pattern of the head is variable, entirely black as in the original description (
<bibRefCitation id="EC574B34FFFFEC02B485FEE8FDECD2FE" author="Hamalainen" box="[390,595,367,392]" pageId="26" pageNumber="177" refString="Hamalainen, M. (1985) Euphaea pahyapi sp. nov. from Peninsular Thailand (Zygoptera: Euphaeidae). Odonatologica, 14 (4), 341 - 344." type="journal article" year="1985">Hämäläinen 1985</bibRefCitation>
) or with dirty blue markings as in Asahina's (1985) description and our specimens from
<collectingCountry id="F0D17655FFFFEC02B452FE13FE0FD2DB" box="[337,432,404,429]" name="Vietnam" pageId="26" pageNumber="177">Vietnam</collectingCountry>
(Phu Quoc Island) (
<figureCitation id="10FD2A40FFFFEC02B795FE13FD47D2DB" box="[662,760,404,429]" captionStart="FIGURE 19" captionStartId="27.[151,250,1020,1042]" captionTargetBox="[168,1415,193,965]" captionTargetId="figure@27.[153,1433,193,998]" captionTargetPageId="27" captionText="FIGURE 19. Euphaea pahyapi. [19ad] ♂, Phu Quoc. (19a) head and thorax; (19b) head, antero-dorsal view; (19c) appendages, lateral view; (19d) wings." httpUri="https://zenodo.org/record/3096125/files/figure.png" pageId="26" pageNumber="177">Fig. 19b</figureCitation>
). Abdomen and anal appendages are as in
<emphasis id="BAB2EAD7FFFFEC02B1DAFE13FA23D2DA" box="[1241,1436,404,429]" italics="true" pageId="26" pageNumber="177">
<taxonomicName id="4FC64D46FFFFEC02B1DAFE13FA27D2DA" box="[1241,1432,404,429]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="masoni">Euphaea masoni</taxonomicName>
,
</emphasis>
with hairs only on S3 (
<figureCitation id="10FD2A40FFFFEC02B496FE30FE49D2A6" box="[405,502,439,464]" captionStart="FIGURE15" captionStartId="24.[151,250,929,951]" captionTargetBox="[186,1410,221,879]" captionTargetId="figure@24.[153,1433,193,907]" captionTargetPageId="24" captionText="FIGURE15.Schematic drawingsof abdominal hair tufts inlateral view ofmale Euphaeaspp.(15a) E.saola sp.nov.; (15b)E. guerini; (15c) E.hirta; (15d) E.masoni and E. pahyapi. IX show abdominal segments S110." httpUri="https://zenodo.org/record/3096107/files/figure.png" pageId="26" pageNumber="177">Fig. 15d</figureCitation>
). Vesicle of
<taxonomicName id="4FC64D46FFFFEC02B783FE30FCFAD2A6" authorityName=": Hamalainen" authorityYear="1985" box="[640,837,439,464]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="pahyapi">
<emphasis id="BAB2EAD7FFFFEC02B783FE30FCFAD2A6" box="[640,837,439,464]" italics="true" pageId="26" pageNumber="177">Euphaea pahyapi</emphasis>
</taxonomicName>
differs from that of
<taxonomicName id="4FC64D46FFFFEC02B124FE3EFB28D2A6" box="[1063,1175,441,464]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="masoni">
<emphasis id="BAB2EAD7FFFFEC02B124FE3EFB28D2A6" box="[1063,1175,441,464]" italics="true" pageId="26" pageNumber="177">E. masoni</emphasis>
</taxonomicName>
by being heart-shaped, pointed at its posterior margin (
<figureCitation id="10FD2A40FFFFEC02B4FCFE5BFDE3D283" box="[511,604,476,501]" captionStart="FIGURE 18" captionStartId="26.[151,250,1894,1916]" captionTargetBox="[335,1233,745,1853]" captionTargetId="figure@26.[313,1273,735,1872]" captionTargetPageId="26" captionText="FIGURE 18. Euphaea spp. [18af] vesicle of male and [18gj] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc." httpUri="https://zenodo.org/record/3096117/files/figure.png" pageId="26" pageNumber="177">Fig. 18f</figureCitation>
) whereas it is ax-shaped with the posterior margin rounded in
<taxonomicName id="4FC64D46FFFFEC02B029FE5AFA23D282" box="[1322,1436,477,500]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="masoni">
<emphasis id="BAB2EAD7FFFFEC02B029FE5AFA23D282" box="[1322,1436,477,500]" italics="true" pageId="26" pageNumber="177">E. masoni</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFFEC02B59CFE78FF40D16E" box="[159,255,511,536]" captionStart="FIGURE 18" captionStartId="26.[151,250,1894,1916]" captionTargetBox="[335,1233,745,1853]" captionTargetId="figure@26.[313,1273,735,1872]" captionTargetPageId="26" captionText="FIGURE 18. Euphaea spp. [18af] vesicle of male and [18gj] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc." httpUri="https://zenodo.org/record/3096117/files/figure.png" pageId="26" pageNumber="177">Fig. 18e</figureCitation>
). Female of this species is still undescribed.
</paragraph>
<paragraph id="887936C5FFFFEC02B5C4FDA3FADFD116" blockId="26.[151,1436,151,680]" pageId="26" pageNumber="177">
The record of
<taxonomicName id="4FC64D46FFFFEC02B475FDA3FDF1D14B" authorityName=": Bui" authorityYear="2007" box="[374,590,548,573]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="ochracea">
<emphasis id="BAB2EAD7FFFFEC02B475FDA3FDF1D14B" box="[374,590,548,573]" italics="true" pageId="26" pageNumber="177">Euphaea ochracea</emphasis>
</taxonomicName>
from Kbal Chhay Cascades of Preah Sihanouk Province,
<collectingCountry id="F0D17655FFFFEC02B004FDA3FAC5D14B" box="[1287,1402,548,573]" name="Cambodia" pageId="26" pageNumber="177">Cambodia</collectingCountry>
in
<bibRefCitation id="EC574B34FFFFEC02B594FDC0FE37D116" box="[151,392,583,608]" pageId="26" pageNumber="177" refString="Kosterin, O. E., Chartier, G., Holden, J. &amp; Mey, F. S. (2012) New records of Odonata from Cambodia, based mostly on photographs. Cambodian Journal of Natural History, 2012 (2), 150 - 163." type="journal article">
Kosterin
<emphasis id="BAB2EAD7FFFFEC02B5FDFDCEFE89D116" box="[254,310,583,608]" italics="true" pageId="26" pageNumber="177">et al.</emphasis>
(2012)
</bibRefCitation>
is a misidentification and concerns in fact
<taxonomicName id="4FC64D46FFFFEC02B66BFDCEFC5ED116" authorityName=": Hamalainen" authorityYear="1985" box="[872,993,583,608]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="pahyapi">
<emphasis id="BAB2EAD7FFFFEC02B66BFDCEFC5ED116" box="[872,993,583,608]" italics="true" pageId="26" pageNumber="177">E. pahyapi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EC574B34FFFFEC02B6F3FDC0FAE8D116" author="Kosterin" box="[1008,1367,583,608]" pageId="26" pageNumber="177" refString="Kosterin, O. E. (2016) A survey of Odonata of Mondulkiri, the elevated eastern province of Cambodia, for ten days in June 2016. International Dragonfly Fund Report. Journal of the International Dragonfly Fund, 98, 1 - 85." type="journal article" year="2016">
Kosterin 2016, and
<emphasis id="BAB2EAD7FFFFEC02B1CCFDCEFAE8D116" box="[1231,1367,585,608]" italics="true" pageId="26" pageNumber="177">pers. comm.</emphasis>
</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C0DC654EFFFFEC02B5C4FDEBFB05D1DE" pageId="26" pageNumber="177" type="distribution">
<paragraph id="887936C5FFFFEC02B5C4FDEBFB05D1DE" blockId="26.[151,1436,151,680]" pageId="26" pageNumber="177">
<emphasis id="BAB2EAD7FFFFEC02B5C4FDEBFEDFD1F3" bold="true" box="[199,352,620,645]" pageId="26" pageNumber="177">Distribution.</emphasis>
<collectingCountry id="F0D17655FFFFEC02B46EFDEBFE73D1F3" box="[365,460,620,645]" name="Vietnam" pageId="26" pageNumber="177">Vietnam</collectingCountry>
(Phu Quoc Island),
<collectingCountry id="F0D17655FFFFEC02B7CBFDEBFC93D1F3" box="[712,812,620,645]" name="Thailand" pageId="26" pageNumber="177">Thailand</collectingCountry>
(
<collectingRegion id="4A02F827FFFFEC02B643FDEBFC23D1F2" box="[832,924,620,644]" country="Thailand" name="Ranong" pageId="26" pageNumber="177">Ranong</collectingRegion>
,
<collectingRegion id="4A02F827FFFFEC02B6AEFDEBFC4FD1F3" box="[941,1008,620,645]" country="Thailand" name="Krabi" pageId="26" pageNumber="177">Krabi</collectingRegion>
, and
<collectingRegion id="4A02F827FFFFEC02B134FDEAFBC6D1F3" box="[1079,1145,621,645]" country="Thailand" name="Trang" pageId="26" pageNumber="177">Trang</collectingRegion>
Provinces) (
<bibRefCitation id="EC574B34FFFFEC02B010FDEBFF6CD1DE" author="Hamalainen" pageId="26" pageNumber="177" refString="Hamalainen, M. (2017) The Caloptera damselflies of Thailand - Distribution maps by provinces (Odonata: Calopterygoidea). Faunistic studies in South-East Asian and Pacific Island Odonata, 19, 1 - 28." type="journal article" year="2017">Hämäläinen 2017</bibRefCitation>
),
<collectingCountry id="F0D17655FFFFEC02B5E5FD08FEE6D1DE" box="[230,345,655,680]" name="Cambodia" pageId="26" pageNumber="177">Cambodia</collectingCountry>
(Preah Sihanouk Province) (
<bibRefCitation id="EC574B34FFFFEC02B79FFD08FCC0D1DE" author="Kosterin" box="[668,895,655,680]" pageId="26" pageNumber="177" refString="Kosterin, O. E., Chartier, G., Holden, J. &amp; Mey, F. S. (2012) New records of Odonata from Cambodia, based mostly on photographs. Cambodian Journal of Natural History, 2012 (2), 150 - 163." type="journal article" year="2012">
Kosterin
<emphasis id="BAB2EAD7FFFFEC02B606FD16FC82D1DE" box="[773,829,655,680]" italics="true" pageId="26" pageNumber="177">et al.</emphasis>
2012
</bibRefCitation>
, and Kosterin
<emphasis id="BAB2EAD7FFFFEC02B122FD16FB15D1DE" box="[1057,1194,657,680]" italics="true" pageId="26" pageNumber="177">pers. comm.</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>