162 lines
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162 lines
17 KiB
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@prefix dc: <http://purl.org/dc/elements/1.1/> .
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@prefix xlink: <http://www.w3.org/1999/xlink/> .
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# Warning: Failed to output a material citation, could not create identifier
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# Warning: Failed to output a material citation, could not create identifier
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# Warning: Failed to output a material citation, could not create identifier
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<http://treatment.plazi.org/id/B551AE7B841AF701FE9A3713FEE4CD1D>
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cito:cites <http://dx.doi.org/10.5281/zenodo.2640607>, <http://dx.doi.org/10.5281/zenodo.2640605>, <http://dx.doi.org/10.5281/zenodo.2640609>, <http://dx.doi.org/10.5281/zenodo.2640611>, <http://dx.doi.org/10.5281/zenodo.2640614>, <http://dx.doi.org/10.5281/zenodo.2640626>, <http://dx.doi.org/10.5281/zenodo.2640628>, <http://dx.doi.org/10.5281/zenodo.2640630>, <http://dx.doi.org/10.5281/zenodo.2640632>, <http://dx.doi.org/10.5281/zenodo.2640634> ;
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dc:creator "Détroit, Florent; Mijares, Armand Salvador; Corny, Julien; Daver, Guillaume; Zanolli, Clément; Dizon, Eusebio; Robles, Emil; Grün, Rainer; Piper, Philip J." ;
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dc:title "Homo luzonensis Détroit & Mijares & Corny & Daver & Zanolli & Dizon & Robles & Grün & Piper 2019, sp. nov." ;
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trt:definesTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Homo_luzonensis_D%C3%A9troit_2019> ;
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trt:publishedIn <http://doi.org/10.5281/zenodo.5931871> ;
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a trt:Treatment .
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<http://doi.org/10.5281/zenodo.5931871>
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bibo:endPage "186" ;
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bibo:journal "Nature" ;
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bibo:pubDate "2019-04-10" ;
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bibo:startPage "181" ;
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bibo:volume "568" ;
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dc:creator "Détroit, Florent; Mijares, Armand Salvador; Corny, Julien; Daver, Guillaume; Zanolli, Clément; Dizon, Eusebio; Robles, Emil; Grün, Rainer; Piper, Philip J." ;
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dc:date "2019" ;
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dc:title "A new species of Homo from the Late Pleistocene of the Philippines" ;
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fabio:hasPart <http://dx.doi.org/10.5281/zenodo.2640607>, <http://dx.doi.org/10.5281/zenodo.2640605>, <http://dx.doi.org/10.5281/zenodo.2640609>, <http://dx.doi.org/10.5281/zenodo.2640611>, <http://dx.doi.org/10.5281/zenodo.2640614>, <http://dx.doi.org/10.5281/zenodo.2640626>, <http://dx.doi.org/10.5281/zenodo.2640628>, <http://dx.doi.org/10.5281/zenodo.2640630>, <http://dx.doi.org/10.5281/zenodo.2640632>, <http://dx.doi.org/10.5281/zenodo.2640634> ;
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a fabio:JournalArticle .
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<http://taxon-concept.plazi.org/id/Animalia/Homo_luzonensis_D%C3%A9troit_2019>
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dwc:authority "Détroit & Mijares & Corny & Daver & Zanolli & Dizon & Robles & Grün & Piper, 2019" ;
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dwc:authorityName "Détroit & Mijares & Corny & Daver & Zanolli & Dizon & Robles & Grün & Piper" ;
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dwc:authorityYear "2019" ;
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dwc:class "Mammalia" ;
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dwc:family "Hominidae" ;
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dwc:genus "Homo" ;
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dwc:kingdom "Animalia" ;
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dwc:order "Primates" ;
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dwc:phylum "Chordata" ;
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dwc:rank "species" ;
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dwc:scientificNameAuthorship "Détroit & Mijares & Corny & Daver & Zanolli & Dizon & Robles & Grün & Piper, 2019" ;
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dwc:species "luzonensis" ;
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trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Homo_luzonensis> ;
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a dwcFP:TaxonConcept .
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<http://taxon-name.plazi.org/id/Animalia/Homo_luzonensis>
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dwc:class "Mammalia" ;
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dwc:family "Hominidae" ;
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dwc:genus "Homo" ;
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dwc:kingdom "Animalia" ;
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dwc:order "Primates" ;
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dwc:phylum "Chordata" ;
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dwc:rank "species" ;
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dwc:species "luzonensis" ;
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dwc:class "Mammalia" ;
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dwc:family "Hominidae" ;
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dwc:genus "Homo" ;
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dwc:kingdom "Animalia" ;
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dwc:order "Primates" ;
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dwc:class "Mammalia" ;
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dwc:family "Hominidae" ;
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dwc:order "Primates" ;
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dwc:class "Mammalia" ;
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dwc:kingdom "Animalia" ;
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dwc:order "Primates" ;
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dwc:class "Mammalia" ;
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dwc:kingdom "Animalia" ;
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dwc:phylum "Chordata" ;
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a dwcFP:TaxonName .
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trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
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a dwcFP:TaxonName .
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<http://taxon-name.plazi.org/id/Animalia>
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dwc:kingdom "Animalia" ;
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a dwcFP:TaxonName .
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<http://dx.doi.org/10.5281/zenodo.2640607>
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dc:description "Fig. 2 | Fossil remains of H. luzonensis from Late Pleistocene sediments at Callao Cave. a, Holotype CCH6: postcanine maxillary teeth in occlusal (left) and buccal (right) aspects, with three-dimensional rendering of enamel (dark blue), dentine and cement (light brown), and pulp cavity (dark grey) for CCH6-b–CCH6-e. b, Intermediate manual phalanx CCH2 (dorsal, lateral and palmar aspects). c, Distal manual phalanx CCH5 (dorsal, lateral/medial and palmar aspects). d, Proximal pedal phalanx CCH4 (dorsal, lateral and plantar aspects). e, Intermediate pedal phalanx CCH3 (dorsal, medial and plantar aspects). f, Left P 3 or P 4 CCH8: occlusal (top) and buccal (bottom) aspects, with three-dimensional rendering of enamel, dentine and cement, and pulp cavity. g, Right M3 CCH9: occlusal (top) and buccal (bottom) aspects. h, Juvenile femoral shaft CCH7 (anterior, lateral and posterior aspects). Scale bars, 10 mm (a–g) and 20 mm (h); additional views are shown in Extended Data Figs. 1, 5, 7–10." ;
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fabio:hasRepresentation <https://zenodo.org/record/2640607/files/figure.svg> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.2640605>
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dc:description "Fig. 1 | Geographical location of Callao Cave. Map showing the location of Callao Cave on Luzon Island (the Philippines), emerged lands at 50 and 120 m below present sea level (adapted from ref. 46, H. K. Voris, Field Museum of Natural History) and the major biogeographical boundaries recognized in the area. A, Wallace’s Line modified by Huxley; B, Wallace’s Line; C, Lydekker’s Line. Luzon Island lies in between the original Wallace’s Line and the Wallace’s Line modified by Huxley and was never connected to mainland Asia during the Quaternary." ;
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fabio:hasRepresentation <https://zenodo.org/record/2640605/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.2640609>
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dc:description "Fig. 3 | Dental metrics. a, Square root of the computed occlusal surface area of P3 (n = 166), P4 (n = 167), M1 (n = 216), M2 (n = 185) and M3 (n = 129). Dots correspond to the group average, vertical bars and shaded areas correspond to ±2 s.e.m. √OS, square root of the occlusal surface. b, Bivariate scatter plot of the summed square root of computed occlusal surface areas of premolars (P3 and P4) versus molars (M1 and M2), with regression lines (solid lines) and 95% confidence intervals (shaded areas) for all groups, except H. floresiensis and H. luzonensis. Sample sizes for a, b, respectively: Australopithecus, n = 119, 6; Paranthropus, n = 111, 5; African and European early Homo, n = 114, 13; Asian early Homo, n = 74, 5; H. neanderthalensis, n = 126, 12; H. sapiens, n = 307,47; H. floresiensis, n = 4, 1; H. luzonensis, n = 8∗, 1 (∗CCH8 treated as P3 and P4 in a). A detailed list of specimens can be found in Supplementary Table 4." ;
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fabio:hasRepresentation <https://zenodo.org/record/2640609/files/figure.svg> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.2640611>
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dc:description "Fig. 4 | Premolar EDJ of H. luzonensis. a, EDJ of the P 3 of H. luzonensis (CCH6-e) compared to the EDJs of H. floresiensis (Liang Bua 1 (LB1)), H. sapiens, H. erectus (Sangiran 4) and H. neanderthalensis (KRD 53). Horns of dentine were reconstructed for CCH6-e, LB1 and Sangiran 4; see Methods. Scale bar, 5 mm. b, c, Between-group principal component analyses (bgPCAs) of the three-dimensional landmark Procrustes- registered shape coordinates of the P 3 s (b) and P 4 s (c). Sample sizes for b, c, respectively: H. erectus, n = 2, 3; H. neanderthalensis, n = 5, 6; fossil H. sapiens, n = 3, 3; extant H. sapiens, n = 8, 9; H. floresiensis, n = 1, 0; H. luzonensis, n = 2, 2. A detailed list of specimens can be found in Supplementary Table 6" ;
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fabio:hasRepresentation <https://zenodo.org/record/2640611/files/figure.svg> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.2640614>
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dc:description "Fig. 5 | Proximal pedal phalanx of H. luzonensis (CCH4). CCH4 compared with specimens attributed to Australopithecus (A. afarensis, n = 6; A. africanus, n = 1), recent H. sapiens (n = 64) and H. floresiensis (n = 2). a, bgPCA of Procrustes-registered landmarks and semilandmarks: scatter plot of individual scores for bgPC1 versus bgPC2. b, Shape variation associated with bgPC1 and bgPC2: CCH4 and Australopithecus phalanges are elongated and curved (see bgPC1 max.). A detailed list of specimens can be found in Supplementary Table 7." ;
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fabio:hasRepresentation <https://zenodo.org/record/2640614/files/figure.svg> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.2640626>
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dc:description "Extended Data Fig. 6 | Procrustes analyses of the intermediate manual phalanx of H. luzonensis. CCH2 compared to specimens attributed to Australopithecus (A. afarensis, n = 1; A. africanus, n = 1; A. sediba, n = 2), Paranthropus/early Homo (Swartkrans, Member 1, n = 3; Member 3, n = 2), H. naledi (Hand 1, n = 2), H. floresiensis (LB1/48 and LB6/9) and recent H. sapiens separated into 3 samples corresponding to ray number (n = 15, 21 and 19 for rays 2, 3 and 4, respectively). A detailed list of specimens can be found in Supplementary Table 9. a–c, bgPCA of Procrustes-registered landmarks and semilandmarks (H. sapiens sample includes ray 3 only): scatter plot of individual scores for bgPC1 versus bgPC2 (a); scatter plot of individual scores for bgPC2 versus bgPC3 (b); shape variation associated with bgPC1, bgPC2 and bgPC3 (c). d, bgPCA of Procrustes-registered landmarks and semilandmarks (H. sapiens sample includes ray 4 only): scatter plot of individual scores for bgPC1 versus bgPC2. e, bgPCA of Procrustes-registered landmarks and semilandmarks (H. sapiens sample includes ray 2 only): scatter plot of individual scores for bgPC1 versus bgPC2. f, Box-and-whisker plot depicting centroid size (Procrustes registration of all specimens together). Box, 25–75th percentiles; centre line, median; whiskers, non-outlier range; dots, outliers; dotted line, value for CCH2. g, Protocol for three-dimensional Procrustes analysis: landmarks placed on the main anatomical features (n = 23, in red) and equally spaced semilandmarks (n = 250, in green) placed on the whole surface of the intermediate manual phalanx (template shown on the intermediate manual phalanx, ray 3, of the recent H. sapiens 17980)." ;
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fabio:hasRepresentation <https://zenodo.org/record/2640626/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.2640628>
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dc:description "Extended Data Fig. 7 | CCH5, distal manual phalanx of H. luzonensis. a, Photograph of the original specimen CCH5 in palmar view. b, Three- dimensional rendering of CCH5. From left to right: palmar, lateral/medial, dorsal, medial/lateral, distal (top), proximal (bottom), disto-lateral/ medial and proximo-lateral/medial aspects. c, Comparison of CCH5 with Pliocene (A.L.333-11 and A.L.333-50), Lower Pleistocene (OH7 (FLK- NN-B) and SKX 27504), Upper Pleistocene (LB6/12) and recent distal manual phalanges (PAPO-74-53 and PAPO-74-11) in palmar (top) and side (bottom) views. All specimens are from rays 2–5 of unknown side, except for OH7 (second to fourth ray, probably from the right hand of a juvenile individual). d, e, Box-and-whisker plots depicting the expansion index ((apical tuft maximum medio-lateral width/maximum mediolateral width of the base) × 100)) (d) and the robusticity index ((apical tuft maximum medio-lateral width/biomechanical length) × 100) (e) of the distal manual phalanx CCH5. Box, 25–75th percentiles, centre line, median; whiskers, non-outlier range; dots, outliers; dotted line, value for CCH5. A detailed list of specimens can be found in Supplementary Table 10. For conservative reasons, taxonomic assignation of hand remains OH7 and SKX 27504 are considered to be uncertain (see main text and Methods). Scale bars, 10 mm." ;
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fabio:hasRepresentation <https://zenodo.org/record/2640628/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.2640630>
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dc:description "Extended Data Fig. 8 | CCH4, proximal pedal phalanx of H. luzonensis. a, Photograph of the original specimen CCH4 in lateral view. b, Threedimensional rendering of CCH4. From left to right: plantar, lateral, dorsal, medial, distal (top), proximal (bottom), disto-lateral and proximo-lateral aspects. c, Comparison of CCH4 with Pliocene (A.L.333-115h, left third proximal phalanx, mirrored), Pleistocene (LB1/36, unknown side and rays 2–5) and recent (‘Negrito’ 9764, right third proximal pedal phalanx), proximal pedal phalanges in plantar (top) and side (bottom) views. d, Transverse (1, 2, 3) and mid-sagittal (4) micro-CT slices of CCH4 (plantar aspect of three-dimensional rendering, during the segmentation process; 1, 2, 3, dorsal is up, plantar is down, lateral is left, medial is right; 4, distal is up, proximal is down, dorsal is left, plantar is right). e, Protocol for three-dimensional Procrustes analysis: landmarks placed on the main anatomical features (n = 20, in red) and equally spaced semilandmarks (n = 250, in green) placed on the whole surface of the proximal pedal phalanx (template shown on the proximal pedal phalanx, ray 2 of the right foot of the recent H. sapiens 35071). f, g, Box-and-whisker plots depicting comparisons of the articular angle a (f) and the dorsal canting angle α (g) of the proximal pedal phalanx CCH4. Box, 25–75th percentiles; centre line, median; whiskers, non-outlier range; dots, outliers; dotted line, value for CCH4. A detailed list of specimens can be found in Supplementary Table 11. Scale bars, 10 mm." ;
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fabio:hasRepresentation <https://zenodo.org/record/2640630/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.2640632>
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dc:description "Extended Data Fig. 9 | CCH3, intermediate pedal phalanx of H. luzonensis. a, Photograph of the original specimen CCH3 in plantar view. b, Three-dimensional rendering of CCH3. From left to right: plantar, medial, dorsal, lateral, distal (top), proximal (bottom), disto-medial and proximo-medial aspects. c, Comparison of CCH3 with Pliocene (A.L.333- 21a, unknown side and rays 2–5, and A.L.333-115k, fourth intermediate phalanx), Upper Pleistocene (LB1/56, LB1-15 and LB1/39: unknown side and rays 2–5) and recent (PAPO-74-150) intermediate pedal phalanges in plantar (top) and side (bottom) views. Note the variation in shape and size both between taxa (for example, H. sapiens and H. floresiensis) and in the same individual (for example, LB1 and PAPO-74-150). A detailed list of specimens can be found in Supplementary Table 12. Scale bars, 10 mm." ;
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fabio:hasRepresentation <https://zenodo.org/record/2640632/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.2640634>
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dc:description "Extended Data Fig. 10 | CCH7, femoral shaft of a juvenile individual of H. luzonensis. a, Photograph of the original specimen CCH7 (posterior aspect). b, Three-dimensional rendering of CCH7. From left to right: anterior, medial, posterior and lateral aspects. Scale bar, 20 mm. c, Transverse micro-CT slices of CCH7 at proximal diaphysis (top), midshaft (middle) and distal diaphysis (bottom), and posterior aspect of the three-dimensional rendering of the femoral shaft, during the segmentation process (orientation of slices: anterior is up, posterior is down, lateral is left, medial is right)." ;
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fabio:hasRepresentation <https://zenodo.org/record/2640634/files/figure.png> ;
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a fabio:Figure .
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