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dc:creator "Moreno-González, Jairo A.; O, Ranulfo González; D, Eduardo Flórez" ;
dc:title "Tityus columbianus" ;
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bibo:endPage "94" ;
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bibo:pubDate "2019-08-26" ;
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bibo:volume "4660" ;
dc:creator "Moreno-González, Jairo A.; O, Ranulfo González; D, Eduardo Flórez" ;
dc:date "2019" ;
dc:title "Taxonomic revision of the Colombian Tityus (Archaeotityus) (Scorpiones, Buthidae) species: a morphological and morphometric approach, with a description of a new species" ;
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dc:description "FIGURE 5. Tityus (Archaeotityus) Lourenço, 2006, carapace coloration, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. B. Male. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). C. Female. D. Male. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Female paratype. F. Male holotype." ;
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dc:description "FIGURE 6. Tityus (Archaeotityus) Lourenço, 2006, metasoma coloration pattern, dorsal (left) and ventral (right). A, D. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. D. Male. B, E. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). B. Female. E. Male. C, F. Tityus guane sp. nov. (ICN-As-1003). C. Female paratype. F. Male holotype." ;
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dc:description "FIGURE 7. Tityus (Archaeotityus) Lourenço, 2006, chelicerae, dorsal aspect, showing reticulation pattern. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. B. Male. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). C. Female. D. Male. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Female paratype. F. Male holotype." ;
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<http://dx.doi.org/10.5281/zenodo.3377151>
dc:description "FIGURE 8. Tityus (Archaeotityus) Lourenço, 2006, carapace sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984. A. Female (ICN-As-740). B. Male (ICN-As-741). C, D. Tityus columbianus (Thorell, 1876). C. Female (ICN-As-739). D. Male (ICN-As-524). E, F. Tityus guane sp. nov. (ICN-As-1003). E. Female paratype. F. Male holotype. Scale bars= 1.5 mm." ;
fabio:hasRepresentation <https://zenodo.org/record/3377151/files/figure.png> ;
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<http://dx.doi.org/10.5281/zenodo.3377153>
dc:description "FIGURE 9. Tityus (Archaeotityus) Lourenço, 2006, female pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN- As-1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." ;
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dc:description "FIGURE 10. Tityus (Archaeotityus) Lourenço, 2006, male pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-741). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN-As- 1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." ;
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dc:description "FIGURE 11. Tityus (Archaeotityus) Lourenço, 2006, right female pedipalp sculpturing. AC. Tityus bastosi Lourenço, 1984 (ICN-As-740). A. External aspect. B. Dorsal aspect. C. Ventral aspect. DF. Tityus columbianus (Thorell, 1876) (ICN-As-739). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." ;
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<http://dx.doi.org/10.5281/zenodo.3377177>
dc:description "FIGURE 21. Tityus (Archaeotityus) columbianus (Thorell, 1876), habitus. A, B. Female (ICN-As-739). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Villa de Leyva, Boyacá) (ICN-As-739). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." ;
fabio:hasRepresentation <https://zenodo.org/record/3377177/files/figure.png> ;
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dc:description "FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments IV sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." ;
fabio:hasRepresentation <https://zenodo.org/record/3377165/files/figure.png> ;
a fabio:Figure .
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dc:description "FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments IV sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." ;
fabio:hasRepresentation <https://zenodo.org/record/3377167/files/figure.png> ;
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dc:description "FIGURE 17. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segment V sculpturing, showing macrosetae, ventral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Abbreviations: ML, median lateral; VL, ventrolateral; VSM, ventrosubmedian. Scale bars= 1.0 mm." ;
fabio:hasRepresentation <https://zenodo.org/record/3377169/files/figure.png> ;
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dc:description "FIGURE 18. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segment V sculpturing, showing macrosetae, ventral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As- 181). Abbreviations: ML, median lateral; VL, ventrolateral; VSM, ventrosubmedian. Scale bars= 1.5 mm." ;
fabio:hasRepresentation <https://zenodo.org/record/3377171/files/figure.png> ;
a fabio:Figure .
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dc:description "FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." ;
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<http://dx.doi.org/10.5281/zenodo.3377175>
dc:description "FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." ;
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<http://dx.doi.org/10.5281/zenodo.3377179>
dc:description "FIGURE 22. Tityus (Archaeotityus) columbianus (Thorell, 1876), male pedipalp chela. AC. Male (Villa de Leyva, Boyacá) (ICN-As-524). A. External aspect. B. Dorsal aspect. C. Ventral aspect. DF. Male (Villa de Leyva, Boyacá) (ICN-As-739). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." ;
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<http://dx.doi.org/10.5281/zenodo.3377199>
dc:description "FIGURE 31. Tityus (Archaeotityus) parvulus Kraepelin, 1914, habitus. A, B. Female (ICN-As-1004). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Chinchiná, Caldas) (ICN-As-1004). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Chinchiná, Caldas) (ICN-As-1004). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." ;
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dc:description "FIGURE 33. Tityus (Archaeotityus) Lourenço, 2006, metasoma coloration pattern, dorsal (left) and ventral (right). A, C. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). A. Female. C. Male. B, D. Tityus tayrona Lourenço, 1991 (ICN-As-181). B. Female. D. Male." ;
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<http://dx.doi.org/10.5281/zenodo.3377215>
dc:description "FIGURE 39. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments IV sculpturing, lateral aspect. A. Tityus parvulus Kraepelin, 1914 (ICN-As-104). B. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bar= 3.0 mm." ;
fabio:hasRepresentation <https://zenodo.org/record/3377215/files/figure.png> ;
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<http://dx.doi.org/10.5281/zenodo.3377217>
dc:description "FIGURE 40. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments IV sculpturing, lateral aspect. A, B. Tityus parvulus Kraepelin, 1914. A. Male (Chinchiná, Caldas) (ICN-As-1004). B. Male (Chinchiná, Caldas) (ICN-As-1004). C, D. Tityus tayrona Lourenço, 1991. C. Male (Zambrano, Bolívar) (ICN-As-181). D. Male (Zambrano, Bolívar) (ICN-As-181). Scale bars= 3.0 mm." ;
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dc:description "FIGURE 46. Tityus (Archaeotityus) Lourenço, 2006, habitus in life. A. Tityus bastosi Lourenço, 1984, female from El Carmen, Villavicencio, Meta, Colombia. B. Tityus columbianus (Thorell, 1876), female from Soacha, Cundinamarca, Colombia. C. Tityus guane sp. nov., male from Vereda Pasochico, Finca La Ceiba, Los Santos, Santander. D, E. Tityus parvulus Kraepelin, 1914, females. D. Female from Regional Natural Reserve El Vínculo [Natural Regional Park El Vínculo], Buga, Valle del Cauca, Colombia. E. Female from the Botanical Garden Maria Cespedes [Jardín Botánico Maria Cespedes], Tulúa, Valle del Cauca, Colombia. F. Tityus tayrona Lourenço, 1991, female from National Natural Park Tayrona [Parque Nacional Natural Tayrona], Santa Marta, Magdalena, Colombia. Photo B, courtesy of N. Herreño." ;
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dc:description "FIGURE 3. Tityus (Archaeotityus) Lourenço, 2006, right pedipalp, showing trichobothrial pattern. AC. Pedipalp chela. A. External aspect. B. Dorsal aspect. C. Ventral aspect. D. Femur, dorsal aspect. E. Patella, dorsal aspect. Abbreviations: d, dorsal; db, dorsal basal; dt, dorsal terminal; e, external; eb, Eb, external basal; esb, Esb, external subbasal; est, Est, external subterminal; et, Et, external terminal; i, internal; it, internal terminal; v, ventral. Scale bars= AC= 2.0 mm; D, E= 1.0 mm." ;
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dc:description "FIGURE 2. Tityus (Archaeotityus) guane sp. nov., sternum and pectines, showing the shape of the basal pectinal piece. A. Female paratype (ICN-As-1003). B. Male holotype (ICN-As-1003). Abbreviations: A, anterior region; P, posterior region." ;
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dc:description "FIGURE 14. Tityus (Archaeotityus) Lourenço, 2006, left pectine with peg sensillae details, ventral aspect. AD. Tityus bastosi Lourenço, 1984.A, B. Female (MZSP 71740). A. Pectine. B. Peg sensillae shape. C, D. Male (MZSP 71740). C. Pectine. D. Peg sensillae shape. E, F. Tityus columbianus (Thorell, 1876) (ICN-As-739). E. Female pectine. F. Female peg sensillae shape. GJ. Tityus guane sp. nov.. G, H. Female (MZSP 71736). G. Pectine. H. Peg sensillae shape. I, J. Male (MZSP 71736). I. Pectine. J. Peg sensillae shape. Scale bars= A, C, E, G, I= 400 µm; B, D, F, H, J= 10 µm." ;
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dc:description "FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." ;
fabio:hasRepresentation <https://zenodo.org/record/3377181/files/figure.png> ;
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<http://dx.doi.org/10.5281/zenodo.3377136>
dc:description "FIGURE 1. Tityus (Archaeotityus) Lourenço, 2006, metasomal segments IV showing distribution of macrosetae.A. Segments IV, ventral aspect. B. Segment V, lateral aspect. Abbreviations: ML, median lateral; VL, ventrolateral; VSM, ventrosubmedian." ;
fabio:hasRepresentation <https://zenodo.org/record/3377136/files/figure.png> ;
a fabio:Figure .
<http://dx.doi.org/10.5281/zenodo.3377195>
dc:description "FIGURE 30. Distribution of Tityus (Archaeotityus) columbianus (Thorell, 1876) and Tityus (Archaeotityus) guane sp. nov., showing known records and new locality records." ;
fabio:hasRepresentation <https://zenodo.org/record/3377195/files/figure.png> ;
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<http://dx.doi.org/10.5281/zenodo.3377231>
dc:description "FIGURE 47. Tityus (Archaeotityus) Lourenço, 2006, habitat. A. Vereda El Carmen, Villavicencio, Meta, Colombia, habitat of Tityus bastosi Lourenço, 1984. B, C. Soacha, Cundinamarca, Colombia, habitat of Tityus columbianus (Thorell, 1876). D. Botanical Garden Maria Cespedes [Jardín Botánico Maria Cespedes], Tulúa, Valle, Colombia, habitat of Tityus parvulus Kraepelin, 1914. E. Vereda Pasochico, Finca La Ceiba, Los Santos, Santander, habitat of Tityus guane sp. nov. F. Natural National Park Tayrona [Parque Natural Nacional Tayrona], Santa Marta, Magdalena, Colombia, habitat of Tityus tayrona Lourenço, 1991. Photos B and C, courtesy of N. Herreño." ;
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dwc:locality "America merid., Columbia" ;
dwc:typeStatus "holotype" ;
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dwc:catalogNumber "ICN-As- 188" ;
dwc:eventDate "1996-07" ;
dwc:locality "Santuario de Flora y Fauna Los Colorados" ;
dwc:recordedBy "F. Escobar & W. R. Lourenco" ;
dwc:verbatimElevation "390" ;
dwc:verbatimLatitude "9.956667" ;
dwc:verbatimLongitude "-75.08667" ;
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dwc:catalogNumber "ICN-As-364" ;
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dwc:locality "Vereda Montesuarez" ;
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dwc:verbatimLatitude "5.75" ;
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dwc:catalogNumber "ICN-As-130" ;
dwc:eventDate "1991-04" ;
dwc:locality "Vereda Chipa" ;
dwc:recordedBy "L. G. Baptiste" ;
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dwc:verbatimLatitude "6.1894445" ;
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dwc:catalogNumber "ICN-As-288" ;
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dwc:catalogNumber "ICN-As-388" ;
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dwc:catalogNumber "ICN-As-739" ;
dwc:eventDate "2002-08-25" ;
dwc:locality "Villa de Leyva" ;
dwc:recordedBy "E. Gonzalez" ;
dwc:verbatimElevation "2128" ;
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dwc:verbatimLongitude "-73.53333" ;
trt:gbifOccurrenceId "2365686013" ;
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dwc:catalogNumber "ICN- As-524" ;
dwc:collectionCode "ICN" ;
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dwc:verbatimLatitude "5.6833334" ;
dwc:verbatimLongitude "-73.48333" ;
trt:gbifOccurrenceId "2365685927" ;
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dwc:catalogNumber "ICN-As-352" ;
dwc:eventDate "1985-10" ;
dwc:locality "El Roble" ;
dwc:recordedBy "R. Gonzalez" ;
dwc:verbatimElevation "2200" ;
dwc:verbatimLatitude "5.638611" ;
dwc:verbatimLongitude "-73.528336" ;
trt:gbifOccurrenceId "2365686372" ;
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dwc:catalogNumber "ICN-As-314" ;
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dwc:locality "near river Cebada and river Cene" ;
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dwc:catalogNumber "ICN-As-851" ;
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dwc:verbatimLatitude "5.566703" ;
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