Novak, Tone; Novak, Ljuba Slana; Kozel, Peter; Schaider, Miriam Gudrun; Komposch, Christian; Lipovšek, Saška; Podlesnik, Jan; Paušič, Igor; Raspotnig, Günther
Nemastoma bidentatum subsp. gruberi Novak, Slana Novak, Kozel & Raspotnig 2021, ssp. nov.
Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence
Novak, Slana Novak, Kozel & Raspotnig, 2021
harvestmen
Fig. 1. Representatives of the Nemastoma bidentatum Roewer, 1914 complex. A. The habitus of a male Nemastoma bidentatum gruberi Novak, Slana Novak, Kozel & Raspotnig ssp. nov.; Svete gore, Slovenia (photo: C. Komposch); inset: femur IV with two pseudo-articles (apparent articles, i.e., sections delimited with pale, indented rings on the proximal femur portion), and a saw on the latero-posterior margin. B. The body of a male Nemastoma relictum Gruber & Martens, 1968 stat. nov. showing the tuberculate surface of glittering scutum compositum, and position of the ocular mound (tuber oculorum), supracheliceral lamellae, chelicerae with cheliceral apophysis, and pedipalps and leg. The frontal portion of the scutum is smeared over with a hardened secretum of the integumental glands (cf. Wolff et al. 2016); Mt. Grosses Karr, the Koralpen Mts, Austria. C. The body of a male Nemastoma bidentatum gruberi ssp. nov., smeared over with the secretum; Kozje, Slovenia.
Fig. 4. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Ch basal articles (medial, lateral views).
Fig. 5. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Pa (medial view).
Fig. 6. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Synchroscopic images of the male Ch basal article and Pa-Fe (medial views, except G, right figure, which is in dorso-medial view).
Fig. 7. Nemastoma bidentatum Roewer, 1914 complex. Penes (lateral view). Note: The penis of N. pluridentatum (Hadži, 1973) stat. nov. is unknown.
Fig. 8. Nemastoma bidentatum Roewer, 1914 complex. Glans (dorsal and lateral views). Note: The penis of N. pluridentatum (Hadži, 1973) stat. nov. is unknown.
Fig. 9. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Variation of the Ch basal article (medial view) in species, subspecies and hybrids.
Fig. 10. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Variation of the Pa-Fe (A–B first image, C, D–E first image, F–G first image; H–J, L–M right, N–O: lateral view; the others in dorso-medial view to show the armament).
Fig. 11. Nemastoma bidentatum Roewer, 1914 complex, ♀♀; Rec sem (lateral view, right portion, except F left figure, which is in dorsal view). A. The frame encloses the ovipositor portion with Rec sem. Note: Female N. pluridentatum (Hadži, 1973) stat. nov. is unknown.
Fig. 12. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. From left to right: Ch basal article (medial, lateral views), Pa (medial view) and Pe tip (dorsal, lateral views). Arrows indicate the most identifying characters. Note: Pe of N. pluridentatum (Hadži, 1973) stat. nov. is unknown.
Fig. 13. Nemastoma bidentatum Roewer, 1914 complex, ♀♀. From left to right: Silhouettes of the Ch basal article, Pa (both medial view) and Rec sem. Arrows indicate the most identifying characters. Note: female N. pluridentatum (Hadži, 1973) stat. nov. is unknown.
B. Bertoncelj & U. Bertoncelj & T. Novak
T. Novak & M. Slana Novak & L. Slana Novak
L. Slana Novak & M. Slana Novak & T. Novak
V. Germovsek & T. Tkavc & A. Kocuvan & T. Novak