@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . cito:cites , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; dc:creator "Starkevich, Pavel; Podenas, Sigitas; Gelhaus, Jon K." ; dc:title "Tipula Linnaeus 1758" ; trt:augmentsTaxonConcept ; trt:citesTaxonName , , , , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "88" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:pubDate "2020-08-24" ; bibo:startPage "1" ; bibo:volume "4837" ; dc:creator "Starkevich, Pavel; Podenas, Sigitas; Gelhaus, Jon K." ; dc:date "2020" ; dc:title "Taxonomic review of Tipula (Vestiplex Bezzi) crane flies (Diptera: Tipulidae) in Mongolia" ; fabio:hasPart , , , , , , , , , , , , , , , , , , , , , , , , , ; a fabio:JournalArticle . dwc:authorityName "Linnaeus" ; dwc:authorityYear "1758" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Linnaeus, 1758" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Bezzi" ; dwc:authorityYear "1924" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:scientificNameAuthorship "Bezzi, 1924" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Schummel" ; dwc:authorityYear "1833" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Schummel, 1833" ; dwc:species "excisa" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Mik Mannheims" ; dwc:authorityYear "1889" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Mik Mannheims, 1889" ; dwc:species "leucoprocta" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "(Starkevich 2012)" ; dwc:baseAuthorityName "Starkevich" ; dwc:baseAuthorityYear "2012" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Starkevich, 2012)" ; dwc:species "virgatula" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Edwards, 1928" ; dwc:authorityName "Edwards" ; dwc:authorityYear "1928" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Edwards, 1928" ; dwc:species "avicularia" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Nielsen, 1929" ; dwc:authorityName "Nielsen" ; dwc:authorityYear "1929" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Nielsen, 1929" ; dwc:species "longitudinalis" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Alexander, 1934" ; dwc:authorityName "Alexander" ; dwc:authorityYear "1934" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Alexander, 1934" ; dwc:species "tchukchi" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Savchenko, 1960" ; dwc:authorityName "Savchenko" ; dwc:authorityYear "1960" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Savchenko, 1960" ; dwc:species "kiritshenkoi" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Lundstrom and Frey, 1915" ; dwc:authorityName "Lundstrom and Frey" ; dwc:authorityYear "1915" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lundstrom & Frey, 1915" ; dwc:species "laccata" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Mik, 1889" ; dwc:authorityName "Mik" ; dwc:authorityYear "1889" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Mik, 1889" ; dwc:species "mediovittata" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Meigen, 1804" ; dwc:authorityName "Meigen" ; dwc:authorityYear "1804" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Meigen, 1804" ; dwc:species "nubeculosa" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "virgatula Riedel, 1913" ; dwc:authorityName "virgatula Riedel" ; dwc:authorityYear "1913" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "virgatula Riedel, 1913" ; dwc:species "virgatula" ; dwc:subGenus "Vestiplex" ; dwc:subSpecies "virgatula" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Alexander" ; dwc:authorityName ": Alexander" ; dwc:authorityYear "1935" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Alexander, 1935" ; dwc:species "subcarinata" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Alexander, 1934" ; dwc:authorityName "Alexander" ; dwc:authorityYear "1934" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Alexander, 1934" ; dwc:species "pallitergata" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Alexander, 1918" ; dwc:authorityName "Alexander" ; dwc:authorityYear "1918" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Alexander, 1918" ; dwc:species "subcentralis" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Savchenko, 1960" ; dwc:authorityName "Savchenko" ; dwc:authorityYear "1960" ; dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Savchenko, 1960" ; dwc:species "coronifera" ; dwc:subGenus "Vestiplex" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Bibionidae" ; dwc:genus "Tipula" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Bibionidae" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:order "Diptera" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "FIGURES 1–3. Adults of Mongolian T. (Vestiplex), general view. 1. T. (Vestiplex) subcentralis, male habitus, left lateral view; 2. T. (Vestiplex) mediovittata, female habitus, left lateral view; 3. T. (Vestiplex) longitudinalis, female habitus, left lateral view." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 4–9. Gonocoxal fragments of Mongolian T. (Vestiplex), epandrium and hypandrium removed, dorsal view. 4. T. (Vestiplex) longitudinalis; 5. T. (Vestiplex) mediovittata; 6. T. (Vestiplex) scripta; 7. T. (Vestiplex) pallitergata; 8. T. (V). balioptera; 9. T. (Vestiplex) leucoprocta. Abbreviations: aa, anterior apodeme of medial sclerite of gonocoxal fragment; pa, posterior apodeme of medial sclerite of gonocoxal fragment; b, base of medial sclerite of gonocoxal fragment; gcx, gonocoxite; ig, inner gonostylus; il, inner lobe of lateral sclerite of gonocoxal fragment; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; ol, outer lobe of lateral sclerite of gonocoxal fragment. Scales of 4–6, 8–9 = 0.5 mm, 7 = 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 129–136. Hypopygium of male T. (Vestiplex) nubeculosa. 129. Tergite 9, dorsal view; 130. Gonocoxite, lateral view; 131. Left outer gonostylus, lateral view; 132. Left inner gonostylus, lateral view; 133. Gonocoxal fragment, dorsal view; 134. Aedeagal guide, lateral view; 135. Sperm pump, dorsal view; 136. Distal part of aedeagus, lateral view. Scale of 129–135 = 0.5 mm, 136 = 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 10–13. Aedeagus examples of Mongolian T. (Vestiplex), lateral view. 10. T. (Vestiplex) scripta; 11. T. (Vestiplex) mediovittata; 12. T. (Vestiplex) virgatula; 13. T. (Vestiplex) kiritshenkoi." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 163–170. Hypopygium of male T. (Vestiplex) longitudinalis. 163. Tergite 9, dorsal view; 164. Gonocoxite, lateral view; 165. Left outer gonostylus, lateral view; 166. Left inner gonostylus, lateral view; 167. Gonocoxal fragment, dorsal view; 168. Aedeagal guide, lateral view; 169. Sperm pump, dorsal view; 170. Distal part of aedeagus, lateral view. Scale of 163–169 = 0.5 mm, 170 = 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 174–181. Hypopygium of male T. (Vestiplex) virgatula. 174. Tergite 9, dorsal view; 175. Gonocoxite, lateral view; 176. Left outer gonostylus, lateral view; 177. Left inner gonostylus, lateral view; 178. Gonocoxal fragment, dorsal view; 179. Aedeagal guide, lateral view; 180. Sperm pump, dorsal view; 181. Distal part of aedeagus, lateral view. Abbreviation: gon, gonapophysis. Scale of 174–180 = 0.5 mm, 181 = 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 28–35. Hypopygium of male T. (Vestiplex) mediovittata. 28. Tergite 9, dorsal view; 29. Gonocoxite, lateral view; 30. Left outer gonostylus, lateral view; 31. Left inner gonostylus, lateral view; 32. Gonocoxal fragment, dorsal view; 33. Aedeagal guide, ventral view; 34. Sperm pump, dorsal view; 35. Distal part of aedeagus, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; pia, posterior immovable apodeme. Scales of 28–34 = 0.5 mm, 35 = 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 149–151. Ovipositor of female T. (Vestiplex) pallitergata. 149. Ovipositor, left lateral view; 150. Sternite 8 with hypovalvae, ventral view; 151. Sternite 9, furca and bursa copulatrix, dorsal view. Scale of 149 = 0.8 mm, 150–151 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 39–46. Hypopygium of male T. (Vestiplex) balioptera. 39. Tergite 9, dorsal view; 40. Gonocoxite, lateral view; 41. Left outer gonostylus, lateral view; 42. Left inner gonostylus, lateral view; 43. Gonocoxal fragment, dorsal view; 44. Aedeagal guide, lateral view; 45. Sperm pump, dorsal view; 46. Distal part of aedeagus, lateral view. Scales of 39–45 = 0.5 mm, 46 = 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 50–57. Hypopygium of male T. (Vestiplex) coronifera. 50. Tergite 9, dorsal view; 51. Gonocoxite, lateral view; 52. Left outer gonostylus, lateral view; 53. Left inner gonostylus, lateral view; 54. Gonocoxal fragment, dorsal view; 55. Aedeagal guide, lateral view; 56. Sperm pump, dorsal view; 57. Distal part of aedeagus, lateral view. Abbreviation: t, mid-dorsal teeth of inner gonostylus. Scales of 50–56 = 0.5 mm, 57 = 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 63–70. Hypopygium of male T. (Vestiplex) kiritshenkoi. 63. Tergite 9, dorsal view; 64. Gonocoxite, lateral view; 65. Left outer gonostylus, lateral view; 66. Left inner gonostylus, lateral view; 67. Gonocoxal fragment, dorsal view; 68. Aedeagal guide, lateral view; 69. Sperm pump, dorsal view; 70. Distal part of aedeagus, lateral view. Scales of 63–69 = 0.5 mm, 70 = 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 75–82. Hypopygium of male T. (Vestiplex) laccata. 75. Tergite 9, dorsal view; 76. Gonocoxite, lateral view; 77. Left outer gonostylus, lateral view; 78. Left inner gonostylus, lateral view; 79. Gonocoxal fragment, dorsal view; 80. Aedeagal guide, lateral view; 81. Sperm pump, dorsal view; 82. Distal part of aedeagus, lateral view. Scalse of 75–81 = 0.5 mm, 82 = 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 36–38. Ovipositor of female T. (Vestiplex) mediovittata. 36. Ovipositor, left lateral view; 37. Sternite 8 with hypovalvae, ventral view; 38. Sternite 9, furca and bursa copulatrix, dorsal view. Abbreviations: bc, bursa copulatrix; c, cercus; cds, cul-de-sac of bursa copulatrix; f, furca; h, hypovalva; s8, sternite 8; s9, sternite 9; t10, tergite 10. Scales of 36 = 0.8 mm, 37–38 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 47–49. Ovipositor of female T. (Vestiplex) balioptera. 47. Ovipositor, left lateral view; 48. Sternite 8 with hypovalvae, ventral view; 49. Sternite 9, furca and bursa copulatrix, dorsal view. Abbreviations: bp sd, basal part of spermathecal duct; li, lateral incision. Scales of 47 = 0.8 mm, 48–49 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 58–62. Ovipositor of female T. (Vestiplex) coronifera. 58. Ovipositor, left lateral view; 59. Sternite 8 with hypovalvae, ventral view; 60. Sternite 9, dorsal view; 61. Furca, dorsal view; 62. Bursa copulatrix, dorsal view. Scales of 58 = 0.8 mm, 59–62 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 71–74. Ovipositor of female T. (Vestiplex) kiritshenkoi. 71. Ovipositor, left lateral view; 72. Sternite 8 with hypovalvae, ventral view; 73. Sternite 9 and furca, dorsal view; 74. Bursa copulatrix, dorsal view. Scales of 71 = 0.8 mm, 72–74 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 83–85. Ovipositor of female T. (Vestiplex) laccata. 83. Ovipositor, left lateral view; 84. Sternite 8 with hypovalvae, ventral view; 85. Sternite 9, furca and bursa copulatrix, dorsal view. Scales of 83 = 0.8 mm, 84–85 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 93–95. Ovipositor of female T. (Vestiplex) leucoprocta. 93. Ovipositor, left lateral view; 94. Sternite 8 with hypovalvae, ventral view; 95. Sternite 9, furca and bursa copulatrix, dorsal view. Scales of 93 = 0.8 mm, 94–95 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 103–106. Ovipositor of female T. (Vestiplex) sintenisi. 103. Ovipositor, left lateral view; 104. Sternite 8 with hypovalvae, ventral view; 105. Sternite 9 and furca, dorsal view; 106. Bursa copulatrix, dorsal view. Scales of 103 = 0.8 mm, 104–106 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 114–116. Ovipositor of female T. (Vestiplex) subcentralis. 114. Ovipositor, left lateral view; 115. Sternite 8 with hypovalvae, ventral view; 116. Sternite 9, furca and bursa copulatrix, dorsal view. Scale of 114 = 0.5 mm, 115–116 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 125–128. Ovipositor of female T. (Vestiplex) tchukchi. 125. Ovipositor, left lateral view; 126. Sternite 8 with hypovalvae, ventral view; 127. Sternite 9, dorsal view; 128. Furca and bursa copulatrix, dorsal view. Scale of 125 = 0.8 mm, 126–127 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 137–141. Ovipositor of female T. (Vestiplex) nubeculosa. 137. Ovipositor, left lateral view; 138. Sternite 8 with hypovalvae, ventral view; 139. Sternite 9, dorsal view; 140. Furca, dorsal view; 141. Bursa copulatrix, dorsal view. Scale of 137 = 0.8 mm, 138–141 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 160–162. Ovipositor of female T. (Vestiplex) scripta. 160. Ovipositor, left lateral view; 161. Sternite 8 with hypovalvae, ventral view; 162. Sternite 9, furca and bursa copulatrix, dorsal view. Abbreviation: li, lateral incision. Scale of 160 = 0.7 mm, 161–162 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 171–173. Ovipositor of female T. (Vestiplex) longitudinalis. 171. Ovipositor, left lateral view; 172. Sternite 8 with hypovalvae, ventral view; 173. Sternite 9, furca and bursa copulatrix, dorsal view. Scale of 171 = 0.7 mm, 172–173 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 182–184. Ovipositor of female T. (Vestiplex) virgatula. 182. Ovipositor, left lateral view; 183. Sternite 8 with hypovalvae, ventral view; 184. Sternite 9, furca and bursa copulatrix, dorsal view. Scale of 182 = 0.8 mm, 183–184 = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 14–27. Spermathecae of Mongolian T. (Vestiplex). 14. T. (Vestiplex) longitudinalis; 15. T. (Vestiplex) virgatula; 16. T. (Vestiplex) pallitergata; 17. T. (Vestiplex) laccata; 18. T. (Vestiplex) balioptera; 19. T. (Vestiplex) nubeculosa; 20. T. (Vestiplex) scripta; 21. T. (Vestiplex) kiritshenkoi; 22. T. (Vestiplex) mediovittata; 23. T. (Vestiplex) subcentralis; 24. T. (Vestiplex) sintenisi; 25. T. (Vestiplex) coronifera; 26. T. (Vestiplex) tchukchi; 27. T. (Vestiplex) leucoprocta." ; fabio:hasRepresentation ; a fabio:Figure .