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@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
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@prefix trt: <http://plazi.org/vocab/treatment#> .
@prefix xlink: <http://www.w3.org/1999/xlink/> .

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    dc:creator "Portman, Zachary M.; Neff, John L.; Griswold, Terry" ;
    dc:title "Perdita bellula Timberlake" ;
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    trt:deprecates <http://taxon-concept.plazi.org/id/Animalia/Perdita_bellula_Timberlake_1954> ;
    trt:publishedIn <http://dx.doi.org/10.11646/zootaxa.4214.1.1> ;
    trt:treatsTaxonName <http://taxon-name.plazi.org/id/Animalia/Perdita_bellula> ;
    a trt:Treatment .

<http://dx.doi.org/10.11646/zootaxa.4214.1.1>
    bibo:endPage "97" ;
    bibo:issue "1" ;
    bibo:journal "Zootaxa" ;
    bibo:startPage "1" ;
    bibo:volume "4214" ;
    dc:creator "Portman, Zachary M.; Neff, John L.; Griswold, Terry" ;
    dc:date "2016" ;
    dc:title "Taxonomic revision of Perdita subgenus Heteroperdita Timberlake (Hymenoptera: Andrenidae), with descriptions of two ant-like males" ;
    fabio:hasPart <http://dx.doi.org/10.5281/zenodo.253090>, <http://dx.doi.org/10.5281/zenodo.253093>, <http://dx.doi.org/10.5281/zenodo.253110>, <http://dx.doi.org/10.5281/zenodo.253096>, <http://dx.doi.org/10.5281/zenodo.253109>, <http://dx.doi.org/10.5281/zenodo.253108>, <http://dx.doi.org/10.5281/zenodo.253106>, <http://dx.doi.org/10.5281/zenodo.253107>, <http://dx.doi.org/10.5281/zenodo.253105>, <http://dx.doi.org/10.5281/zenodo.253142>, <http://dx.doi.org/10.5281/zenodo.253144>, <http://dx.doi.org/10.5281/zenodo.253114>, <http://dx.doi.org/10.5281/zenodo.253101> ;
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<http://taxon-concept.plazi.org/id/Animalia/Perdita_bellula_Timberlake_1954>
    dwc:authority "Timberlake, 1954: 369" ;
    dwc:authorityName "Timberlake" ;
    dwc:authorityPageNumber "369" ;
    dwc:authorityYear "1954" ;
    dwc:class "Insecta" ;
    dwc:family "Andrenidae" ;
    dwc:genus "Perdita" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Hymenoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Timberlake, 1954" ;
    dwc:species "bellula" ;
    dwc:subGenus "Heteroperdita" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/6V3CV> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Perdita_bellula> ;
    a dwcFP:TaxonConcept .

<http://taxon-name.plazi.org/id/Animalia/Perdita_bellula>
    dwc:class "Insecta" ;
    dwc:family "Andrenidae" ;
    dwc:genus "Perdita" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Hymenoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:species "bellula" ;
    dwc:subGenus "Heteroperdita" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Perdita>, <http://taxon-name.plazi.org/id/Animalia/Perdita_Heteroperdita> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Perdita>
    dwc:class "Insecta" ;
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    dwc:genus "Perdita" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Hymenoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Andrenidae> ;
    a dwcFP:TaxonName .

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    dwc:class "Insecta" ;
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    dwc:order "Hymenoptera" ;
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    dwc:rank "family" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Hymenoptera> ;
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<http://taxon-name.plazi.org/id/Animalia/Hymenoptera>
    dwc:class "Insecta" ;
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    dwc:order "Hymenoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "order" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Insecta> ;
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    dwc:class "Insecta" ;
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    dwc:rank "class" ;
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    dwc:kingdom "Animalia" ;
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    dwc:rank "phylum" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
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<http://taxon-name.plazi.org/id/Animalia>
    dwc:kingdom "Animalia" ;
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<http://taxon-name.plazi.org/id/Animalia/Perdita_Heteroperdita>
    dwc:class "Insecta" ;
    dwc:family "Andrenidae" ;
    dwc:genus "Perdita" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Hymenoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Heteroperdita" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Perdita> ;
    a dwcFP:TaxonName .

<http://dx.doi.org/10.5281/zenodo.253090>
    dc:description "FIGURE 4. Second submedial cell of forewing. (A) Perdita vesca female, present (BBSL 816135; Harney Co., OR). (B) P. prodigiosa female, 2 m-cu vein weakened dorsally (paratype, BBSL 344037; Clark Co., NV). (C) P. bellula female, absent / weakened (BBSL 316794; Clark Co., NV). (D) P. wasbaueri male, absent / weakened (BBSL 842242; Imperial Co., CA)." ;
    fabio:hasRepresentation <https://zenodo.org/record/253090/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.253093>
    dc:description "FIGURE 7. Spotted female metasomas. (A) Perdita arenaria (BBSL 813432; Imperial Co., CA). (B) P. frontalis (BBSL 813797; Imperial Co., CA). (C) P. maculosa (BBSL 638406; Clark Co., NV). (D) P. nuttalliae (paratype, BBSL 816167; Harney Co., OR). (E) P. sycorax (paratype, BBSL 389685; Emery Co., UT). (F) P. titania (paratype, BBSL 342256; San Bernardino Co., CA). (G) P. wasbaueri (141836; Imperial Co., CA). (H) P. yanegai (BBSL 282782; Clark Co., NV). (I) P. bellula with distinct spots (FDP 161540; Riverside Co., CA). All scale bars 0.5 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/253093/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.253110>
    dc:description "FIGURE 24. Female pygidial plates. (A) Perdita arenaria (BBSL 289244; Clark Co., NV). (B) P. bellula (103744; San Bernardino Co., CA). (C) P. coldeniae (BBSL 813873; Imperial Co., CA). (D) P. desdemona (holotype, BBSL 289742; San Bernardino Co., CA). (E) P. exusta (paratype, 141946; Clark Co., NV). (F) P. frontalis (BBSL 813797; Imperial Co., CA). (G) P. hippolyta (paratype, BBSL 844647; Culberson Co., TX). (H) P. hooki (paratype, 141840; Terrell Co., TX). (I) P. maculosa (UCRC 179125; Riverside Co., CA). (J) P. nuttalliae (142005; Box Elder Co., UT). (K) P. optiva (BBSL 284974; Clark Co., NV). (L) P. pilonotata (neotype, UCRC 182770; Otero Co., NM). (M) P. prodigiosa (paratype, BBSL 319169; Clark Co., NV). (N) P. rhodogastra, pointed (FDP 102612; Maricopa Co., AZ). (O) P. rhodogastra, rounded (BBSL 271674; Clark Co., NV). (P) P. scutellaris (BBSL 904845; Imperial Co., CA). (Q) P. sexfasciata (BBSL 295303; Clark Co., NV). (R) P. sycorax (paratype, 37238; Emery Co., UT). (S) P. titania (holotype, BBSL 326372; San Bernardino Co., CA). (T) P. trifasciata (FDP 75046; Brewster Co., TX). (U) P. vesca (BBSL 339200; Nye Co., NV). (V) P. wasbaueri (141836; Imperial Co., CA). (W) P. yanegai (UCRC 221828; San Bernardino Co., CA). Scale bar = 0.1 mm. All images are at the same scale." ;
    fabio:hasRepresentation <https://zenodo.org/record/253110/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.253096>
    dc:description "FIGURE 10. Striped male metasomas. (A) Perdita optiva (BBSL 288698; Clark Co., NV). (B) P. sexfasciata (141743; Mojave Co., AZ). (C) P. trifasciata (141760; Terrell Co., TX). (D) P. bellula dark form (FDP 161544; Riverside Co., CA). (E) P. bellula typical light form (BBSL 817812; Mojave Co., AZ). (F) P. coldeniae (UCRC 169099; Imperial Co., CA). (G) P. scutellaris (DEVA 55309; San Bernardino Co., CA). All scale bars 0.5 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/253096/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.253109>
    dc:description "FIGURE 23. Male pygidial plates. (A) Perdita arenaria (BBSL 814703; Imperial Co., CA). (B) P. bellula (103717; San Bernardino Co., CA). (C) P. coldeniae (UCRC 277998; Imperial Co., CA). (D) P. exusta (141888; Clark Co., NV). (E) P. frontalis (BBSL 989204; Imperial Co., CA). (F) P. hippolyta (paratype, BBSL 844677; Culberson Co., TX). (G) P. hooki (note: the pygidial plate is from a dissected specimen and the lateral edges have been artificially enhanced to give a better idea of the lateral edges, paratype, BBSL 852902; Coahuila de Zaragoza, MEX). (H) P. maculosa (BBSL 668379; Clark Co., NV). (I) P. nuttalliae (paratype, BBSL 816190; Harney Co., OR). (J) P. optiva (BBSL 813428; Imperial Co., CA). (K) P. pilonotata (CAVE 17654; Eddy Co., NM). (L) P. prodigiosa (paratype, BBSL 344085; Clark Co., NV). (M) P. rhodogastra (UCRC 277717; Yuma Co., AZ). (N) P. scutellaris (DEVA 55305; San Bernardino Co., CA). (O) P. sexfasciata (141736; San Bernardino Co., CA). (P) P. sycorax (note: the picture is taken at an angle but it still accurately represents the shape the pygidial plate, paratype, 37225; Emery Co., UT). (Q) P. titania (pygidial plate is artificially outlined for better visibility, paratype, UCRC 166000; Imperial Co., CA). (R) P. trifasciata (141767; Chihuahua, MEX). (S) P. vesca (BBSL 816156; Harney Co., OR). (T) P. wasbaueri (BBSL 904842; Imperial Co., CA). (U) P. yanegai (UCRC 50407; Riverside Co., CA). Scale bar = 0.1 mm. All images are at the same scale." ;
    fabio:hasRepresentation <https://zenodo.org/record/253109/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.253108>
    dc:description "FIGURE 22. Miscellaneous female faces. (A) Perdita bellula (BBSL 813459; Imperial Co., CA). (B) P. desdemona (paratype, BBSL 289006; San Bernardino Co., CA). (C) P. exusta (paratype, 141952; Clark Co., NV). (D) P. vesca, dark coloration (BBSL 816172; Harney Co., OR). (E) P. vesca, light coloration (BBSL 904881; Nye Co., NV). (F) P. prodigiosa, dark coloration (paratype, BBSL 813546; Clark Co., NV). (G) P. prodigiosa, light coloration (paratype, BBSL 318844; Clark Co., NV). All scale bars 0.25 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/253108/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.253106>
    dc:description "FIGURE 20. Miscellaneous female metasomas. (A) Perdita bellula, typical coloration (103744; San Bernardino Co., CA). (B) P. bellula, melanized (BBSL 316794; Clark Co., NV). (C) P. desdemona (paratype, UCRC 185342; San Bernardino Co., CA). (D) P. exusta (paratype, 141946; Clark Co., NV). (E) P. prodigiosa, light coloration (BBSL 344016). (F) P. prodigiosa, dark coloration (paratype, BBSL 344044; Clark Co., NV). (G) P. rhodogastra (BBSL 271666; Clark Co., NV). (H) P. vesca, dark typical coloration (BBSL 816172; Harney Co., OR). (I) P. vesca, light coloration (BBSL 904881; Nye Co., NV). All scale bars 0.5 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/253106/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.253107>
    dc:description "FIGURE 21. Miscellaneous male faces. (A) Perdita bellula, large specimen (BBSL 842249; Imperial Co., CA). (B) P. bellula, small specimen (BBSL 817812; Mojave Co., AZ). (C) P. rhodogastra (BBSL 817150; Mojave Co., AZ). (D) P. exusta (141888; Clark Co., NV). (E) P. vesca, dark coloration (BBSL 816156; Harney Co., OR). (F) P. vesca, light coloration (BBSL 904888; Nye Co., NV). All scale bars 0.25 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/253107/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.253105>
    dc:description "FIGURE 19. Miscellaneous male metasomas. (A) Perdita bellula, typical form (141678; Yuma Co., AZ). (B) P. bellula, variation (BBSL 817827; Riverside Co., CA). (C) P. exusta (141895; Clark Co., NV). (D) P. rhodogastra (BBSL 309170; Clark Co., NV). (E) P. rhodogastra, dark form with light band (BBSL 290131; Clark Co., NV). (F) P. vesca, light coloration (BBSL 904888; Nye Co., NV). (G) P. vesca, spotted (BBSL 816156; Harney Co., NV). (H) P. vesca, solid coloration (141789; Clark Co., NV). All scale bars 0.5 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/253105/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.253142>
    dc:description "FIGURE 56. Male S 8 in dorsal view. (A) Perdita arenaria (BBSL 814693; Imperial Co., CA). (B) P. bellula (BBSL 842486; Riverside Co., CA). (C) P. coldeniae (BBSL 813338; Yuma Co., AZ). (D) P. exusta (BBSL 578345; Clark Co., NV). (E) P. frontalis (BBSL 813814; Imperial Co., CA). (F) P. hippolyta (paratype, CAVE 4578; Eddy Co., NM). (G) P. hooki (paratype, BBSL 852902; Coahuila, MEX). (H) P. maculosa (BBSL 638408; Clark Co., NV). (I) P. nuttalliae (BBSL 816192; Harney Co., OR). (J) P. optiva (BBSL 814695; Imperial Co., CA). (K) P. pilonotata (CAVE 17654; Eddy Co., NM). (L) P. prodigiosa (BBSL 344096; Clark Co., NV). Scale bar = 0.25 mm. All images are at the same scale." ;
    fabio:hasRepresentation <https://zenodo.org/record/253142/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.253144>
    dc:description "FIGURE 58. Male genital capsules in ventral and dorsal view. (A) Perdita arenaria dorsal and (B) ventral (BBSL 814693; Imperial Co., CA). (C) P. bellula dorsal and (D) ventral (BBSL 307306; Clark Co., NV). (E) P. coldeniae dorsal and (F) ventral (BBSL 817844; Riverside Co., CA). (G) P. exusta dorsal and (H) ventral (141875; Clark Co., NV). (I) P. frontalis dorsal and (J) ventral (BBSL 813814; Imperial Co., CA). (K) P. hippolyta dorsal and (L) ventral (paratype, CAVE 4578; Eddy Co., NM). (M) P. hooki dorsal and (N) ventral (paratype, BBSL 852902; Coahuila, MEX). Scale bar = 0.25 mm. All images are at the same scale." ;
    fabio:hasRepresentation <https://zenodo.org/record/253144/files/figure.png> ;
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<http://dx.doi.org/10.5281/zenodo.253114>
    dc:description "FIGURE 28. Distribution of (A) Perdita bellula and (B) P. coldeniae. Scale bar = 100 km." ;
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<http://dx.doi.org/10.5281/zenodo.253101>
    dc:description "FIGURE 15. Amber male metasomas. (A) Perdita rhodogastra, typical form (BBSL 817862; Riverside Co., CA). (B) P. rhodogastra, dark with medial bars (paratype, UCRC 185330; Imperial Co., CA). (C) P. hippolyta (paratype, BBSL 844677 Culberson Co., TX). (D) P. hooki (paratype, 141837; Terrell Co., TX). (E) P. bellula, cyanided (BBSL 315311; Clark Co., NV). All scale bars 0.5 mm." ;
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    dwc:collectionCode "CAS" ;
    dwc:locality "Palm Springs" ;
    dwc:typeStatus "holotype" ;
    trt:gbifOccurrenceId "1425219645" ;
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    dwc:collectionCode "CAS" ;
    dwc:eventDate "1941-06-28" ;
    dwc:locality "Palm Springs" ;
    dwc:recordedBy "P. H. Timberlake" ;
    dwc:typeStatus "holotype" ;
    trt:gbifOccurrenceId "1425219640" ;
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    dwc:collectionCode "CAS" ;
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    dwc:locality "Palm Springs" ;
    dwc:recordedBy "P. H. Timberlake" ;
    dwc:typeStatus "allotype" ;
    trt:gbifOccurrenceId "1425219656" ;
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    dwc:collectionCode "UCRC" ;
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    dwc:locality "Palm Springs" ;
    dwc:recordedBy "P. H. Timberlake" ;
    dwc:typeStatus "paratype" ;
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    dwc:locality "Gillespie Dam" ;
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    dwc:eventDate "2010-08-14" ;
    dwc:locality "Gila River" ;
    dwc:recordedBy "M. E. Irwin" ;
    trt:gbifOccurrenceId "1425219659" ;
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    dwc:eventDate "2010-08-07" ;
    dwc:locality "Mohave Co." ;
    dwc:recordedBy "M. E. Irwin & Littlefield & Interstate" ;
    trt:gbifOccurrenceId "1425219654" ;
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    dwc:eventDate "1983-05-13" ;
    dwc:locality "Yuma Co." ;
    dwc:recordedBy "P. F. Torchio & Yuma" ;
    trt:gbifOccurrenceId "1425219648" ;
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    dwc:eventDate "1957-09-02" ;
    dwc:locality "Imperial Co." ;
    dwc:recordedBy "T. R. Haig & Algodones Dunes & Glamis" ;
    trt:gbifOccurrenceId "1425219637" ;
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    dwc:eventDate "2012-09-07" ;
    dwc:locality "Glamis" ;
    dwc:recordedBy "Z. M. Portman & Glamis Dunes" ;
    trt:gbifOccurrenceId "1425219644" ;
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    dwc:eventDate "1983-09-13" ;
    dwc:locality "Riverside Co." ;
    dwc:recordedBy "P. F. & D. M. Torchio" ;
    trt:gbifOccurrenceId "1425219665" ;
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    dwc:eventDate "1976-10-05" ;
    dwc:locality "Baristo Rd." ;
    dwc:recordedBy "J. D. Pinto & Cathedral City, N" ;
    trt:gbifOccurrenceId "1425219667" ;
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    dwc:eventDate "2013-09-24" ;
    dwc:locality "Ramon Rd." ;
    dwc:recordedBy "Z. M. Portman & Cathedral City, N" ;
    trt:gbifOccurrenceId "1425219677" ;
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    dwc:eventDate "2000-04-22" ;
    dwc:locality "San Bernardino Co." ;
    dwc:recordedBy "G. R. Ballmer & Baker" ;
    trt:gbifOccurrenceId "1425219680" ;
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    dwc:eventDate "2003-09-21" ;
    dwc:locality "Mojave National Preserve" ;
    dwc:recordedBy "D. Yanega & Kelso Dunes & Devils Playground Wash & Kelso Dunes Road" ;
    trt:gbifOccurrenceId "1425219663" ;
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    dwc:eventDate "2012-09-06" ;
    dwc:locality "San Diego Co." ;
    dwc:recordedBy "Z. M. Portman & Borrego Springs" ;
    trt:gbifOccurrenceId "1425219678" ;
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    dwc:eventDate "2013-04-28" ;
    dwc:locality "West Valley" ;
    dwc:recordedBy "Z. M. Portman & Beehive Rock, W & Fire State Park" ;
    trt:gbifOccurrenceId "1425219683" ;
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    dwc:eventDate "1998-08-05" ;
    dwc:locality "Jean Lake" ;
    dwc:recordedBy "M. Andres & C. Schultz & Jean Lake, N" ;
    trt:gbifOccurrenceId "1425219675" ;
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    dwc:eventDate "1998-08-11" ;
    dwc:locality "Jean Lake" ;
    dwc:recordedBy "T. L. Griswold & Jean Lake, N" ;
    trt:gbifOccurrenceId "1425219661" ;
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    dwc:eventDate "1998-08-11" ;
    dwc:locality "Jean Lake" ;
    dwc:recordedBy "M. Andres & T. L. Griswold & C. Schultz & Jean Lake, NE" ;
    trt:gbifOccurrenceId "1425219676" ;
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    dwc:eventDate "1998-10-08" ;
    dwc:locality "Jean Lake" ;
    dwc:recordedBy "T. L. Griswold & Jean Lake, NE" ;
    trt:gbifOccurrenceId "1425219682" ;
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    dwc:eventDate "1998-05-13" ;
    dwc:locality "Mesquite Lake" ;
    dwc:recordedBy "K. Receveur & C. Schultz & Mesquite Lake, E" ;
    trt:gbifOccurrenceId "1425219671" ;
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    dwc:eventDate "1998-08-05" ;
    dwc:locality "Virgin Valley" ;
    dwc:recordedBy "M. Andres & C. Schultz & Riverside & New Gold Butte Rd" ;
    trt:gbifOccurrenceId "1425219681" ;
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    dwc:eventDate "2012-05-16" ;
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    dwc:eventDate "2012-07-23" ;
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    trt:gbifOccurrenceId "1425219672" ;
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    dwc:eventDate "2005-07-18" ;
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    dwc:recordedBy "E. Green" ;
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    dwc:eventDate "2005-05-25" ;
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    dwc:recordedBy "S. M. Higbee & E. Ahlstrom" ;
    trt:gbifOccurrenceId "1425219673" ;
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    dwc:eventDate "1998-06-07" ;
    dwc:locality "St. Thomas Gap" ;
    dwc:recordedBy "F. D. Parker" ;
    trt:gbifOccurrenceId "1425219669" ;
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    dwc:eventDate "1998-08-27" ;
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    dwc:recordedBy "O. J. Messinger & S. Messinger & C. Schultz" ;
    trt:gbifOccurrenceId "1425219670" ;
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    dwc:eventDate "1998-06-29" ;
    dwc:locality "St. Thomas Gap" ;
    dwc:recordedBy "M. Andres & K. Keen & K. Receveur & C. Schultz" ;
    trt:gbifOccurrenceId "1425219684" ;
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    dwc:eventDate "1998-08-04" ;
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    dwc:recordedBy "M. Andres & C. Schultz" ;
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    dwc:eventDate "1998-05-21" ;
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