@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , ; dc:creator "Kemper, Jenny M." ; dc:title "Chimaera willwatchi Kemper 2017, sp. nov." ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , ; trt:citesTaxonName , , , , , ; trt:definesTaxonConcept ; trt:deprecates , , , , , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "37" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:pubDate "2017-08-28" ; bibo:startPage "1" ; bibo:volume "4312" ; dc:creator "Kemper, Jenny M." ; dc:date "2017" ; dc:title "New species of Chimaera (Chondrichthyes: Holocephali: Chimaeriformes: Chimaeridae) from the Southwestern Indian Ocean" ; fabio:hasPart , , , , , ; a fabio:JournalArticle . dwc:authority "Kemper, 2017" ; dwc:authorityName "Kemper" ; dwc:authorityYear "2017" ; dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Kemper, 2017" ; dwc:species "willwatchi" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Linnaeus" ; dwc:authorityYear "1758" ; dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Linnaeus, 1758" ; dwc:species "monstrosa" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Tanaka" ; dwc:authorityYear "1905" ; dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Tanaka, 1905" ; dwc:species "owstoni" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Didier" ; dwc:authorityYear "1998" ; dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Didier, 1998" ; dwc:species "panthera" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Last, White & Pogonoski" ; dwc:authorityYear "2008" ; dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Last, White & Pogonoski, 2008" ; dwc:species "argiloba" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Didier, Last & White" ; dwc:authorityYear "2008" ; dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Didier, Last & White, 2008" ; dwc:species "fulva" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Kemper, Ebert, Didier & Compagno" ; dwc:authorityYear "2010" ; dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Kemper, Ebert, Didier & Compagno, 2010" ; dwc:species "bahamaensis" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Tanaka" ; dwc:authorityYear "1905" ; dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Tanaka, 1905" ; dwc:species "jordani" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "willwatchi" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "monstrosa" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "owstoni" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "panthera" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "argiloba" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "cubana" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "fulva" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "phantasma" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "notafricana" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "obscura" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "opalescens" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "bahamaensis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "carophila" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Holocephali" ; dwc:family "Chimaeridae" ; dwc:genus "Chimaera" ; dwc:kingdom "Animalia" ; dwc:order "Chimaeriformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "jordani" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURE 8. Chimaera willwatchi, sp. nov., holotype CAS 242336, mature male 834 mm TL, 492 mm BDL illustrations of a) Frontal tenaculum, b) Lateral VieW of pelVic claspers, c) Pre-pelVic tenacula, d) Tooth plates, and e) Skeletonized skull." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 10. Chimaera willwatchi, sp. nov., morph A a) Holotype CAS 242336, mature male 834 mm TL, 492 mm BDL, b) Paratype, immature male, 620 mm TL, 323 mm TL, c) Paratype, neWly hatched female, CAS 242351, 290 mm TL, 117.58 mm BDL." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 9. Map of study area denoting Where Chimaera willwatchi, sp. nov., specimens Were collected, With subpopulations represented by circles (morph A), star (morph B), and triangles (morph C)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 11. Chimaera willwatchi, sp. nov., morph B a) Non-type CAS, immature male, 694 mm TL, 369 mm BDL, b) Nontype CAS 242354, immature female, 477 mm TL, 252 mm BDL." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 12. Chimaera willwatchi, sp. nov., morph C a) Non-type CAS 242358, immature female, 975 mm TL, 542 mm BDL, b) Non-type CAS 242342, immature female, 620 mm TL, 323 mm BDL," ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 13. MaXimum likelihood tree topology based on a general-time reVersible (GTR) substitution model + gamma distribution for Chimaera willwatchi, sp. nov., Chimaera didierae, sp. nov., Chimaera buccanigella, sp. nov., and comparartiVe species based on aligned NADH 2 DNA sequences. GenBank accession numbers folloW species (see AppendiX 2). Bootstrap support Values of main clades shoWn on tree. Specimens indicated in bold are designated holotypes." ; fabio:hasRepresentation ; a fabio:Figure . dwc:catalogNumber "CAS 242336, 834" ; dwc:collectionCode "BDL" ; dwc:eventDate "2014-05-24" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "850" ; dwc:verbatimLatitude "-33.916668" ; dwc:verbatimLongitude "55.266666" ; trt:gbifOccurrenceId "1587010221" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "CAS 242339" ; dwc:collectionCode "BDL" ; dwc:eventDate "2014-04-28" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "Southwest Indian Ocean Ridge & P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1008" ; dwc:verbatimLatitude "-35.133335" ; dwc:verbatimLongitude "55.283333" ; trt:gbifOccurrenceId "1587010244" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "CAS 242354" ; dwc:collectionCode "BDL" ; dwc:eventDate "2014-04-30" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1240" ; dwc:verbatimLatitude "-29.85" ; dwc:verbatimLongitude "46.05" ; trt:gbifOccurrenceId "1587010222" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "USNM 440273" ; dwc:collectionCode "BDL" ; dwc:eventDate "2014-04-31" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1200" ; dwc:verbatimLatitude "-35.716667" ; dwc:verbatimLongitude "53.716667" ; trt:gbifOccurrenceId "1587010230" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "MCZ 171972" ; dwc:collectionCode "BDL" ; dwc:eventDate "2014-04-23" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1110" ; dwc:verbatimLatitude "-38.4" ; dwc:verbatimLongitude "48.366665" ; trt:gbifOccurrenceId "1587010231" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "SIO 16-67" ; dwc:collectionCode "BDL" ; dwc:eventDate "2012-03-18" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "970" ; dwc:verbatimLatitude "-35.133335" ; dwc:verbatimLongitude "53.7" ; trt:gbifOccurrenceId "1587010228" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "CAS 242338" ; dwc:collectionCode "BDL" ; dwc:eventDate "2014-03-17" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1180" ; dwc:verbatimLatitude "-35.133335" ; dwc:verbatimLongitude "53.7" ; trt:gbifOccurrenceId "1587010235" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "CAS 242337" ; dwc:collectionCode "BDL" ; dwc:eventDate "2014-03-26" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1118" ; dwc:verbatimLatitude "-33.933334" ; dwc:verbatimLongitude "55.283333" ; trt:gbifOccurrenceId "1587010233" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "CAS 242343" ; dwc:collectionCode "BDL" ; dwc:eventDate "2014-04-28" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1190" ; dwc:verbatimLatitude "-39.033333" ; dwc:verbatimLongitude "46.55" ; trt:gbifOccurrenceId "1587010236" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "CAS 242367" ; dwc:collectionCode "BDL" ; dwc:eventDate "2012-03-17" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1178" ; dwc:verbatimLatitude "-34.733334" ; dwc:verbatimLongitude "43.733334" ; trt:gbifOccurrenceId "1587010224" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "USNM 440274" ; dwc:collectionCode "BDL" ; dwc:eventDate "2014-04-04" ; dwc:locality "Southwestern Indian Ocean Ridge" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1180" ; dwc:verbatimLatitude "-35.133335" ; dwc:verbatimLongitude "53.7" ; trt:gbifOccurrenceId "1587010240" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "SAIAB 203575" ; dwc:collectionCode "BDL" ; dwc:eventDate "2014-04-30" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1240" ; dwc:verbatimLatitude "-35.15" ; dwc:verbatimLongitude "53.716667" ; trt:gbifOccurrenceId "1587010243" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "SIO 16-68" ; dwc:collectionCode "BDL" ; dwc:eventDate "2014-04-23" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1200" ; dwc:verbatimLatitude "-35.133335" ; dwc:verbatimLongitude "53.7" ; trt:gbifOccurrenceId "1587010226" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BDL" ; dwc:eventDate "2014-04-30" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1240" ; dwc:verbatimLatitude "-38.366665" ; dwc:verbatimLongitude "47.583332" ; trt:gbifOccurrenceId "1587010239" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BDL" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "B. Walkins & Januaray" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "960" ; dwc:verbatimLatitude "-39.833332" ; dwc:verbatimLongitude "45.783333" ; trt:gbifOccurrenceId "1587010242" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BDL" ; dwc:eventDate "2001-05-03" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "B. Walkins" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "982" ; dwc:verbatimLatitude "-39.833332" ; dwc:verbatimLongitude "45.783333" ; trt:gbifOccurrenceId "1587010241" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BDL" ; dwc:eventDate "2001-05-03" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "B. Walkins" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "982" ; dwc:verbatimLatitude "-39.433334" ; dwc:verbatimLongitude "41.333332" ; trt:gbifOccurrenceId "1587010225" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BDL" ; dwc:eventDate "2001-05-06" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "B. Walkins" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "890" ; dwc:verbatimLatitude "-39.433334" ; dwc:verbatimLongitude "41.316666" ; trt:gbifOccurrenceId "1587010237" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BDL" ; dwc:locality "Males" ; dwc:verbatimElevation "715" ; dwc:verbatimLatitude "-39.533333" ; dwc:verbatimLongitude "53.7" ; trt:gbifOccurrenceId "1587010234" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BDL" ; dwc:eventDate "2014-03-23" ; dwc:locality "Southwestern Indian Ocean" ; dwc:recordedBy "P. J. Clerkin & R. Downie" ; dwc:verbatimElevation "1328" ; dwc:verbatimLatitude "-39.033333" ; dwc:verbatimLongitude "55.266666" ; trt:gbifOccurrenceId "1587010223" ; trt:httpUri ; a dwc:MaterialCitation .