@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; dc:creator "Kuwahara, Gregory K.; Marshall, Stephen A." ; dc:title "Howickia Richards 1951" ; trt:augmentsTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "152" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:pubDate "2022-10-03" ; bibo:startPage "1" ; bibo:volume "5192" ; dc:creator "Kuwahara, Gregory K.; Marshall, Stephen A." ; dc:date "2022" ; dc:title "A revision of the Australian species of Howickia Richards (Diptera: Sphaeroceridae: Limosininae)" ; fabio:hasPart , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; a fabio:JournalArticle . dwc:authorityName "Richards" ; dwc:authorityYear "1951" ; dwc:class "Insecta" ; dwc:family "Sphaeroceridae" ; dwc:genus "Howickia" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Richards, 1951" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Sphaeroceridae" ; dwc:genus "Howickia" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Sphaeroceridae" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Diptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "FIGURES 24.1–24.3. Howickia wilsoni (Richards). 24.1, female habitus, lateral; 24.2, female head, anterolateral, 24.3, left wing. Scale bars indicate 0.5 mm. Photo credit: R. Cox." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 25.1–25.5. Howickia xanthocephala sp. nov. 25.1, male habitus, lateral; 25.2, male head, anterolateral; 25.3 male S5–8, ventral; 25.4, male terminalia, lateral; 25.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 8.1–8.6. Howickia erythrocephala sp. nov. 8.1, male habitus, lateral; 8.2, male head, anterolateral; 8.3, left wing; 8.4, male S5–8, ventral; 8.5, male terminalia, lateral; 8.6, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 13.1–13.5. Howickia kurandensis (Richards). 13.1, male habitus, lateral; 13.2, male head, anterolateral; 13.3, male S5–8, ventral; 13.4, male terminalia, lateral; 13.4a, left surstylus, posterior; 13.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 14.1–14.5. Howickia myrmecophila (Knab & Malloch). 14.1, male habitus, lateral; 14.2, male head, anterolateral; 14.3, male S5–8, ventral; 14.4, male terminalia, lateral; 14.5, aedeagus and associated structures, lateral; 14.5a, apex of distiphallus, anterior. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 15.1–15.5. Howickia nitidipleura (Richards). 15.1, male habitus, lateral; 15.2, male head, anterolateral; 15.3, male S5, ventral; 15.4, male terminalia, lateral; 15.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 22.1–22.5. Howickia vernalis (Richards). 22.1, male habitus, lateral; 22.2, male head, anterolateral; 22.3, male S5–8, ventral; 22.4, male terminalia, lateral; 22.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 3.1–3.22. Mid tibial chaetotaxy of macropterous Australian species of Howickia. 3.1, H. acicula sp. nov., a) dorsal, male, b) lateral male, c) lateral, female; 3.2, H. biantenna sp. nov., a) dorsal, male, b) lateral male, c) lateral, female; 3.3, H. brevior (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.4, H. capitalis (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.5, H. erythrocephala sp. nov., a) dorsal, male, b) lateral female; 3.6, H. flaviterga (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.7, H. fuscalis (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.8, H. grandisterna sp. nov., a) dorsal, male, b) lateral male, c) lateral, female; 3.9, H. hardyina (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.10, H. kurandensis (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.11, H. myrmecophila (Knab & Malloch), a) dorsal, male, b) lateral male, c) lateral, female; 3.12, H. nitidipleura (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.13, H. percostata (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.14, H. pertusa sp. nov., a) dorsal, male, b) lateral male, c) lateral, female; 3.15, H. pseudovittata sp. nov., a) dorsal, male, b) lateral male, c) lateral, female; 3.16, H. subsinuata (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.17, H. symmetrica (Richards), a) dorsal, male, b) lateral male; 3.18, H. trivittata (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.19, H. vernalis (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.20, H. wamini sp. nov., a) dorsal, male, b) lateral male, c) lateral, female; 3.21, H. wilsoni (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.22, H. xanthocephala sp. nov., a) dorsal, male, b) lateral male, c) lateral, female. Scale bars indicate 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 21.1–21.5. Howickia trivittata (Richards). 21.1, male habitus, lateral; 21.2, male head, anterolateral; 21.3, male S5–8, ventral; 21.4, male terminalia, lateral; 21.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 19.1–19.5. Howickia subsinuata (Richards). 19.1, male habitus, lateral; 19.2, male head, anterolateral; 19.3, male S5–8, ventral; 19.4, male terminalia, lateral; 19.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 18.1–18.5. Howickia pseudovittata sp. nov. 18.1, male habitus, lateral; 18.2, male head, anterolateral; 18.3, male S5–8, ventral; 18.4, male terminalia, lateral; 18.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 6.1–6.2. Howickia brevior (Richards). 6.1, female habitus, lateral; 6.2, female head, anterolateral.Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 20.1–20.3. Howickia symmetrica (Richards). 20.1, male habitus, lateral; 20.2, male head, anterolateral; 20.3, left wing. Body and wing scale bars indicate 0.5 mm, head scale bar indicates 0.2 mm. Photo credit: J. Lumbers." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 12.1–12.5. Howickia hardyina (Richards). 12.1, male habitus, lateral; 12.2, male head, anterolateral; 12.3, male S5–8, ventral; 12.4, male terminalia, lateral; 12.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 5.1–5.5. Howickia biantenna sp. nov. 5.1, male habitus, lateral; 5.2, male head, anterolateral; 5.3, male S5–8, ventral; 5.4, male terminalia, lateral; 5.4a, left surstylus, posterior; 5.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 17.1–17.5. Howickia pertusa sp. nov. 17.1, male habitus, lateral; 17.2, male head, anterolateral; 17.3, male S5–8, ventral; 17.4, male terminalia, lateral; 17.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 23.1–23.5. Howickia wamini sp. nov. 23.1, male habitus, lateral; 23.2, male head, anterolateral; 23.3, male S5–8, ventral; 23.4, male terminalia, lateral; 23.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 7.1–7.2. Howickia capitalis (Richards). 7.1, female habitus, lateral; 7.2, female head, anterolateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 16.1–16.5. Howickia percostata (Richards). 16.1, male habitus, lateral; 16.2, male head, anterolateral; 16.3, male S5–8, ventral; 16.4, male terminalia, lateral; 16.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 10.1–10.5. Howickia fuscalis (Richards). 10.1, male habitus, lateral; 10.2, male head, anterolateral; 10.3, male S5–8, ventral; 10.4, male terminalia, lateral; 10.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 9.1–9.5. Howickia flaviterga (Richards). 9.1, male habitus, lateral; 9.2, male head, anterolateral; 9.3, male S5–8, ventral; 9.4, male terminalia, lateral; 9.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 4.1–4.5. Howickia acicula sp. nov. 4.1, male habitus, lateral; 4.2, male head, anterolateral; 4.3, male S5–8, ventral; 4.4, male terminalia, lateral; 4.5, aedeagus and associated structures, lateral. Abbreviations: bas—basiphallus, cer—cercus, dist—distiphallus, epa—epandrium, hyp—hypandrium, phal—phallapodeme, post—postgonite, S—sternite, sur—surstylus. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 11.1–11.5. Howickia grandisterna sp. nov. 11.1, male habitus, lateral; 11.2, male head, anterolateral; 11.3, male S5–8, ventral; 11.4, male terminalia, lateral; 11.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 26.1–26.2. Howickia cruspica sp. nov. 26.1, female habitus, lateral; 26.2, female head, anterolateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 27.1–27.5. Howickia pinnula sp. nov. 27.1, male habitus, lateral; 27.2, male head, anterolateral; 27.3, male S5–8, ventral; 27.4, male terminalia, lateral; 27.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 28.1–28.6. Howickia sabina (Richards). 28.1, male habitus, lateral; 28.2, male head, anterolateral; 29.3, right wing; 28.4, male S5–8, ventral; 28.5, male terminalia, lateral; 28.6, aedeagus and associated structures, lateral. Body and head scale bars indicate 0.5 mm, wing scale bar indicates 0.2 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 30.1–30.5. Howickia australis (Papp). 30.1, male habitus, lateral; 30.2, male head, anterolateral; 30.3, male S5–8, ventral; 30.4, male terminalia, lateral; 30.4a, left surstylus, posterior; 30.4b, left cercus, posterior; 30.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 29.1–29.5. Howickia auricoma sp. nov. 29.1, male habitus, lateral; 29.2, male head, anterolateral; 29.3, male S5–8, ventral; 29.4, male terminalia, lateral; 29.4a, left surstylus, posterior; 29.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 42.1–42.6. Howickia regalis (Richards). 42.1, light morph male habitus, lateral; 42.2, light morph male head, anterolateral; 42.3, dark morph male habitus, lateral; 42.4, dark morph male head, anterolateral; 42.5, darkest morph male habitus, lateral; 42.6, darkest morph male head, anterolateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 44.1–44.5. Howickia tricolor sp. nov. 44.1, male habitus, lateral; 44.2, male head, anterior; 44.3, male S5–8, ventral; 44.4, male terminalia, lateral; 44.5, aedeagus and associated structures, lateral. Body scale bar indicates 0.5 mm, head scale bar indicates 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 36.1–36.5. Howickia flavithorax (Papp). 36.1, male habitus, lateral; 36.2, male head, anterolateral; 36.3, male S5–8, ventral; 36.4, male terminalia, lateral; 36.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 43.1–43.5. Howickia robustaseta sp. nov. 43.1, male habitus, lateral; 43.2, male head, anterolateral; 43.3, male S5–8, ventral; 43.4, male terminalia, lateral; 43.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 38.1–38.5. Howickia irwini sp. nov. 38.1, male habitus, lateral; 38.2, male head, anterolateral; 38.3, male S5–8, ventral; 38.4, male terminalia, lateral; 38.4a, left surstylus, posterior; 38.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 34.1–34.5. Howickia flagella sp. nov. 34.1, male habitus, lateral; 34.2, male head, anterolateral; 34.3, male S5–8, ventral; 34.4, male terminalia, lateral; 34.4a, left surstylus, posterior; 34.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 37.1–37.5. Howickia holti sp. nov. 37.1, male habitus, lateral; 37.2, male head, anterolateral; 37.3, male S5–8, ventral; 37.4, male terminalia, lateral; 37.4a, left surstylus, posterior; 37.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 40.1–40.5. Howickia notechis sp. nov. 40.1, female habitus, lateral; 40.2, female head, anterolateral; 40.3, male S5–8, ventral; 40.4, male terminalia, lateral; 40.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 45.1–45.5. Howickia truemani sp. nov. 45.1, male habitus, lateral; 45.2, male head, anterolateral; 45.3, male S5–8, ventral; 45.4, male terminalia, lateral; 45.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 41.1–41.5. Howickia nuda sp. nov. 41.1, male habitus, lateral; 41.2, male head, anterolateral; 41.3, male S5–8, ventral; 41.4, male terminalia, lateral; 41.5, aedeagus and associated structures, lateral. Scale bars indicate 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 32.1–32.5. Howickia dealata (Richards). 32.1, male habitus, lateral; 32.2, male head, anterior; 32.3, epiproct and cercus, dorsal; 32.4, S8 and hypoproct, ventral; 32.5, spermathecae. Body scale bar indicates 0.25 mm, head bar indicates 0.1 mm. Photo credit: G. Thompson." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 31.1–31.5. Howickia cynthia (Richards). 31.1, male habitus, lateral; 31.2, male head, anterior; 31.3, male S5–8, ventral; 31.4, male terminalia, lateral; 31.4a, left surstylus, posterior; 31.5, aedeagus and associated structures, lateral. Body scale bar indicates 0.25 mm, head bar indicates 0.1 mm. Photo credit: G. Thompson." ; fabio:hasRepresentation ; a fabio:Figure .