@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , , , , ; dc:creator "Fernández, Mercedes; Fernicola, Juan C.; Cerdeño, Esperanza" ; dc:title "Protypotherium praerutilum Ameghino 1887" ; dwc:basisOfRecord ; trt:augmentsTaxonConcept ; trt:deprecates , , , , , , , , , , , , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "444" ; bibo:issue "2" ; bibo:journal "Zoological Journal of the Linnean Society" ; bibo:pubDate "2023-07-11" ; bibo:startPage "417" ; bibo:volume "199" ; dc:creator "Fernández, Mercedes; Fernicola, Juan C.; Cerdeño, Esperanza" ; dc:date "2023" ; dc:title "Systematic revision of the species of Protypotherium (Notoungulata: Interatheriidae) from the Santa Cruz Formation (Early-Middle Miocene), Argentinian Patagonia: a new phylogenetic hypothesis for the Interatheriidae" ; fabio:hasPart , , , , , , ; a fabio:JournalArticle . dwc:authority "Ameghino, 1887 b" ; dwc:authorityName "Ameghino" ; dwc:authorityYear "1887" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ameghino, 1887" ; dwc:species "praerutilum" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ameghino, 1891 b" ; dwc:authorityName "Ameghino" ; dwc:authorityYear "1891" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Patriarchus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ameghino, 1891" ; dwc:species "leptocephalus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ameghino, 1891 b" ; dwc:authorityName "Ameghino" ; dwc:authorityYear "1891" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ameghino, 1891" ; dwc:species "convexidens" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ameghino, 1887 b" ; dwc:authorityName "Ameghino" ; dwc:authorityYear "1887" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ameghino, 1887" ; dwc:species "attenuatum" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ameghino, 1891 b" ; dwc:authorityName "Ameghino" ; dwc:authorityYear "1891" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Patriarchus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ameghino, 1891" ; dwc:species "rectus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ameghino, 1891 b" ; dwc:authorityName "Ameghino" ; dwc:authorityYear "1891" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Patriarchus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ameghino, 1891" ; dwc:species "diastematus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ameghino, 1894" ; dwc:authorityName "Ameghino" ; dwc:authorityYear "1894" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Patriarchus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ameghino, 1894" ; dwc:species "icochiloides" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ameghino, 1891 b" ; dwc:authorityName "Ameghino" ; dwc:authorityYear "1891" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ameghino, 1891" ; dwc:species "diversidens" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ameghino, 1891 a" ; dwc:authorityName "Ameghino" ; dwc:authorityYear "1891" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Eudiastatus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ameghino, 1891" ; dwc:species "lingulatus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ameghino, 1894" ; dwc:authorityName "Ameghino" ; dwc:authorityYear "1894" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ameghino, 1894" ; dwc:species "lineare" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ameghino, 1891 b" ; dwc:authorityName "Ameghino" ; dwc:authorityYear "1891" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ameghino, 1891" ; dwc:species "globosum" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Roth, 1899" ; dwc:authorityName "Roth" ; dwc:authorityYear "1899" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Icochilus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Roth, 1899" ; dwc:species "endiadys" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Ameghino" ; dwc:authorityYear "1885" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Ameghino, 1885" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Roth" ; dwc:baseAuthorityYear "1898" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Roth, 1898)" ; dwc:species "endiadys" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Ameghino" ; dwc:baseAuthorityYear "1887" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Ameghino, 1887)" ; dwc:species "australe" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Ameghino" ; dwc:authorityYear "1891" ; dwc:baseAuthorityName "Roth" ; dwc:baseAuthorityYear "1898" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Patriarchus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Roth, 1898)" ; dwc:species "diastematus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Ziflel" ; dwc:authorityYear "1893" ; dwc:class "Mammalia" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "subOrder" ; dwc:scientificNameAuthorship "Ziflel, 1893" ; dwc:subOrder "Typotheria" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Ameghino" ; dwc:authorityYear "1889" ; dwc:baseAuthorityName "Roth" ; dwc:baseAuthorityYear "1898" ; dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Patriarchus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "(Roth, 1898)" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "praerutilum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Patriarchus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "leptocephalus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Patriarchus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "convexidens" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "attenuatum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Patriarchus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "rectus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Patriarchus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "diastematus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Patriarchus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "icochiloides" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "diversidens" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Eudiastatus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "lingulatus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Eudiastatus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "lineare" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "globosum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Icochilus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "endiadys" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Icochilus" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "endiadys" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:family "Interatheriidae" ; dwc:genus "Protypotherium" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "australe" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Mammalia" ; dwc:kingdom "Animalia" ; dwc:order "Notoungulata" ; dwc:phylum "Chordata" ; dwc:rank "subOrder" ; dwc:subOrder "Typotheria" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "Figure 2. Protypotherium australe (D–E, G, I–J, L) and Pr. praerutilum (A–C, F, H, K, M), in occlusal view. (A) YPM-VPPU 15386, incomplete skull; (B) less I3 (alveolus)–C–dP1–P2–M3 of MACN-A 4049; (C) less I3–dP1–M3 of MACN-A 4278; (D) right I1 (alveolus)–dP1–P2–M3 of MACN-A 3882; (E) right I1–dP1–P2–M3 of MACN-A 9623–9633; (F) less I1 (alveolus)–dP1–P2–M3 of AMNH 9187; (G) right I1–2 (alveolus)–dP1–P2–M3 of YPM-VPPU 15643; (H) MLP 12-2886, incomplete skull; (I) right I1 (alveolus)–I2–dP1–P2–M3 of MACN-A 37; (J) MACN-A 3911, less maxillary fragment with C–dP1–P2–M3; (K) MACN-A 1778, right maxillary fragment with dP1–P2 (alveoli)– P3–M3; (L) right I1 (alveolus)–dP1–P2–M3 of AMNH 9149; (M) MACN-A 4016, less maxillary fragment with P4–M3.* indicates alveolus. Scale bar = 10 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 3. Protypotherium australe (B, F–M) and Pr. praerutilum (A, C–E), in occlusal view. (A) MACN-A 3952, right mandibular fragment with p3–m3; (B) less dp1–p2–m3 of AMNH 9286; (C) MACN-A 11191a, right mandibular fragment with m1–3; (D) right p2–m3 of MACN-A 9614; (E) right i1–dp1–p2–m3 of MACN-A 1082; (F) less p4–m3 of AMNH 9565; (G) right p2–m3 of AMNH 9149; (H) less i1–dp1– p2–m3 of MPM-PV 3531; (I) MACN-A 532, right mandibular fragment with m2–3; (J) right p2–m3 of MPM-PV 3739; (K) MPM-PV 19612, right mandibular fragment with m2 (broken)–3; (L) MACN-A 3979, less mandibular fragment with p3–m3; (M) MACN-A 3955, less mandibular fragment with p3–m3. * indicates alveolus; ’ indicates broken tooth. Scale bar = 10 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)–p3–m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)–m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1–dp1–p2–p4, and right i1 and dp1–p2–m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3–M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)–m3; (G) MACN-A 3867, right mandibular fragment with dp1–2–p3–4 (trigonid); (H) MACN-A 9716–9732?g, right mandibular fragment with m1 (talonid)–3; (I) MPM-PV 19761, right maxillary fragment with M1–2; (J) MPM-PV 19781, right mandibular fragment with p2–m1; (K) MPM-PV 19782, less maxillary fragment with P2–M3. Abbreviation: ’, indicates broken tooth. Scale bar = 10 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 7. Box-plot showing the length of I1–3 in: (A) Pr.praerutilum; (B) Pr.australe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 5. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs.width of M1. B, Length vs. width of m1. Abbreviations: L, length; W, width." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 6. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs. width of M2. B, Length vs. width of m2. Abbreviations: L, length; W, width." ; fabio:hasRepresentation ; a fabio:Figure . dwc:catalogNumber "MACN-A 1081, MACN-A 1082" ; dwc:collectionCode "MACN-A" ; dwc:locality "Early - Middle Miocene" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "4425895303" ; trt:httpUri ; a dwc:MaterialCitation .