@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , , , , , ; dc:creator "López-García, Margarita M.; Deloya, Cuauhtémoc" ; dc:title "Ligyrus (Ligyrus) villosus" ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; trt:augmentsTaxonConcept ; trt:deprecates ; trt:publishedIn ; a trt:Treatment . bibo:endPage "119" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:pubDate "2022-11-22" ; bibo:startPage "1" ; bibo:volume "5211" ; dc:creator "López-García, Margarita M.; Deloya, Cuauhtémoc" ; dc:date "2022" ; dc:title "Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" ; fabio:hasPart , , , , , , , , , , ; a fabio:JournalArticle . dwc:authority "(Burmeister, 1847)" ; dwc:baseAuthorityName "Burmeister" ; dwc:baseAuthorityYear "1847" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Burmeister, 1847)" ; dwc:species "villosus" ; dwc:subGenus "Ligyrus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Burmeister, 1847: 120" ; dwc:authorityName "Burmeister" ; dwc:authorityPageNumber "120" ; dwc:authorityYear "1847" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Podalgus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Burmeister, 1847" ; dwc:species "villosus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "villosus" ; dwc:subGenus "Ligyrus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:subGenus "Ligyrus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Podalgus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "villosus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Podalgus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURE 2. Labium. A) Ligyrus (Anagrylius) moroni, B) Ligyrus (Ligyrus) allonasutus, C) L. (Ligyrus) gibbosus, D) L. (Ligyrus) villosus, E) L. (Ligyrodes) relictus, F) Proculigyrus cicatricosus, G) Euligyrus ebenus, H) E. similis, I) Tomarus gyas, J) Tomarus selanderi, K) Tomarus subtropicus. Abbreviations: LL: Ligular lobes, LP: Labial palpus, M: Mentum, S: Suture." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 4. Maxilla. A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) relictus, D) L. (Ligyrus) burmeisteri, E) L. (Ligyrus) villosus, F) Tomarus gyas, G) Tomarus maternus, H) T. selanderi, I) T. subtropicus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 5. Mandible. A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) relictus, D) L. (Ligyrus) burmeisteri, E) L. (Ligyrus) villosus, F) Tomarus rostratus, G) T. gyas, H) T. maternus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 6. Labrum. A) Ligyrus (Ligyrus) burmeisteri, B) L. (Ligyrus) villosus, C) Ligyrus (Ligyrodes) relictus, D) Tomarus gyas, E) T. maternus, F) T. selanderi, G) T. subtropicus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 11. Protibia in frontal view.A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) relictus, E) L. (Ligyrodes) sallaei, F) L. (Ligyrus) allonasutus, G) L. (Ligyrus) villosus, H) Tomarus bituberculatus, I) T. colombianus, J) T. pullus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 12. Mesotibia (A–F). A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei; E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) villosus. Metatibia (G–L). G) Euligyrus similis, H) Proculigyrus cicatricosus, I) Ligyrus (Anagrylius) moroni, J) L. (Ligyrodes) sallaei, K) L. (Ligyrus) allonasutus, L) L. (Ligyrus) villosus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 14. Spiculum gastrale. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) L. (Ligyrodes) relictus, D) L. (Ligyrus) villosus, E) Tomarus pullus, F) T. rostratus, G) T. selanderi, H) T. subtropicus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 18. Parameres in lateral view.A) L. (Ligyrus.) allonasutus, B) L. (Ligyrus) bidentulus, C) L. (Ligyrus) burmeisteri, D) Ligyrus (Ligyrus) fossor, E) L. (Ligyrus) gibbosus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) neglectus, H) L. (Ligyrus) paranaensis, I) L. (Ligyrus) peninsularis, J) L. (Ligyrus) rubripes, K) L. (Ligyrus) ruginasus, L) L. (Ligyrus) villosus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 26. Habitus of the species of Ligyrus. Subgenus Ligyrus: A) Ligyrus fossor, B) L. gibbosus, C) L. nasutus, D) L. neglectus, E) L. paranaensis, F) L. peninsularis, G) L. rubripes, H) L. ruginasus, I) L. villosus. Scale bar: 5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 35. Distributional records of Ligyrus (Ligyrus) villosus, L. bidentulus, and L. rubripes." ; fabio:hasRepresentation ; a fabio:Figure . dwc:locality "Santa Rosa" ; trt:gbifOccurrenceId "3986696714" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "The" ; trt:gbifOccurrenceId "3986697006" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696779" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696311" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696995" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696931" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696954" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696565" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696328" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696794" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696875" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986697040" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696415" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696511" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696723" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696983" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696634" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696681" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986697011" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696501" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696498" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696526" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696476" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986697039" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696688" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696915" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696639" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696630" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696942" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986697014" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696774" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696784" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696724" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696615" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696895" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696900" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696420" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696491" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696925" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696471" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696339" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696534" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696360" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696686" ; trt:httpUri ; a dwc:MaterialCitation .