@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . # Warning: Failed to output a material citation, could not create identifier # Warning: Failed to output a material citation, could not create identifier # Warning: Failed to output a material citation, could not create identifier # Warning: Failed to output a material citation, could not create identifier # Warning: Failed to output a material citation, could not create identifier cito:cites , , , ; dc:creator "Rossini, Michele; Montreuil, Olivier; Grebennikov, Vasily; Tarasov, Sergei" ; dc:title "Helictopleurus sicardi" ; trt:augmentsTaxonConcept ; trt:deprecates ; trt:publishedIn ; a trt:Treatment . bibo:endPage "79" ; bibo:journal "ZooKeys" ; bibo:pubDate "2021-04-22" ; bibo:startPage "63" ; bibo:volume "1033" ; dc:creator "Rossini, Michele; Montreuil, Olivier; Grebennikov, Vasily; Tarasov, Sergei" ; dc:date "2021" ; dc:title "Genome sequencing reveals extraordinary cephalic horns in the Madagascan dung beetle genus Helictopleurus (Coleoptera, Scarabaeinae): insight from a revision of fungicola species group" ; fabio:hasPart , ; a fabio:JournalArticle . dwc:LSID "35214DD4-6AC9-52B0-A271-E3EF3027DBFD" ; dwc:authority "(Paulian, 1975)" ; dwc:baseAuthorityName "Paulian" ; dwc:baseAuthorityYear "1975" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Helictopleurus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Paulian, 1975)" ; dwc:species "sicardi" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Paulian" ; dwc:authorityYear "1975" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Heterosyphus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Paulian, 1975" ; dwc:species "sicardi" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Paulian" ; dwc:authorityYear "1975" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Heterosyphus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Paulian, 1975" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Orbigny" ; dwc:authorityYear "1915" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Helictopleurus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Orbigny, 1915" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Helictopleurus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "sicardi" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Helictopleurus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Heterosyphus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "sicardi" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Heterosyphus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "Figure 4. Helictopleurus sicardi A, B major (A) and minor (B) male dorsal habitus C major male lateral habitus D female dorsal habitus (lectotype) and original labels E propleural groove (PpG) F VIII interstrial ridge (IR) G, H lateral and anterior view of the aedeagus; basoventral lobes (BVLb) I endophallus: endophallic spines (EpS); superior (SpLC) and inferior (IpLC) part of the lamella copulatrix." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 1. Molecular phylogenies and morphological synapomorphies of the Helictopleurus fungicola species group A phylogenetic position of H. sicardi within Helictopleurus. Helictopleurus neoamplicollis and H. carbonarius are highlighted with colored dashed line to indicate their previous placement B COI-based phylogeny: magnification of the Helictopleurus fungicola clade and phylogenetic position of H. peyrierasi stat. rest. C synapomorphies of the Helictopleurus fungicola species group: basal pygidial ridge (BPyR); mesotarsal teeth (MsT) on the first tarsomere; parameres (P) elongated and evenly curved downward, basoventral lobes of the parameres bell shaped (BVLb); superior side of the membranous sac of the endophallus with spines (EpS); lamella copulatrix very simple (LC) D detail of the clypeal horns of the male of H. sicardi." ; fabio:hasRepresentation ; a fabio:Figure .