@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , ; dc:creator "Angus, Robert B.; Jia, Fenglong; Chen, Zhen-ning; Zhang, Ying; Vondráček, Dominik; Fikáček, Martin; Sw, London" ; dc:title "Helophorus (Lihelophorus) ser Zaitzev 1908" ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; trt:augmentsTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "148" ; bibo:issue "1" ; bibo:journal "Acta Entomologica Musei Nationalis Pragae" ; bibo:pubDate "2016-07-15" ; bibo:startPage "109" ; bibo:volume "56" ; dc:creator "Angus, Robert B.; Jia, Fenglong; Chen, Zhen-ning; Zhang, Ying; Vondráček, Dominik; Fikáček, Martin; Sw, London" ; dc:date "2016" ; dc:title "Taxonomy, larval morphology and cytogenetics of Lihelophorus, the Tibetan endemic subgenus of Helophorus (Coleoptera: Hydrophiloidea)" ; fabio:hasPart , , , , , ; a fabio:JournalArticle . dwc:authority "Zaitzev, 1908" ; dwc:authorityName "Zaitzev" ; dwc:authorityYear "1908" ; dwc:class "Insecta" ; dwc:family "Helophoridae" ; dwc:genus "Helophorus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Zaitzev, 1908" ; dwc:species "ser" ; dwc:subGenus "Lihelophorus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Helophoridae" ; dwc:genus "Helophorus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "ser" ; dwc:subGenus "Lihelophorus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Helophoridae" ; dwc:genus "Helophorus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:subGenus "Lihelophorus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Helophoridae" ; dwc:genus "Helophorus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Helophoridae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "Fig. 1. Adult morphology of Helophorus (Lihelophorus). a–b – H. yangae sp. nov. (a – ventral view of the head; b – detail of mentum and labial palps); c–d – maxillary galea and lacinia (c – H. yangae sp. nov.; d – H. ser Zaitzev, 1908); e–f – mandible (e – H. yangae sp. nov.; f – H. ser); g – labial palpus of H. ser; h–l – hind legs in dorsal view (h – H. yangae sp. nov.; i – H. ser; j – H. lamicola Zaitzev, 1908; k – detail of apex of tibia and base of tarsus of H. lamicola; l – detail of tarsomere 5 and claw of H. ser). Abbreviations: sh – swimming hairs. Not to scale." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 2. Adult morphology of Helophorus (Lihelophorus). a–f – details of left half of pronotum (a,c,e) and base of left elytron (b, d, f): a–b – H. yangae sp. nov.; c–d – H. ser Zaitzev, 1908; e–f – H. lamicola Zaitzev, 1908; arrow in (a) shows remnants of pronotal granulations, arrows in (b, d, f) show scutellary stria. g–h – prothoracic hypomeron and ‘suprapleural area’ (g – H. yangae sp. nov.; h – H. ser). i – detail of posterior margin of abdominal ventrite 5 of H. ser, showing the irregular denticulation. j – ventral view of the thorax of H. ser. k – dorsolateral portion of left elytron of H. lamicola. Abbreviations: ep – epipleuron; li – last elytral interval. Not to scale." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 3. Habitus of the species of Helophorus (Lihelophorus). a–c – H. lamicola Zaitzev, 1908 (a – Qinghai: Zhaling Hu (lectotype); b–c – Quinghai: Golo).d–g – H.ser Zaitzev, 1908 (d–e – Qinghai:Golo; f – Xizang, Nyainqentanglha Feng; g – Gansu: Xiahe env.). h–k – H. yangae sp. nov. (h – Xizang: 90 km W of Amdo; i – Xizang: 25 km W of Amdo; j – Qinghai: Golo; k – India: Kar Tso (Orchymont coll.)). All specimens to scale." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig.4. Details of the head and pronotum of the Helophorus (Lihelophorus) specimens.a–c – H. lamicola Zaitzev,1908 (a – Qinghai: Zhaling Hu (lectotype); b–c – Qinghai: Golo). d–h – H. ser Zaitzev, 1908 (d – Qinghai: valley of Alag Hu lake (lectotype); e–f – Qinghai: Golo; g – Xizang, Nyainqentanglha Feng; h – Gansu: Xiahe env.). Not to scale." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. a–e – H. (L.) ser Zaitzev, 1908 (a – lectotype, Qinghai: valley of Alag Hu lake; b–c – Xizang: valley SE of Dong La pass; d – Qinghai: Golo, 20 km E of Maduo; e – Qinghai: Zuimatan). f–j – H. (L.) lamicola Zaitzev, 1908 (f – lectotype, Qinghai: Djarin Nor; g–i – Qinghai: Zuimatan; j – paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). k–o – H. (L.) yangae sp. nov. (k – holotype, Qinghai: Zuimatan; l–m – paratypes, same locality; n – Qinghai: Golo, Yematan; o – Xizang: Gongbujiangda)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 5. Details of the head and pronotum of the Helophorus (Lihelophorus) yangae sp. nov. (a – Xizang: 90 km W of Amdo; b,d – Qinghai: Golo; c – Xizang: 25 km W of Amdo; e – India: Kar Tso (Orchymont coll.). Not to scale." ; fabio:hasRepresentation ; a fabio:Figure . dwc:catalogNumber "ANGUS 1995" ; dwc:collectionCode "ZIN" ; dwc:eventDate "1900-05-30" ; dwc:locality "Bydda" ; dwc:recordedBy "Ekcp. & Eksp. Kozlova. & Hel." ; dwc:typeStatus "lectotype" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IRSNB" ; dwc:eventDate "1906-08" ; dwc:locality "Westtibet" ; dwc:recordedBy "Zugmayer & Orchymont & Knisch" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SE" ; dwc:eventDate "2011-06-18" ; dwc:locality "Dongo La Pass" ; dwc:recordedBy "S. Tibet & J. Schmidt" ; dwc:verbatimLatitude "28.766666" ; dwc:verbatimLongitude "87.95" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NMPC" ; dwc:eventDate "2010-07-20" ; dwc:locality "Lha Tsu Vall" ; dwc:recordedBy "Schmidt" ; dwc:verbatimElevation "5175" ; dwc:verbatimLatitude "30.379723" ; dwc:verbatimLongitude "90.72278" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NMPC" ; dwc:eventDate "2005-06-11" ; dwc:locality "Monda La" ; dwc:recordedBy "J. Schmidt & J. Hajek & D. Kral & J. Ruzicka & Gangca Dasi" ; dwc:verbatimElevation "5060" ; dwc:verbatimLatitude "35.191666" ; dwc:verbatimLongitude "102.51" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NMPC" ; dwc:eventDate "2005-07-13" ; dwc:locality "Haibu" ; dwc:recordedBy "J. Hajek & D. Kral & Ruzicka" ; dwc:verbatimElevation "3230" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NMPC" ; dwc:eventDate "2005-07-03" ; dwc:locality "Hairag" ; dwc:recordedBy "Hajek & Kral & Ruzicka" ; dwc:verbatimElevation "3240" ; dwc:verbatimLatitude "37.168335" ; dwc:verbatimLongitude "100.41167" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2013-06-08" ; dwc:locality "Maduo" ; dwc:recordedBy "R. B. Angus & F. L. Jia & Zhang" ; dwc:verbatimElevation "4290" ; dwc:verbatimLatitude "34.85472" ; dwc:verbatimLongitude "98.28833" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2013-06-07" ; dwc:locality "Zuimatan" ; dwc:recordedBy "Angus & Jia & Zhang" ; dwc:verbatimElevation "4141" ; dwc:verbatimLatitude "35.331112" ; dwc:verbatimLongitude "99.065834" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NMPC" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "1990-08-15" ; dwc:locality "Kangzhagri" ; dwc:recordedBy "Xuezhong Zhang" ; dwc:verbatimElevation "5130" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "1990-08-02" ; dwc:locality "Mt." ; dwc:recordedBy "Kekexili & Xinqingfeng & Xuezhong Zhang" ; dwc:verbatimElevation "4900" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "1990-08-01" ; dwc:locality "Malanshan" ; dwc:recordedBy "Xuezhong Zhang" ; dwc:verbatimElevation "4950" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "1990-08-10" ; dwc:locality "Kusaihu lake" ; dwc:recordedBy "Xuezhong Zhang" ; dwc:verbatimElevation "4600" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU, IRSNB" ; dwc:eventDate "1932-08-10" ; dwc:locality "Yematan" ; dwc:recordedBy "R. B. Angus & F. L. Jia & Y. Zhang & Indian Tibet, L & Chushol & Orchymont" ; dwc:verbatimElevation "4240" ; dwc:verbatimLatitude "34.67972" ; dwc:verbatimLongitude "99.065834" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "2013-06-07" ; dwc:locality "Zuimatan" ; dwc:recordedBy "R. B. Angus & F. L. Jia & Zhang" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "4141" ; dwc:verbatimLatitude "35.331112" ; dwc:verbatimLongitude "99.065834" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "2013-06-12" ; dwc:locality "Zuimatan" ; dwc:recordedBy "R. B. Angus & Angus" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "4141" ; dwc:verbatimLatitude "35.331112" ; dwc:verbatimLongitude "99.065834" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "2013-06-07" ; dwc:locality "Zuimatan" ; dwc:recordedBy "R. B. Angus & F. L. Jia & Zhang" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "4141" ; dwc:verbatimLatitude "35.331112" ; dwc:verbatimLongitude "99.065834" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2013-06-08" ; dwc:locality "Yematan" ; dwc:recordedBy "R. B. Angus & F. L. Jia & Y. Zhang & Angus" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "4240" ; dwc:verbatimLatitude "34.67972" ; dwc:verbatimLongitude "99.065834" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:typeStatus "paratype" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NMPC" ; dwc:typeStatus "paratype" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "2013-06-08" ; dwc:locality "Heihexiang" ; dwc:recordedBy "R. B. Angus & F. L. Jia & Zhang" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "4240" ; dwc:verbatimLatitude "34.67972" ; dwc:verbatimLongitude "99.065834" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2013-06-08" ; dwc:locality "Maduo" ; dwc:recordedBy "R. B. Angus & F. L. Jia & Zhang" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "4290" ; dwc:verbatimLatitude "34.85472" ; dwc:verbatimLongitude "98.28833" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "1986-07-20" ; dwc:locality "Gangca" ; dwc:recordedBy "F. L. Jia & Y. Zhang & R. B. Angus & Rikaze & Liang Ge-Qiu" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "3370" ; dwc:verbatimLatitude "37.3" ; dwc:verbatimLongitude "100.183334" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "1986-07-27" ; dwc:locality "Jiangzi" ; dwc:recordedBy "Liang Ge-Qiu" ; dwc:typeStatus "paratype" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "2013-08-06" ; dwc:locality "Yangbajing town" ; dwc:recordedBy "Jia Yue & Jia Fenglong" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "4320" ; dwc:verbatimLatitude "30.09533" ; dwc:verbatimLongitude "90.53926" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "2013-08-02" ; dwc:locality "Sangye Fery" ; dwc:recordedBy "Jia Yue & Jia Fenglong" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "3658" ; dwc:verbatimLatitude "29.32309" ; dwc:verbatimLongitude "91.509384" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "2013-08-06" ; dwc:locality "Sangye" ; dwc:recordedBy "Jia Yue & Jia Fenglong" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "3575" ; dwc:verbatimLatitude "29.323748" ; dwc:verbatimLongitude "91.50373" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:locality "Sangye town" ; dwc:recordedBy "Jia Yue & Jia Fenglong" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "3620" ; dwc:verbatimLatitude "29.32309" ; dwc:verbatimLongitude "91.509384" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "2013-08-05" ; dwc:locality "Langkazi" ; dwc:recordedBy "Jia Yue & Jia Fenglong" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "5030" ; dwc:verbatimLatitude "28.96803" ; dwc:verbatimLongitude "90.39797" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "2013-08-03" ; dwc:locality "Yangzhuoyongcuo lake" ; dwc:recordedBy "Jia Fenglong & Jia Yue" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "4447" ; dwc:verbatimLatitude "28.915985" ; dwc:verbatimLongitude "90.72477" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "2013-08-02" ; dwc:locality "Gongbujiangda" ; dwc:recordedBy "Jia Fenglong & Jia Yue" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "4155" ; dwc:verbatimLatitude "29.88528" ; dwc:verbatimLongitude "93.24607" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SYSU" ; dwc:eventDate "2013-08-02" ; dwc:locality "Jiamudui village" ; dwc:recordedBy "Jia Yue" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "3872" ; dwc:verbatimLatitude "29.153214" ; dwc:verbatimLongitude "89.0113" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NHMW" ; dwc:locality "90 km W Amdo" ; dwc:recordedBy "Cigetang & X. Guo" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "4400" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NHMW, IRSNB" ; dwc:locality "Couna Lake" ; dwc:recordedBy "X. Guo & Indian Tibet & Tzo-Kar & Kar Tso & Startsakpuk Tso & Zaitzev'" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "4400" ; trt:httpUri ; a dwc:MaterialCitation .