@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , ; dc:creator "Génier, François; Moretto, Philippe" ; dc:title "Digitonthophagus namaquensis Genier, new species" ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; trt:definesTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "110" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:startPage "1" ; bibo:volume "4248" ; dc:creator "Génier, François; Moretto, Philippe" ; dc:date "2017" ; dc:title "Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)" ; fabio:hasPart , , , , , ; a fabio:JournalArticle . dwc:authority "Genier" ; dwc:authorityName "Genier" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Digitonthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Genier" ; dwc:species "namaquensis" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Digitonthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "namaquensis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Digitonthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "FIGURES 19 – 22. Male dorsal habitus. 19. Digitonthophagus fimator, holotype; 20. D. lusinganus, lectotype; 21. D. ulcerosus, holotype; 22. D. namaquensis, holotype." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 47 – 54. 47, 49, 51, 53. Male head, dorsal view; 48, 50, 52, 54. Male head, frontal view. 47 – 48. Digitonthophagus fimator, holotype; 49 – 50. D. lusinganus, lectotype; 51 – 52. D. ulcerosus, holotype; 53 – 54. D. namaquensis, holotype." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 63 – 70. Male pronotum, dorsal view. 63. D. petilus, holotype; 64. Digitonthophagus viridicollis, holotype; 65. D. aksumensis, holotype; 66. D. dilatatus, holotype; 67. D. fimator, holotype; 68. D. lusinganus, lectotype; 69. D. ulcerosus, holotype 70. D. namaquensis, holotype." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 79 – 86. Female head, dorsal view. 79. Digitonthophagus petilus, allotype; 80. D. viridicollis, allotype; 81. D. aksumensis, allotype; 82. D. dilatatus, allotype; 83. D. fimator, allotype; 84. D. lusinganus, paralectotype; 85. D. ulcerosus, allotype 86. D. namaquensis, allotype." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 119 – 134. Parameres, dorsal view. 119. D. bonasus, alloreferent (Tamil Nadu State, India); 120. Digitonthophagus uks, holotype; 121. D. sahelicus, holotype; 122. D. catta, alloreferent (West Bengal, India); 123. D. eucatta, holotype; 124. D. falciger, holotype; 125. D. biflagellatus, holotype; 126. D. gazella, non type (Antisiranana Province, Madagascar); 127. D. petillus, holotype; 128. D. viridicollis, holotype; 129. D. aksumensis, holotype; 130. D. dilatatus, holotype; 131. D. fimator, holotype; 132. D. lusinganus, lectotype; 133. D. ulcerosus, holotype 134. D. namaquensis, holotype." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 171 – 182. Internal sac sclerites (FLP, A + SA). 171, 174, 177, 180. Dorsal view; 172, 175, 178, 181. Ventral view. 173, 176, 179, 182. Right lateral view. 171 – 173. Digitonthophagus fimator, holotype; 174 – 176. D. lusinganus, lectotype; 177 – 179. D. ulcerosus, holotype; 180 – 182. D. namaquensis, holotype." ; fabio:hasRepresentation ; a fabio:Figure . dwc:locality "Northern Cape" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "957" ; dwc:verbatimLatitude "-28.87137" ; dwc:verbatimLongitude "21.30944" ; trt:gbifOccurrenceId "1455384277" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Material" ; trt:gbifOccurrenceId "1455384287" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IRSNB" ; dwc:locality "Benguela" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-12.583333" ; dwc:verbatimLongitude "13.416667" ; trt:gbifOccurrenceId "1455384279" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "1972-01-31" ; dwc:locality "Karibib" ; dwc:recordedBy "Ameib Farm" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-21.783333" ; dwc:verbatimLongitude "15.633333" ; trt:gbifOccurrenceId "1455384286" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:locality "Southern" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455384283" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "ZMUC" ; dwc:eventDate "1978-02-10" ; dwc:locality "Gobabeb" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-23.566668" ; dwc:verbatimLongitude "15.033334" ; trt:gbifOccurrenceId "1455384291" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "1972-01-30" ; dwc:locality "River" ; dwc:recordedBy "N Usakos" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-22.0" ; dwc:verbatimLongitude "15.6" ; trt:gbifOccurrenceId "1455384295" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1972-01-22" ; dwc:locality "Southern" ; dwc:recordedBy "Kuiseb Canyon" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-23.3" ; dwc:verbatimLongitude "15.75" ; trt:gbifOccurrenceId "1455384292" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:locality "Southern" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455384273" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMNS" ; dwc:eventDate "2010-02-16" ; dwc:locality "Tiras Mountains" ; dwc:recordedBy "Farm Lovedale, N" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1600" ; dwc:verbatimLatitude "-25.539999" ; dwc:verbatimLongitude "16.39" ; trt:gbifOccurrenceId "1455384275" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1972-01-19" ; dwc:locality "Maltahoe District" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-24.183332" ; dwc:verbatimLongitude "15.983334" ; trt:gbifOccurrenceId "1455384284" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:locality "Voigtgrund" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-24.816668" ; dwc:verbatimLongitude "17.433332" ; trt:gbifOccurrenceId "1455384294" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OUMNH" ; dwc:eventDate "2000-01-16" ; dwc:locality "Mann" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-24.049444" ; dwc:verbatimLongitude "16.153889" ; trt:gbifOccurrenceId "1455384297" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OUMNH" ; dwc:eventDate "2000-01-17" ; dwc:locality "Mann" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-24.075556" ; dwc:verbatimLongitude "16.4725" ; trt:gbifOccurrenceId "1455384280" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "1972-01-17" ; dwc:locality "KARAS" ; dwc:recordedBy "Barby Farm & W Helmeringhausen" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-25.883333" ; dwc:verbatimLongitude "16.816668" ; trt:gbifOccurrenceId "1455384281" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:locality "Southern" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455384276" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMF" ; dwc:locality "Keetmannshoop" ; dwc:recordedBy "S. Hardt" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-26.566668" ; dwc:verbatimLongitude "18.133333" ; trt:gbifOccurrenceId "1455384290" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1972-01-10" ; dwc:locality "Grunau" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-27.383333" ; dwc:verbatimLongitude "18.466667" ; trt:gbifOccurrenceId "1455384293" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "1972-01-08" ; dwc:locality "RIVER" ; dwc:recordedBy "Onseepkans" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-28.733334" ; dwc:verbatimLongitude "19.3" ; trt:gbifOccurrenceId "1455384282" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:locality "Southern" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455384278" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "1972-01-14" ; dwc:locality "Plateau" ; dwc:recordedBy "E Aus" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-26.633333" ; dwc:verbatimLongitude "16.566668" ; trt:gbifOccurrenceId "1455384288" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:locality "Southern" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455384289" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1934-02-16" ; dwc:locality "Seeheim" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-26.816668" ; dwc:verbatimLongitude "17.783333" ; trt:gbifOccurrenceId "1455384285" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OUMNH" ; dwc:eventDate "1999-12-19" ; dwc:locality "Mann" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-17.865278" ; dwc:verbatimLongitude "19.93" ; trt:gbifOccurrenceId "1455384311" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OUMNH" ; dwc:eventDate "2002-03-23" ; dwc:locality "Sovo area" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-18.320557" ; dwc:verbatimLongitude "19.257778" ; trt:gbifOccurrenceId "1455384300" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMNS" ; dwc:eventDate "2010-02-03" ; dwc:locality "Spreetshoogte" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1800" ; dwc:verbatimLatitude "-23.37" ; dwc:verbatimLongitude "16.14" ; trt:gbifOccurrenceId "1455384305" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1972-01-22" ; dwc:locality "Solitaire" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-23.883333" ; dwc:verbatimLongitude "16.0" ; trt:gbifOccurrenceId "1455384308" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMF" ; dwc:locality "Windhoek" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-22.570002" ; dwc:verbatimLongitude "17.083612" ; trt:gbifOccurrenceId "1455384304" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1977-04-02" ; dwc:locality "Kamanjab" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-19.816668" ; dwc:verbatimLongitude "14.166667" ; trt:gbifOccurrenceId "1455384318" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMNS" ; dwc:eventDate "2004-03" ; dwc:locality "Orupembe" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-18.166666" ; dwc:verbatimLongitude "12.566667" ; trt:gbifOccurrenceId "1455384314" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMNS" ; dwc:eventDate "2005-04-18" ; dwc:locality "Palmwag Lodge" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-19.88611" ; dwc:verbatimLongitude "13.936945" ; trt:gbifOccurrenceId "1455384312" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMNS" ; dwc:eventDate "2005-04-15" ; dwc:locality "Hoaruzsib Valley" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-18.766666" ; dwc:verbatimLongitude "12.95" ; trt:gbifOccurrenceId "1455384313" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OUMNH" ; dwc:eventDate "1999-12-28" ; dwc:locality "Khoanib" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-19.245556" ; dwc:verbatimLongitude "13.342777" ; trt:gbifOccurrenceId "1455384299" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OUMNH" ; 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a dwc:MaterialCitation . dwc:collectionCode "NMPC" ; dwc:eventDate "1916-02" ; dwc:locality "Willowmore" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-33.3" ; dwc:verbatimLongitude "23.483334" ; trt:gbifOccurrenceId "1455384322" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2004-03-12" ; dwc:locality "NORTHERN CAPE" ; dwc:recordedBy "Farm Klipkoppies" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "957" ; dwc:verbatimLatitude "-28.87137" ; dwc:verbatimLongitude "21.309439" ; trt:gbifOccurrenceId "1455384301" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BDGC, CMNC, FGIC, OUMNH, PMOC" ; dwc:eventDate "2001-03-23" ; dwc:locality "A" ; dwc:recordedBy "Groblershoop & Farm Brulpan" ; dwc:typeStatus "holotype" ; dwc:verbatimLatitude "-28.89889" ; dwc:verbatimLongitude "21.780277" ; trt:gbifOccurrenceId "1455384309" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "2004-03-06" ; dwc:locality "A" ; dwc:recordedBy "Keograbie Farm" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "1038" ; dwc:verbatimLatitude "-29.076" ; dwc:verbatimLongitude "21.76823" ; trt:gbifOccurrenceId "1455384319" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC, PMOC" ; dwc:eventDate "1980-01-14" ; dwc:locality "A" ; dwc:recordedBy "Rooiplato" ; dwc:typeStatus "holotype" ; dwc:verbatimLatitude "-31.433332" ; dwc:verbatimLongitude "19.4" ; trt:gbifOccurrenceId "1455384320" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1933-12-07" ; dwc:locality "Upington" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-28.45" ; dwc:verbatimLongitude "21.25" ; trt:gbifOccurrenceId "1455384302" ; trt:httpUri ; a dwc:MaterialCitation .