@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .

<http://treatment.plazi.org/id/03DD87EDFFDBFFAC1917FA05559AFF05>
    cito:cites <http://dx.doi.org/10.5281/zenodo.5392169>, <http://dx.doi.org/10.5281/zenodo.5392171>, <http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_Du_>, <http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_aeneoniger_Lebis_1960>, <http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_cambeforti_Montreuil_2005>, <http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_dorbignyi_Montreuil_2005>, <http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_niger_dOrb_>, <http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_pseudoniger_Montreuil_2005>, <http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_seminiger_Montreuil_>, <http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_seminiger_dOrbigny_1915>, <http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_semivirens_dOrbigny_1915>, <http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_steineri_Paulian_1991>, <http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_tamatavensis_Montreuil_2005>, <http://taxon-concept.plazi.org/id/Animalia/Insecta_Linnaeus_1758>, <http://taxon-concept.plazi.org/id/Plantae/Coleoptera_Linnaeus_1758> ;
    dc:creator "Montreuil, Olivier" ;
    dc:title "Helictopleurus niger d'Orbigny 1915" ;
    dwc:basisOfRecord <http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFDBFFA318D5F9AB5594F931>, <http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC1A88FDD25000FCAD>, <http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC1A88FBF250DDFBF5>, <http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC1A88FB305330FB50>, <http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC1A88FB6453C9FB6A>, <http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC1A88FB5B5099FA83>, <http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC183BFB5B5349FA9C>, <http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC182CFAB250B5FAB6>, <http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC182FFAA85334FACF>, <http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC1A88FAE653E2FA02> ;
    trt:augmentsTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_niger_dOrbigny_1915> ;
    trt:publishedIn <http://dx.doi.org/10.5281/zenodo.5392165> ;
    a trt:Treatment .

<http://dx.doi.org/10.5281/zenodo.5392165>
    bibo:endPage "135" ;
    bibo:issue "1" ;
    bibo:journal "Zoosystema" ;
    bibo:startPage "123" ;
    bibo:volume "27" ;
    dc:creator "Montreuil, Olivier" ;
    dc:date "2005" ;
    dc:title "Nouveaux Helictopleurus d’Orbigny, 1915 de Madagascar et révision du « groupe semivirens » sensu Lebis, 1960 (Insecta, Coleoptera, Scarabaeidae, Oniticellini)" ;
    fabio:hasPart <http://dx.doi.org/10.5281/zenodo.5392169>, <http://dx.doi.org/10.5281/zenodo.5392171> ;
    a fabio:JournalArticle .

<http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_niger_dOrbigny_1915>
    dwc:authority "d'Orbigny, 1915" ;
    dwc:authorityName "d'Orbigny" ;
    dwc:authorityYear "1915" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "dOrbigny, 1915" ;
    dwc:species "niger" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/3K9TF> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Helictopleurus_niger> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_niger_dOrb_>
    dwc:authority "d'Orb., Montreuil" ;
    dwc:authorityName "d'Orb., Montreuil" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "dOrb., Montreuil" ;
    dwc:species "niger" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Helictopleurus_niger> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_seminiger_dOrbigny_1915>
    dwc:authority "d'Orbigny, 1915", "d'Orbigny, 1915: 431" ;
    dwc:authorityName "d'Orbigny" ;
    dwc:authorityPageNumber "431" ;
    dwc:authorityYear "1915" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "dOrbigny, 1915" ;
    dwc:species "seminiger" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Helictopleurus_seminiger> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_seminiger_Montreuil_>
    dwc:authority "d'Orb. O. Montreuil" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "dOrb. Montreuil" ;
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    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_Du_>
    dwc:authority "du" ;
    dwc:authorityName "DU" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
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<http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_semivirens_dOrbigny_1915>
    dwc:authority "d'Orbigny, 1915", "d'Orbigny, 1915: 430" ;
    dwc:authorityName "d'Orbigny" ;
    dwc:authorityPageNumber "430" ;
    dwc:authorityYear "1915" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "dOrbigny, 1915" ;
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    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Helictopleurus_semivirens> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_aeneoniger_Lebis_1960>
    dwc:authority "Lebis, 1960" ;
    dwc:authorityName "Lebis, O. Montreuil" ;
    dwc:authorityYear "1960" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Lebis, Montreuil, 1960" ;
    dwc:species "aeneoniger" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Helictopleurus_aeneoniger> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_cambeforti_Montreuil_2005>
    dwc:authority "Montreuil, 2005" ;
    dwc:authorityName "Montreuil" ;
    dwc:authorityYear "2005" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Montreuil, 2005" ;
    dwc:species "cambeforti" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Helictopleurus_cambeforti> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_tamatavensis_Montreuil_2005>
    dwc:authority "Montreuil, 2005" ;
    dwc:authorityName "Montreuil" ;
    dwc:authorityYear "2005" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Montreuil, 2005" ;
    dwc:species "tamatavensis" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Helictopleurus_tamatavensis> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_pseudoniger_Montreuil_2005>
    dwc:authority "Montreuil, 2005" ;
    dwc:authorityName "Montreuil" ;
    dwc:authorityYear "2005" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Montreuil, 2005" ;
    dwc:species "pseudoniger" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Helictopleurus_pseudoniger> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_dorbignyi_Montreuil_2005>
    dwc:authority "Montreuil, 2005" ;
    dwc:authorityName "Montreuil" ;
    dwc:authorityYear "2005" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Montreuil, 2005" ;
    dwc:species "dorbignyi" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Helictopleurus_dorbignyi> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Helictopleurus_steineri_Paulian_1991>
    dwc:authority "Paulian & Cambefort, 1991" ;
    dwc:authorityName "Paulian & Cambefort" ;
    dwc:authorityYear "1991" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Paulian & Cambefort, 1991" ;
    dwc:species "steineri" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Helictopleurus_steineri> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Insecta_Linnaeus_1758>
    dwc:authorityName "Linnaeus" ;
    dwc:authorityYear "1758" ;
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    dwc:kingdom "Animalia" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "class" ;
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<http://taxon-concept.plazi.org/id/Plantae/Coleoptera_Linnaeus_1758>
    dwc:authorityName "Linnaeus" ;
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    dwc:order "Coleoptera" ;
    dwc:phylum "Tracheophyta" ;
    dwc:rank "order" ;
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<http://taxon-name.plazi.org/id/Animalia/Helictopleurus_niger>
    dwc:class "Insecta" ;
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    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
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    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Helictopleurus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Helictopleurus>
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Helictopleurus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
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    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Scarabaeidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Scarabaeidae>
    dwc:family "Scarabaeidae" ;
    dwc:rank "family" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Coleoptera> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Coleoptera>
    dwc:order "Coleoptera" ;
    dwc:rank "order" ;
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    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Insecta>
    dwc:class "Insecta" ;
    dwc:kingdom "Animalia" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "class" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Arthropoda> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Arthropoda>
    dwc:phylum "Arthropoda" ;
    dwc:rank "phylum" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia>
    dwc:kingdom "Animalia" ;
    dwc:rank "kingdom" ;
    a dwcFP:TaxonName .

<http://dx.doi.org/10.5281/zenodo.5392169>
    dc:description "FIG. 2. — Édéage des Helictopleurus du « groupe semivirens », paramère gauche, vue dorsale et paramère droit; A, H. dorbignyi n. sp.; B, H. tamatavensis n. sp.; C, H. cambeforti n. sp.; D, H. pseudoniger n. sp.; E, H. niger d’Orbigny, 1915; F, H. seminiger d’Orbigny, 1915; G, H. aeneoniger Lebis, 1960; H, H. semivirens d’Orbigny, 1915; I, H. steineri Paulian & Cambefort, 1991. Échelle: A-H, 1 mm; I, 0,5 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/5392169/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.5392171>
    dc:description "FIG. 3. — Tête des Helictopleurus du « groupe semivirens », mâle, vue frontale; A, H. dorbignyi n. sp.; B, H. tamatavensis n. sp.; C, H. cambeforti n. sp.; D, H. pseudoniger n. sp.; E, H. niger d’Orbigny, 1915; F, H. seminiger d’Orbigny, 1915; G, H. aeneoniger" ;
    fabio:hasRepresentation <https://zenodo.org/record/5392171/files/figure.png> ;
    a fabio:Figure .

<http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFDBFFA318D5F9AB5594F931>
    dwc:collectionCode "MATERIEL, TYPE" ;
    dwc:locality "Madagascar" ;
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<http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC1A88FDD25000FCAD>
    dwc:collectionCode "MATERIEL, TYPE, MNHN" ;
    dwc:locality "Madagascar" ;
    dwc:typeStatus "lectotype" ;
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<http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC1A88FBF250DDFBF5>
    dwc:collectionCode "MATERIEL, TYPE, MNHN" ;
    dwc:locality "Tananarive" ;
    dwc:typeStatus "holotype" ;
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<http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC1A88FB305330FB50>
    dwc:collectionCode "AUTRE, MATERIEL, EXAMINE" ;
    dwc:eventDate "1990-03-null" ;
    dwc:locality "Prov." ;
    dwc:recordedBy "W. E. Steiner" ;
    dwc:verbatimElevation "1000" ;
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<http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC1A88FB6453C9FB6A>
    dwc:eventDate "1955-03" ;
    dwc:locality "Beanana" ;
    dwc:recordedBy "J. Vadon" ;
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<http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC1A88FB5B5099FA83>
    dwc:locality "Perinet" ;
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<http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC183BFB5B5349FA9C>
    dwc:eventDate "1972-11" ;
    dwc:locality "Perinet" ;
    dwc:recordedBy "A. Peyrieras" ;
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<http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC182CFAB250B5FAB6>
    dwc:eventDate "1951-01" ;
    dwc:locality "Mandritsara" ;
    dwc:recordedBy "Michel" ;
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<http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC182FFAA85334FACF>
    dwc:eventDate "1951-03" ;
    dwc:locality "Ambohitsitondroma" ;
    dwc:recordedBy "Michel" ;
    trt:httpUri <https://treatment.plazi.org/id/03DD87EDFFDBFFAC1917FA05559AFF05#3B1C3CA6FFD4FFAC182FFAA85334FACF> ;
    a dwc:MaterialCitation .

<http://tb.plazi.org/GgServer/dwcaRecords/03DD87EDFFDBFFAC1917FA05559AFF05.mc.3B1C3CA6FFD4FFAC1A88FAE653E2FA02>
    dwc:collectionCode "MNHN" ;
    dwc:eventDate "1951-11" ;
    dwc:locality "Ambohitsitondroma" ;
    dwc:recordedBy "Vadon" ;
    trt:httpUri <https://treatment.plazi.org/id/03DD87EDFFDBFFAC1917FA05559AFF05#3B1C3CA6FFD4FFAC1A88FAE653E2FA02> ;
    a dwc:MaterialCitation .