@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , ; dc:creator "Duarte, Paulo R. M.; Grossi, Paschoal C.; Dupuis, Fabien" ; dc:title "Bothynus spinophallicus Duarte & Grossi & Dupuis 2022, new combination" ; dwc:basisOfRecord , , ; trt:definesTaxonConcept ; trt:deprecates ; trt:publishedIn ; a trt:Treatment . bibo:endPage "66" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:pubDate "2022-01-25" ; bibo:startPage "49" ; bibo:volume "5093" ; dc:creator "Duarte, Paulo R. M.; Grossi, Paschoal C.; Dupuis, Fabien" ; dc:date "2022" ; dc:title "A taxonomic revision of the Bothynus villiersi Endrödi, 1968 species group (Coleoptera: Scarabaeidae: Dynastinae)" ; fabio:hasPart , , , ; a fabio:JournalArticle . dwc:authority "(Ratcliffe & Cave, 2021)" ; dwc:authorityName "Duarte & Grossi & Dupuis" ; dwc:authorityYear "2022" ; dwc:baseAuthorityName "Ratcliffe & Cave" ; dwc:baseAuthorityYear "2021" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Bothynus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Ratcliffe & Cave, 2021) Duarte & Grossi & Dupuis, 2022" ; dwc:species "spinophallicus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ratcliffe & Cave 2021: 45" ; dwc:authorityName "Ratcliffe & Cave" ; dwc:authorityPageNumber "45" ; dwc:authorityYear "2021" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Coelosis" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ratcliffe & Cave, 2021" ; dwc:species "spinophallicus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Bothynus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "spinophallicus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Bothynus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Coelosis" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "spinophallicus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Coelosis" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record." ; fabio:hasRepresentation ; a fabio:Figure . dwc:collectionCode "UNSM" ; dwc:locality "Paraguay" ; dwc:recordedBy "R. C. Aguilar & Aguilar J." ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3455072332" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CERPE" ; dwc:eventDate "2011-05" ; dwc:locality "Parque Estadual do Cerrado" ; dwc:recordedBy "C. S. Costa & Equipe" ; trt:gbifOccurrenceId "3455072345" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Capiibary" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3455072361" ; trt:httpUri ; a dwc:MaterialCitation .