@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .

<http://treatment.plazi.org/id/05B3683E83175DC39567AA5195E1784D>
    cito:cites <http://dx.doi.org/10.3897/zookeys.1075.67459.figure1>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure2>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure3>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure4>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure5>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure6>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure7>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure8>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure12>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure11>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure13>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure9>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure10>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure14> ;
    dc:creator "Moreno-Gonzalez 1, Jairo A.; Pinto-da-Rocha 1, Ricardo; Gallao 2, Jonas E." ;
    dc:title "Tityus spelaeus Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2 2021, sp. nov." ;
    trt:definesTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Tityus_spelaeus_1_1> ;
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<http://dx.doi.org/10.3897/zookeys.1075.67459>
    bibo:endPage "75" ;
    bibo:journal "ZooKeys" ;
    bibo:pubDate "2021-12-07" ;
    bibo:startPage "33" ;
    bibo:volume "1075" ;
    dc:creator "Moreno-Gonzalez 1, Jairo A.; Pinto-da-Rocha 1, Ricardo; Gallao 2, Jonas E." ;
    dc:date "2021" ;
    dc:title "Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species" ;
    fabio:hasPart <http://dx.doi.org/10.3897/zookeys.1075.67459.figure1>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure2>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure3>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure4>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure5>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure6>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure7>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure8>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure12>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure11>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure13>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure9>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure10>, <http://dx.doi.org/10.3897/zookeys.1075.67459.figure14> ;
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<http://taxon-concept.plazi.org/id/Animalia/Tityus_spelaeus_1_1>
    dwc:LSID "http://zoobank.org/3AE5D4E6-C2F1-47A7-9768-046B09B2FF48" ;
    dwc:authority "Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2, 2021" ;
    dwc:authorityName "Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" ;
    dwc:authorityYear "2021" ;
    dwc:class "Insecta" ;
    dwc:family "Buthidae" ;
    dwc:genus "Tityus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Scorpiones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Moreno-González 1, Pinto-da-Rocha 1 & Gallão 2, 2021" ;
    dwc:species "spelaeus" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Tityus_spelaeus> ;
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<http://taxon-name.plazi.org/id/Animalia/Tityus_spelaeus>
    dwc:class "Insecta" ;
    dwc:family "Buthidae" ;
    dwc:genus "Tityus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Scorpiones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:species "spelaeus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Tityus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Tityus>
    dwc:genus "Tityus" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Buthidae> ;
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<http://taxon-name.plazi.org/id/Animalia/Buthidae>
    dwc:family "Buthidae" ;
    dwc:rank "family" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Scorpiones> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Scorpiones>
    dwc:order "Scorpiones" ;
    dwc:rank "order" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Insecta> ;
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<http://taxon-name.plazi.org/id/Animalia/Insecta>
    dwc:class "Insecta" ;
    dwc:rank "class" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Arthropoda> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Arthropoda>
    dwc:phylum "Arthropoda" ;
    dwc:rank "phylum" ;
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<http://taxon-name.plazi.org/id/Animalia>
    dwc:kingdom "Animalia" ;
    dwc:rank "kingdom" ;
    a dwcFP:TaxonName .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure1>
    dc:description "Figure 1. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, and Cytochrome c Oxidase I- COI). Maximum likelihood tree (Log-likelihood = - 12896.086), showing species-groups and subgenera. Values on nodes correspond to ultrafast-bootstrap (Ubst) values." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619807> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure2>
    dc:description "Figure 2. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, and Cytochrome c Oxidase I- COI), showing the distribution of the characters states of the ventral setae of telotarsi I-IV (orange: an irregularly distributed tuft of setae (type I); turquoise: two ventro-submedian rows of setae (type II )) across different Tityus subgenera and species-groups. Boxes on branches and associated values correspond to ultrafast-bootstrap (Ubst) values. Observations = Tityus (Tityus) spelaeus sp. nov. is marked in bold." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619808> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure3>
    dc:description "Figure 3. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, Cytochrome c Oxidase I), showing distribution of the characters states exhibited by the female basal pectinal piece (orange: GR absent; grey: medium-sized GR; turquoise: relatively large GR; blue: very large GR) across different Tityus subgenera and species-groups. Boxes on branches and associated values correspond to ultrafast-bootstrap (Ubst) values. Observations = Tityus (Tityus) spelaeus sp. nov. is marked in bold. Abbreviations = GR, glandular region." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619809> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure4>
    dc:description "Figure 4. Tityus C. L. Koch, 1836, telotarsi IV, showing ventral macrosetae A, C, E, G, I, K white light B, D, F, H, J, L UV light A, B Tityus (Tityus) brazilae Lourenco & Eickstedt, 1984 (type II) (MZSP 75619) C, D Tityus (Archaeotityus) clathratus C. L. Koch, 1844 (type I) (MZSP 31468) E, F Tityus (Atreus) forcipula (Gervais, 1843) (type II) (MZSP) G, H Tityus (Atreus) obscurus Gervais, 1843 (type I) (MNRJ 07610) I, J Tityus (Tityus) serrulatus Lutz & Mello, 1922 (type II) (MZSP 28205) K, L Tityus (Tityus) spelaeus sp. nov. (MZSP 74633) (type II). Observations = telotarsi I-IV ventral setae distribution: Type I = tuft of irregularly distributed setae. Type II = two discrete ventrosubmedian rows of setae. Scale bars: 500 μm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619810> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure5>
    dc:description "Figure 5. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Tityus) brazilae Lourenco & Eickstedt, 1984 (MZSP 75619) C, D Tityus (Archaeotityus) clathratus C. L. Koch, 1844 (MZSP 31468) E, F Tityus (Atreus) forcipula (Gervais, 1843) (MZSP). Abbreviations: BML, basal middle lamellae; GR, glandular region. Scale bars: 500 μm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619811> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure6>
    dc:description "Figure 6. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Atreus) obscurus Gervais, 1843 (MNRJ 07610) C, D Tityus (Tityus) serrulatus Lutz & Mello, 1922 (MZSP 28205) E, F Tityus (Tityus) spelaeus sp. nov. (MZSP 74633). Abbreviations: BML, basal middle lamellae; GR, glandular region of the pectinal basal piece. Scale bars: 500 μm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619812> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure7>
    dc:description "Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619813> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure8>
    dc:description "Figure 8. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), carapace, dorsal view A white light B UV light. Scale bars: 2 mm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619814> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure12>
    dc:description "Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D)." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619818> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure11>
    dc:description "Figure 11. Tityus (Tityus) spelaeus sp. nov., female paratype (MZSP 74633) genital area and pectines A genital operculum and pectinal basal piece B closeup of the pectinal basal piece, showing cuticular pores on glandular region C right pectine D peg sensillae, distribution E closeup of a peg sensilla. Scale bars: 800 µm (A); 60 µm (B); 1000 µm (C); 20 µm (D); 3 µm (E)." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619817> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure13>
    dc:description "Figure 13. Tityus (Tityus) spelaeus sp. nov., female paratypes under laboratory conditions A, B female paratype with scorpionlings A feeding upon scorpionlings B litter on female's back C specimens feeding on a cricket." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619819> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure9>
    dc:description "Figure 9. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), right pedipalp segments A femur, dorsal view B, C patella B dorsal view C external view. Scale bars: 1.5 mm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619815> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure10>
    dc:description "Figure 10. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), chela A external view B dorsal view C ventral view. Scale bar: 3 mm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619816> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1075.67459.figure14>
    dc:description "Figure 14. Habitat of Tityus (Tityus) spelaeus sp. nov. in the Russao cave A inside landscape of the cave B females on the cave walls." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/619820> ;
    a fabio:Figure .