@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .
@prefix xlink: <http://www.w3.org/1999/xlink/> .

<http://treatment.plazi.org/id/90E91BD08620595391E3513E8E367030>
    cito:cites <http://dx.doi.org/10.3897/zookeys.1207.117225.figure3>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure6>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure16>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure18>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure24>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure26>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure8>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure17>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure21>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure29> ;
    dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
    dc:title "Aphodius (Liothorax) isikdagensis" ;
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<http://dx.doi.org/10.3897/zookeys.1207.117225>
    bibo:endPage "299" ;
    bibo:journal "ZooKeys" ;
    bibo:pubDate "2024-07-22" ;
    bibo:startPage "205" ;
    bibo:volume "1207" ;
    dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
    dc:date "2024" ;
    dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
    fabio:hasPart <http://dx.doi.org/10.3897/zookeys.1207.117225.figure3>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure6>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure16>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure18>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure24>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure26>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure8>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure17>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure21>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure29> ;
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<http://taxon-concept.plazi.org/id/Animalia/Aphodius_isikdagensis_Balthasar_1952>
    dwc:authority "(Balthasar, 1952)", "Balthasar, 1952: 22" ;
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    dwc:authorityPageNumber "22" ;
    dwc:authorityYear "1952" ;
    dwc:baseAuthorityName "Balthasar" ;
    dwc:baseAuthorityYear "1952" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Aphodius" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Balthasar, 1952)", "Balthasar, 1952" ;
    dwc:species "isikdagensis" ;
    dwc:subGenus "Liothorax", "Ataeniomorphus" ;
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<http://taxon-name.plazi.org/id/Animalia/Aphodius_isikdagensis>
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<http://taxon-name.plazi.org/id/Animalia/Aphodius_Liothorax>
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<http://taxon-name.plazi.org/id/Animalia/Aphodius_Ataeniomorphus>
    dwc:class "Insecta" ;
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<http://dx.doi.org/10.3897/zookeys.1207.117225.figure3>
    dc:description "Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/1097461> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1207.117225.figure6>
    dc:description "Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/1097465> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1207.117225.figure16>
    dc:description "Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/1097478> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1207.117225.figure18>
    dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/1097482> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1207.117225.figure24>
    dc:description "Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/1097494> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1207.117225.figure26>
    dc:description "Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/1097499> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1207.117225.figure8>
    dc:description "Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/1097467> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1207.117225.figure17>
    dc:description "Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/1097480> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1207.117225.figure21>
    dc:description "Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/1097488> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.1207.117225.figure29>
    dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/1097509> ;
    a fabio:Figure .

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    dwc:locality "Isik" ;
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