Leiurus haenggii Lowe & Yağmur & Kovařík 2014, sp. n.
cites: zenodo.6505721
cites: zenodo.6505745
cites: zenodo.6505747
cites: zenodo.6505806
cites: zenodo.6505808
cites: zenodo.6505810
cites: zenodo.6505812
cites: zenodo.6505814
cites: zenodo.6505816
cites: zenodo.6505818
cites: zenodo.6505820
cites: zenodo.6505822
creator: Lowe, Graeme; Yağmur, Ersen Aydın; Kovařík, František
cites taxon name: Leiurus quinquestriatus hebraeus
defines taxon concept: Leiurus haenggii Lowe 2014
deprecates: Buthus quinquestriatus Pocock 1895
deprecates: Buthus quinquestriatus brachycentrus Birula 1937
deprecates: Buthus quinquestriatus Whittick 1941
deprecates: Leiurus quinquestriatus voelschowi Kovarik 1997
deprecates: Leiurus quinquestriatus Stahnke 1972
published in: A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula
type: Treatment

A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula
end page: 129
journal: Euscorpius
pub date: 2014-12-31
start page: 1
volume: 191
creator: Lowe, Graeme; Yağmur, Ersen Aydın; Kovařík, František
date: 2014
has part: zenodo.6505721
has part: zenodo.6505745
has part: zenodo.6505747
has part: zenodo.6505806
has part: zenodo.6505808
has part: zenodo.6505810
has part: zenodo.6505812
has part: zenodo.6505814
has part: zenodo.6505816
has part: zenodo.6505818
has part: zenodo.6505820
has part: zenodo.6505822
type: JournalArticle

Leiurus haenggii Lowe 2014
authority: Lowe & Yağmur & Kovařík, 2014
authority name: Lowe & Yağmur & Kovařík
authority year: 2014
class: Arachnida
family: Buthidae
genus: Leiurus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: species
scientific name authorship: Lowe & Yağmur & Kovařík, 2014
species: haenggii
has taxon name: Leiurus haenggii
type: TaxonConcept

Buthus quinquestriatus Pocock 1895
authority: : Pocock, 1895: 292
authority name: Pocock
authority page number: 292
authority year: 1895
class: Arachnida
family: Buthidae
genus: Buthus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: species
scientific name authorship: Pocock, 1895
species: quinquestriatus
has taxon name: Buthus quinquestriatus
type: TaxonConcept

Buthus quinquestriatus brachycentrus Birula 1937
authority: : Birula 1937: 102 - 105
authority name: Birula
authority page number: 102 - 105
authority year: 1937
class: Arachnida
family: Buthidae
genus: Buthus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: subSpecies
scientific name authorship: Birula, 1937
species: quinquestriatus
sub species: brachycentrus
has taxon name: Buthus quinquestriatus brachycentrus
type: TaxonConcept

Buthus quinquestriatus Whittick 1941
authority: : Whittick, 1941: 43
authority name: Whittick
authority page number: 43
authority year: 1941
class: Arachnida
family: Buthidae
genus: Buthus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: species
scientific name authorship: Whittick, 1941
species: quinquestriatus
sub genus: Buthus
has taxon name: Buthus quinquestriatus
type: TaxonConcept

Leiurus quinquestriatus voelschowi Kovarik 1997
authority: : Kovarik, 1997: 180
authority name: Kovarik
authority page number: 180
authority year: 1997
class: Arachnida
family: Buthidae
genus: Leiurus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: subSpecies
scientific name authorship: Kovarik, 1997
species: quinquestriatus
sub species: voelschowi
has taxon name: Leiurus quinquestriatus voelschowi
type: TaxonConcept

Leiurus quinquestriatus Stahnke 1972
authority: : Stahnke, 1972: 130
authority name: : Stahnke
authority page number: 130
authority year: 1972
class: Arachnida
family: Buthidae
genus: Leiurus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: species
scientific name authorship: Stahnke, 1972
species: quinquestriatus
has taxon name: Leiurus quinquestriatus
type: TaxonConcept

Leiurus haenggii
class: Arachnida
family: Buthidae
genus: Leiurus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: species
species: haenggii
has parent name: Leiurus
type: TaxonName

Leiurus
class: Arachnida
family: Buthidae
genus: Leiurus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: genus
has parent name: Buthidae
type: TaxonName

Buthidae
class: Arachnida
family: Buthidae
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: family
has parent name: Scorpiones
type: TaxonName

Scorpiones
class: Arachnida
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: order
has parent name: Arachnida
type: TaxonName

Arachnida
class: Arachnida
kingdom: Animalia
phylum: Arthropoda
rank: class
has parent name: Arthropoda
type: TaxonName

Arthropoda
kingdom: Animalia
phylum: Arthropoda
rank: phylum
has parent name: Animalia
type: TaxonName

Animalia
kingdom: Animalia
rank: kingdom
type: TaxonName

Buthus quinquestriatus
class: Arachnida
family: Buthidae
genus: Buthus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: species
species: quinquestriatus
sub genus: Buthus
has parent name: Buthus
has parent name: Buthus Buthus
type: TaxonName

Buthus
class: Arachnida
family: Buthidae
genus: Buthus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: genus
has parent name: Buthidae
type: TaxonName

Buthus quinquestriatus brachycentrus
class: Arachnida
family: Buthidae
genus: Buthus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: subSpecies
species: quinquestriatus
sub species: brachycentrus
has parent name: Buthus quinquestriatus
type: TaxonName

Buthus Buthus
class: Arachnida
family: Buthidae
genus: Buthus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: subGenus
sub genus: Buthus
has parent name: Buthus
type: TaxonName

Leiurus quinquestriatus voelschowi
class: Arachnida
family: Buthidae
genus: Leiurus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: subSpecies
species: quinquestriatus
sub species: voelschowi
has parent name: Leiurus quinquestriatus
type: TaxonName

Leiurus quinquestriatus
class: Arachnida
family: Buthidae
genus: Leiurus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: species
species: quinquestriatus
has parent name: Leiurus
type: TaxonName

Leiurus quinquestriatus hebraeus
class: Arachnida
family: Buthidae
genus: Leiurus
kingdom: Animalia
order: Scorpiones
phylum: Arthropoda
rank: subSpecies
species: quinquestriatus
sub species: hebraeus
has parent name: Leiurus quinquestriatus
type: TaxonName

zenodo.6505721
description: Figure 46: Morphometric differentiation between female Leiurus arabicus sp. n. and L. haenggii sp. n. A. Scatter plot of the compound morphometric ratio (slenderness factor) Fs = a.b.c (where a = metasoma III L/W, b = pedipalp patella L/W, c = leg III patella L/D) vs. Carapace L (N = 30 L. arabicus, N = 43 L. haenggii). The distribution of ratios of the two species were overlapped for juveniles and young, and were increasingly separated in subadults and adults, as segments of L. arabicus become more slender by allometric growth. B, C, D. Log-log scatter plots of metasoma III L vs. W, pedipalp patella L vs. W, and leg III patella L vs. D, respectively. Linear regression was applied separately to data sets split into upper and lower size ranges for each species (red lines). Numbers show fitted slope ± standard deviation (SD), with SD ranges in brackets. Slopes were similar between L. arabicus and L. haenggii over the lower size range, and diverged over the upper size range. For all three segments, allometry was significantly diphasic in L. arabicus, and only weakly so, or monophasic, in L. haenggii.
type: Figure

zenodo.6505745
description: Figure 58: Hemispermatophores of new Leiurus species. A. L. macroctenus sp. n., paratype male, Duhai, Oman. B. L. heberti sp. n., holotype male, Wadi Andur, Oman. C. L. haenggii sp. n., paratype male, Jabal Qara, Oman. Distal portions hemispermatophores illustrated to show flagellum and lobes (right hemispermatophores, convex aspect). Scale bars: 1 mm.
type: Figure

zenodo.6505747
description: Figure 59: Chelicerae of Leiurus species. A, B. L. macroctenus sp. n., paratype male, Thumrait, Oman. Right chelicera, ventral (A) and dorsal (B) aspect. C, D. L. haenggii sp. n., paratype male, Jabal Qara, Oman. Right chelicera, ventral (C) and dorsal (D) aspect. E, F. L. brachycentrus (Ehrenberg, 1829), female, Al Mansuriah, Yemen. Right chelicera, ventral (E) and dorsal (F) aspect. G, H. L. heberti sp. n., holotype male, Wadi Andur, Oman. Right chelicera, ventral (G) and dorsal (H) aspect. Scale bar in A: 1 mm (also applies to B–H).
type: Figure

zenodo.6505806
description: Figure 87: Metasoma V and telson of Leiurus spp. Males, lateral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n.. C. L. brachycentrus (Ehrenberg, 1829) stat. n.. D. L. haenggii sp. n. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n.. G. L. macroctenus sp. n.. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm.
type: Figure

zenodo.6505808
description: Figure 88: Metasoma V and telson of Leiurus spp. Males, ventral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n.. C. L. brachycentrus (Ehrenberg, 1829) stat. n.. D. L. haenggii sp. n.. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n.. G. L. macroctenus sp. n.. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm.
type: Figure

zenodo.6505810
description: Figure 89: Metasoma V and telson of Leiurus spp. Females, lateral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n. C. L. brachycentrus (Ehrenberg, 1829) stat. n. D. L. haenggii sp. n. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n. G. L. macroctenus sp. n. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm.
type: Figure

zenodo.6505812
description: Figure 90: Metasoma V and telson of Leiurus spp. Females, ventral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n. C. L. brachycentrus (Ehrenberg, 1829) stat. n.. D. L. haenggii sp. n. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n. G. L. macroctenus sp. n. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm.
type: Figure

zenodo.6505814
description: Figure 91: Pigmentation patterns of metasoma IV, V of Leiurus species, ventral aspect. A. L. haenggii sp. n., holotype female, Ta’if, Saudi Arabia (NHMB 17k). B. L. macroctenus sp. n., holotype male, Thumrait, Oman. C. L. heberti sp. n., holotype male, Wadi Andur, Oman. D. L. quinquestriatus (Ehrenberg, 1828), female, Cairo, Egypt (NHMB 17d). E. L. brachycentrus (Ehrenberg, 1829) stat. n., male, Ad Darb, Saudi Arabia (NHMB 17ag). F. L. arabicus sp. n., holotype female, Kushm Dibi, Saudi Arabia (NHMB 17aq). G. L. hebraeus (Birula, 1908) stat. n., female, Israel (NHMB 17a). H. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, male, Eski Sarkaya Village, Turkey. Scale bars: 2 mm.
type: Figure

zenodo.6505816
description: Figure 92: Sternite III of Leiurus spp., female. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, Caybasi Village, Turkey. B. L. arabicus sp. n., holotype, Kushm Dibi, Saudi Arabia (NHMB 17aq). C. L. arabicus sp. n., paratype, Al Amar, Bahrain (NHMB 17bg). D. L. heberti sp. n., paratype, Jabal Samhan. Oman. E. L. haenggii sp. n., paratype, Wadi Turabah, Saudi Arabia (NHMB 17al). F. L. haenggii sp. n., paratype, Adnan, Saudi Arabia (NHMB 17am). G. L. brachycentrus (Ehrenberg, 1829) stat. n., Al Mansuriah, Yemen. H. L. macroctenus sp. n., paratype, Masirah Island, Oman. I. L. macroctenus sp. n., paratype, Thumrait, Oman. J. L. quinquestriatus (Ehrenberg, 1828), Kharga Oasis, Egypt (NHMB 17i). K. L. hebraeus (Birula, 1908) stat. n., Israel (NHMB 17a). L. L. jordanensis Lourenço, Modry et Amr, 2002, al-Tawil, Saudi Arabia.
type: Figure

zenodo.6505818
description: Figure 93: Medial intercarinal areas of tergite III of female Leiurus spp. A. L. arabicus sp. n., paratype, Jeddah, Saudi Arabia. B. L. arabicus sp. n., paratype, Wadi Khumrah, Saudi Arabia (NHMB 17bj). C. L. haenggii sp. n., paratype, Yemen. D. L. haenggii sp. n., paratype, Wadi Maraba, Saudi Arabia (NHMB 17al). E. L. heberti sp. n., paratype, Jabal Samhan. Oman. F. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, Caybasi Village, Turkey. G. L. brachycentrus (Ehrenberg, 1829) stat. n., Al Mansuriah, Yemen. H. L. macroctenus sp. n., paratype, Masirah Island, Oman. I. L. hebraeus (Birula, 1908) stat. n., Israel (NHMB 17a). J. L. hebraeus Birula, 1908, Kurayyima, Jordan. K. L. quinquestriatus (Ehrenberg, 1828), Kharga Oasis, Egypt (NHMB 17i). L. L. jordanensis Lourenço, Modry et Amr, 2002, al-Tawil, Saudi Arabia.
type: Figure

zenodo.6505820
description: Figure 94: Comparative analysis of granulation on tergite III of female Leiurus. A. UV fluorescence image of tergite III medial intercarinal surface of L. quinquestriatus (Egypt). B. Mask of granulation pattern on medial intercarinal surface from image in Figure 94A. C. Enlarged view of tergite III granulation in rectangular area indicated in Figure 94A. Granules were identified as local maxima of fluorescence intensity with contour shadows cast by directional UV illumination, and were modeled by elliptical regions-of-interest (ROIs). Parameters of granule ROIs were measured in ImageJ 1.44 (Rasband, 1997–2011). Area of granulometric analysis was restricted to surfaces bounded by the lateral and posterior marginal carinae, and a line passing through oblique, anterior transverse rows of enlarged granules (granules along carinae and bounding lines were omitted). D. Scatter plot of mean granule diameter vs. total granule area for 4 species of Leiurus (L. arabicus sp. n., L. haenggii sp. n., L. hebraeus stat. n. and L. quinquestriatus). Each point represents granulometric data derived from tergite III of one scorpion specimen (bilaterally, as shown in Figure 94B). Data were extracted from 18,423 ROIs from 36 scorpions. For each specimen, total granule area (a measure of density of granulation) was computed as sum of areas of all ROIs, and mean ROI diameter (a measure of coarseness of granulation) as the mean value of the maximum diameters of all granule ROIs. For comparative analysis, images from different size scorpions were resampled to equalize the distance between left and right posterior marginal granules of the lateral carinae (arbitrarily set to 4,000 units or pixels; linear dimensions expressed as [pixel], areas as [pixel2]). The total granule area separated the species into 2 groups: i.e. sparsely granulated (L. arabicus, L. haenggii) and densely granulated (L. hebraeus, L. quinquestriatus). In contrast, the distributions of ROI diameter were broadly overlapping…
type: Figure

zenodo.6505822
description: Figure 95: Variation in key diagnostic biometrics of adult Leiurus spp. A. Cumulative distributions of position of trichobothrium db relative to est on pedipalp fixed finger for: L. abdullahbayrami (53), L. brachycentrus stat. n. (11), L. hebraeus stat. n. (38), L. macroctenus sp. n. (176), L. quinquestriatus (61), L. haenggii sp. n. (158), L. arabicus sp. n. (87) (number of fingers measured in parentheses). Data for males and females were pooled. The db–est distances were normalized to pedipalp movable finger length (chord length from finger tip to external articular condyle), with positive values indicating db distal to est, negative values indicating db proximal to est. Distributions of L. haenggii (µ = 0.06367) and L. arabicus sp. n. (µ = 0.06671) were not significantly different (P = 0.405148, t test), indicating a close relationship between these species. Distribution of L. quinquestriatus (µ = 0.047313) was significantly different from those of L. haenggii and L. arabicus (P = 0.000102 and 0.000030 respectively). B. Scatter plot of morphometric ratios of mid-pectine sensillar margin L (MPSM, indicated in inset) to metasoma I W, and to carapace L, showing differences in relative pectinal tooth size in females of nine species of Leiurus. C. A subset of the female data in Figure 95B, plotted as cumulative distributions of morphometric ratio of mid-pectine sensillar margin L to metasoma I W. Distributions of L. haenggii (µ = 0.102268) and L. arabicus (µ = 0.10432) were not significantly different (P> 0.1, K-S test). D. Scatter plot of morphometric ratios of mid-pectine sensillar margin L to metasoma I W (relative size of pectine teeth), and metasoma II L/W (slenderness of metasoma II) for females of nine species of Leiurus. A, C, D: symbol key as in Figure 95B.
type: Figure

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collection code: NHMB
event date: 1975-10
locality: Ta'if
recorded by: W. Buttiker
type status: holotype
gbif occurrence id: 3913646530
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collection code: NHMB
event date: 1980-04-11
locality: Wadi Ghaiz
recorded by: J. N. Barnes
type status: paratype
verbatim elevation: 60
verbatim latitude: 16.966667
verbatim longitude: 53.866665
gbif occurrence id: 3913646528
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collection code: NHMB
locality: Sarfait Area
recorded by: Sultan of Oman Land Forces
type status: paratype
verbatim latitude: 16.666666
verbatim longitude: 53.1
gbif occurrence id: 3913646313
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collection code: NHMB
event date: 1989-09-04
locality: Jabal Qara
recorded by: R. P. Whitcomb
type status: paratype
verbatim elevation: 1300
verbatim latitude: 16.716667
verbatim longitude: 53.116665
gbif occurrence id: 3913646523
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collection code: ONHM
event date: 1989-09-06
locality: Nejd
recorded by: M. D. Gallagher
type status: paratype
verbatim elevation: 850
verbatim latitude: 17.316668
verbatim longitude: 54.266666
gbif occurrence id: 3913646377
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF42071196DFC6488F7D9E1F9B5
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collection code: NHMB
event date: 1993-10
locality: Salalah
recorded by: F. Barter
type status: paratype
verbatim latitude: 17.083334
verbatim longitude: 54.166668
gbif occurrence id: 3913646490
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collection code: GL
event date: 1993-10-16
locality: Nejd
recorded by: G. Lowe
type status: paratype
verbatim elevation: 800
verbatim latitude: 17.297167
verbatim longitude: 54.085167
gbif occurrence id: 3913646443
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF42071196DFAB88B13DE82F94B
type: MaterialCitation

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collection code: NHMB
event date: 1993-10-17
locality: Salalah- Thumrait road
recorded by: G. Lowe
sampling protocol: UV detection
type status: paratype
verbatim elevation: 800
verbatim latitude: 17.287666
verbatim longitude: 54.089333
gbif occurrence id: 3913646477
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF42071196DFC018B9ED859F8AF
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collection code: NHMB
event date: 1993-10-17
locality: Salalah-Thumrait road
recorded by: G. Lowe
sampling protocol: UV detection
type status: paratype
verbatim elevation: 800
verbatim latitude: 17.293
verbatim longitude: 54.082832
gbif occurrence id: 3913646351
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collection code: NHMB
event date: 1993-10-18
locality: main road above Khor Rori Beach
recorded by: G. Lowe
sampling protocol: UV detection on ground
type status: paratype
verbatim elevation: 50
verbatim latitude: 17.053667
verbatim longitude: 54.422165
gbif occurrence id: 3913646360
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF42076196AFEA38BB5DCDBF8B5
type: MaterialCitation

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collection code: GL
event date: 1993-10-18
locality: Ayn Hamran
recorded by: G. Lowe
type status: paratype
verbatim elevation: 50
verbatim latitude: 17.059
verbatim longitude: 54.273335
gbif occurrence id: 3913646399
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collection code: BMNH
event date: 1995-04-29
locality: Jabal Qara
recorded by: J. Dundon
type status: paratype
verbatim latitude: 17.0315
verbatim longitude: 54.091
gbif occurrence id: 3913646432
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event date: 1995-04-30
locality: Jabal Qara
recorded by: J. Dundon
type status: paratype
verbatim elevation: 142
verbatim latitude: 17.125668
verbatim longitude: 54.1495
gbif occurrence id: 3913646526
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collection code: ONHM
event date: 2001-12-24
locality: Khor Rori Beach
recorded by: A. Winkler
type status: paratype
verbatim elevation: 22
verbatim latitude: 17.053
verbatim longitude: 54.425167
gbif occurrence id: 3913646466
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event date: 2013-09
locality: Wadi Khor Rori
recorded by: Mazuch & Novak
type status: paratype
verbatim elevation: 22
verbatim latitude: 17.049915
verbatim longitude: 54.424557
gbif occurrence id: 3913646363
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collection code: FKCP
event date: 2014-03-21
locality: Taqah
recorded by: D. Hoferek
type status: paratype
verbatim elevation: 26
verbatim latitude: 17.054333
verbatim longitude: 54.425
gbif occurrence id: 3913646439
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collection code: FKCP
event date: 2014-03-22
locality: Mirbat
recorded by: D. Hoferek
type status: paratype
verbatim elevation: 54
verbatim latitude: 17.0365
verbatim longitude: 54.645832
gbif occurrence id: 3913646368
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collection code: NHMB
locality: Saudi Arabia
recorded by: W. Buttiker
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gbif occurrence id: 3913646469
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locality: Wadi Turabah
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1580
verbatim latitude: 20.476667
verbatim longitude: 41.1675
gbif occurrence id: 3913646525
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collection code: NHMB
locality: Wadi Wajj
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1810
verbatim latitude: 21.166666
verbatim longitude: 40.266666
gbif occurrence id: 3913646461
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collection code: NHMB
locality: Wadi Wajj
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1810
verbatim latitude: 21.166666
verbatim longitude: 40.266666
gbif occurrence id: 3913646361
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collection code: NHMB
event date: 1975-10
locality: Taif
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1670
verbatim latitude: 21.3
verbatim longitude: 40.4
gbif occurrence id: 3913646352
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF42077196BFAD38B9FD901F88D
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collection code: NHMB
event date: 1976-04-21
locality: Abha Gizan km 53
recorded by: W. Buttiker & W. Witt
verbatim elevation: 300
verbatim latitude: 17.85
verbatim longitude: 42.38
gbif occurrence id: 3913646442
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collection code: NHMB
event date: 1977-02-18
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 2050
verbatim latitude: 18.306667
verbatim longitude: 42.595
gbif occurrence id: 3913646450
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collection code: NHMB
event date: 1978-09-21
locality: Adnan
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1650
verbatim latitude: 20.433332
verbatim longitude: 41.516666
gbif occurrence id: 3913646487
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collection code: NHMB
event date: 1978-10-01
locality: Wadi Maraba
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 320
verbatim latitude: 17.896
verbatim longitude: 42.39
gbif occurrence id: 3913646527
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collection code: NHMB
event date: 1979-11-25
locality: Wadi Qatan
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1350
verbatim latitude: 18.114334
verbatim longitude: 44.1505
gbif occurrence id: 3913646471
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collection code: NHMB
event date: 1979-11-25
locality: Wadi Qatan
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1350
verbatim latitude: 18.114334
verbatim longitude: 44.1505
gbif occurrence id: 3913646336
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF420741968FE158B63DC20F918
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collection code: NHMB
event date: 1979-11-26
locality: Wadi Jaddah
recorded by: W. Buttiker
verbatim latitude: 25.866667
verbatim longitude: 48.8
gbif occurrence id: 3913646488
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collection code: NHMB
locality: Adama
recorded by: W. Buttiker
verbatim elevation: 1770
verbatim latitude: 19.338833
verbatim longitude: 42.022835
gbif occurrence id: 3913646486
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collection code: NHMB
event date: 1980-02-13
locality: Wadi Gaanah
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 470
verbatim latitude: 18.43
verbatim longitude: 41.893333
gbif occurrence id: 3913646389
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collection code: NHMB
event date: 1980-03-31
locality: Wadi Yamaniyah
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1100
verbatim latitude: 21.65
verbatim longitude: 40.316666
gbif occurrence id: 3913646331
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collection code: NHMB
event date: 1980-04-04
locality: Wadi Turabah
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1480
verbatim latitude: 20.483334
verbatim longitude: 41.2
gbif occurrence id: 3913646422
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collection code: NHMB
event date: 1980-04-17
locality: Adama
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1770
verbatim latitude: 19.338833
verbatim longitude: 42.022835
gbif occurrence id: 3913646383
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collection code: NHMB
event date: 1980-04-17
locality: Adama
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1770
verbatim latitude: 19.338833
verbatim longitude: 42.022835
gbif occurrence id: 3913646317
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collection code: NHMB
event date: 1980-04-21
locality: Wadi Shuqub
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1480
verbatim latitude: 20.716667
verbatim longitude: 41.166668
gbif occurrence id: 3913646420
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collection code: NHMB
event date: 1980-09-15
locality: Wadi Turabah
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1510
verbatim latitude: 20.483334
verbatim longitude: 41.2
gbif occurrence id: 3913646379
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collection code: NHMB
event date: 1980-09-23
locality: Wadi Qatan
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1350
verbatim latitude: 18.114334
verbatim longitude: 44.1505
gbif occurrence id: 3913646411
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collection code: NHMB
event date: 1980-04-21
locality: Wadi Sanakhah
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1320
verbatim latitude: 18.016666
verbatim longitude: 44.116665
gbif occurrence id: 3913646516
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collection code: NHMB
event date: 1980-10-07
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 2050
verbatim latitude: 19.9095
verbatim longitude: 41.566666
gbif occurrence id: 3913646497
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collection code: NHMB
event date: 1980-10-07
locality: Biljurshi
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1840
verbatim latitude: 19.834333
verbatim longitude: 41.54233
gbif occurrence id: 3913646546
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collection code: NHMB
event date: 1981-09-27
locality: Hesua
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1930
verbatim latitude: 18.083334
verbatim longitude: 42.35
gbif occurrence id: 3913646482
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF420751969FEDC8A23DE4CF8B8
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collection code: NHMB
event date: 1982-02-15
locality: Al Faraah
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 700
verbatim latitude: 20.95
verbatim longitude: 40.2
gbif occurrence id: 3913646452
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF420751969FCC38A0DDFA6F8FA
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collection code: NHMB
event date: 1982-04-23
locality: Wadi Wajj
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1810
verbatim latitude: 21.166666
verbatim longitude: 40.266666
gbif occurrence id: 3913646511
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF420751969FD278A4FD8C8F8A6
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collection code: NHMB
event date: 1983-01-07
locality: Wadi Maharish
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1000
verbatim latitude: 21.358334
verbatim longitude: 40.216667
gbif occurrence id: 3913646345
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF420751969FA468A23D8DFF8D9
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collection code: NHMB
event date: 1983-02-08
locality: Wadi Bani Malek
recorded by: W. Buttiker
type status: paratype
verbatim latitude: 21.574667
verbatim longitude: 39.28583
gbif occurrence id: 3913646400
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF420751966FCE88A4FDF9EF983
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collection code: NHMB
event date: 1983-09-09
locality: Wadi Ellah
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1480
verbatim latitude: 22.583334
verbatim longitude: 41.583332
gbif occurrence id: 3913646549
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collection code: NHMB
event date: 1983-09-10
locality: Wadi Asidah
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1560
verbatim latitude: 20.416666
verbatim longitude: 41.2
gbif occurrence id: 3913646302
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1966FD7C8B00DFE9F9F8
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collection code: NHMB
event date: 1983-09-15
locality: Wadi Tayyah
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 825
verbatim latitude: 18.533333
verbatim longitude: 42.233334
gbif occurrence id: 3913646394
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1966FD678B4DDF40F93B
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collection code: NHMB
event date: 1984-03-08
locality: Wadi Dhiyan
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1050
verbatim latitude: 19.908333
verbatim longitude: 41.506668
gbif occurrence id: 3913646460
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1966FDD98B8EDC8FF97D
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collection code: NHMB
event date: 1984-08-03
locality: Baha
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 2170
verbatim latitude: 20.010834
verbatim longitude: 41.466667
gbif occurrence id: 3913646518
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1966FE0A8BC8DF7FF8B0
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collection code: NHMB
event date: 1984-09-10
locality: Al Foqah
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1630
verbatim latitude: 19.833334
verbatim longitude: 41.85
gbif occurrence id: 3913646365
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1966FD868A35DCC8F8F3
type: MaterialCitation

http://tb.plazi.org/GgServer/dwcaRecords/038C87BF2071197FFC018C0BDED4FD2D.mc.3B4D3CF4207A1966FE478A76DEBDF9A4
collection code: NHMB
event date: 1984-10-20
locality: Al Foqah
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 1630
verbatim latitude: 19.983334
verbatim longitude: 41.85
gbif occurrence id: 3913646359
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1966FE478A76DEBDF9A4
type: MaterialCitation

http://tb.plazi.org/GgServer/dwcaRecords/038C87BF2071197FFC018C0BDED4FD2D.mc.3B4D3CF4207A1966FBC58B21DEE8F9E7
collection code: NHMB
event date: 1985-03-24
locality: Jebel Lebaba
recorded by: J. Grainger
type status: paratype
verbatim elevation: 420
verbatim latitude: 18.0275
verbatim longitude: 42.032166
gbif occurrence id: 3913646495
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1966FBC58B21DEE8F9E7
type: MaterialCitation

http://tb.plazi.org/GgServer/dwcaRecords/038C87BF2071197FFC018C0BDED4FD2D.mc.3B4D3CF4207A1966FC668B62D868F9F8
collection code: NHMB
event date: 1985-04-02
locality: Jebel Warjan
recorded by: S. Collenette
type status: paratype
verbatim elevation: 2100
verbatim latitude: 23.983334
verbatim longitude: 39.166668
gbif occurrence id: 3913646353
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1966FC668B62D868F9F8
type: MaterialCitation

http://tb.plazi.org/GgServer/dwcaRecords/038C87BF2071197FFC018C0BDED4FD2D.mc.3B4D3CF4207A1966FAE68B4DD900F93B
collection code: NHMB
event date: 1985-04-25
locality: Fare
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 850
verbatim latitude: 22.75
verbatim longitude: 39.806667
gbif occurrence id: 3913646428
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1966FAE68B4DD900F93B
type: MaterialCitation

http://tb.plazi.org/GgServer/dwcaRecords/038C87BF2071197FFC018C0BDED4FD2D.mc.3B4D3CF4207A1966FB9F8B8EDEB2F97D
collection code: NHMB
event date: 1985-06-05
locality: Jebel Al Lawz
recorded by: J. Grainger
type status: paratype
verbatim elevation: 2200
verbatim latitude: 28.692
verbatim longitude: 35.298
gbif occurrence id: 3913646330
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1966FB9F8B8EDEB2F97D
type: MaterialCitation

http://tb.plazi.org/GgServer/dwcaRecords/038C87BF2071197FFC018C0BDED4FD2D.mc.3B4D3CF4207A1966FC2B8BC8DEC4F8B0
collection code: NHMB
event date: 1986-11-12
locality: Shumaisy
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 740
verbatim latitude: 25.1
verbatim longitude: 38.716667
gbif occurrence id: 3913646467
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1966FC2B8BC8DEC4F8B0
type: MaterialCitation

http://tb.plazi.org/GgServer/dwcaRecords/038C87BF2071197FFC018C0BDED4FD2D.mc.3B4D3CF4207A1966FC458A35DEFFF8D1
collection code: GL
locality: Sanaa
recorded by: Heule
type status: paratype
gbif occurrence id: 3913646326
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1966FC458A35DEFFF8D1
type: MaterialCitation

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collection code: NHMB
event date: 1985-09-27
locality: Jebel Bagim
recorded by: W. Buttiker
type status: paratype
verbatim elevation: 2000
verbatim latitude: 16.925
verbatim longitude: 43.783333
gbif occurrence id: 3913646337
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207A1967FC1B8A14DC45F983
type: MaterialCitation

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collection code: FKCP
locality: Wadi Dhahr
recorded by: Stastny
type status: paratype
verbatim latitude: 15.408667
verbatim longitude: 44.123165
gbif occurrence id: 3913646354
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207B1967FECE88C6DF91F9A4
type: MaterialCitation

http://tb.plazi.org/GgServer/dwcaRecords/038C87BF2071197FFC018C0BDED4FD2D.mc.3B4D3CF4207B1967FD308B21DC1BF9E7
collection code: GL
event date: 2003-11-17
locality: Wadithar
recorded by: P. Kabatek
type status: paratype
verbatim latitude: 14.443334
verbatim longitude: 44.758335
gbif occurrence id: 3913646529
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207B1967FD308B21DC1BF9E7
type: MaterialCitation

http://tb.plazi.org/GgServer/dwcaRecords/038C87BF2071197FFC018C0BDED4FD2D.mc.3B4D3CF4207B1967FEBE8B62DE47F9F8
collection code: FKCP
locality: Jabal Lawz
recorded by: P. Kabatek
type status: paratype
verbatim elevation: 2828
verbatim latitude: 15.383333
verbatim longitude: 44.483334
gbif occurrence id: 3913646392
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207B1967FEBE8B62DE47F9F8
type: MaterialCitation

http://tb.plazi.org/GgServer/dwcaRecords/038C87BF2071197FFC018C0BDED4FD2D.mc.3B4D3CF4207B1967FCC48B4DDC58F97D
collection code: FKCP
event date: 2007-11-04
locality: Locality No.
recorded by: D. Kral
type status: paratype
verbatim elevation: 1968
verbatim latitude: 14.1405
verbatim longitude: 45.43133
gbif occurrence id: 3913646426
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207B1967FCC48B4DDC58F97D
type: MaterialCitation

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collection code: FKCP
type status: paratype
gbif occurrence id: 3913646309
http uri: https://treatment.plazi.org/id/038C87BF2071197FFC018C0BDED4FD2D#3B4D3CF4207B1967FEF58BC8DCE9F97D
type: MaterialCitation