@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , , ; dc:creator "Mateos, Eduardo; Riutort, Marta; Álvarez-Presas, Marta" ; dc:title "Caenoplana variegata" ; dwc:basisOfRecord , ; trt:augmentsTaxonConcept ; trt:deprecates , , , , , , , , , , , , , , , , , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "222" ; bibo:issue "2" ; bibo:journal "Zootaxa" ; bibo:pubDate "2020-02-05" ; bibo:startPage "193" ; bibo:volume "4731" ; dc:creator "Mateos, Eduardo; Riutort, Marta; Álvarez-Presas, Marta" ; dc:date "2020" ; dc:title "The identity of the invasive yellow-striped terrestrial planarian found recently in Europe: Caenoplana variegata (Fletcher & Hamilton, 1888) or Caenoplana bicolor (Graff, 1899)?" ; fabio:hasPart , , , , , , , ; a fabio:JournalArticle . dwc:authority "(Fletcher & Hamilton, 1888)" ; dwc:baseAuthorityName "Fletcher & Hamilton" ; dwc:baseAuthorityYear "1888" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Caenoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Fletcher & Hamilton, 1888)" ; dwc:species "variegata" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Fletcher & Hamilton, 1888" ; dwc:authorityName "Fletcher & Hamilton" ; dwc:authorityYear "1888" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Fletcher & Hamilton, 1888" ; dwc:species "variegata" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Dendy, 1892" ; dwc:authorityName "Dendy" ; dwc:authorityYear "1892" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Dendy, 1892" ; dwc:species "variegata" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Dendy, 1893 a" ; dwc:authorityName "Dendy" ; dwc:authorityYear "1893" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Dendy, 1893" ; dwc:species "variegata" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Dendy, 1894" ; dwc:authorityName "Dendy" ; dwc:authorityYear "1894" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Dendy, 1894" ; dwc:species "variegata" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Graff, 1899" ; dwc:authorityName "Graff" ; dwc:authorityYear "1899" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Graff, 1899" ; dwc:species "bicolor" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Graff, 1899" ; dwc:authorityName "Graff" ; dwc:authorityYear "1899" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Graff, 1899" ; dwc:species "variegata" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Steel, 1900" ; dwc:authorityName "Steel" ; dwc:authorityYear "1900" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Steel, 1900" ; dwc:species "variegata" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Winsor, 1973" ; dwc:authorityName "Winsor" ; dwc:authorityYear "1973" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Winsor, 1973" ; dwc:species "varigata" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Winsor, 1973" ; dwc:authorityName "Winsor" ; dwc:authorityYear "1973" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Winsor, 1973" ; dwc:species "bicolor" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Winsor, 1977" ; dwc:authorityName "Winsor" ; dwc:authorityYear "1977" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Winsor, 1977" ; dwc:species "varigata" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ogren & Kawakatsu, 1991" ; dwc:authorityName "Ogren & Kawakatsu" ; dwc:authorityYear "1991" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Australopacifica" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ogren & Kawakatsu, 1991" ; dwc:species "bicolor" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ogren & Kawakatsu, 1991" ; dwc:authorityName "Ogren & Kawakatsu" ; dwc:authorityYear "1991" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Australopacifica" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ogren & Kawakatsu, 1991" ; dwc:species "variegata" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Winsor, 1991" ; dwc:authorityName "Winsor" ; dwc:authorityYear "1991" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Caenoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Winsor, 1991" ; dwc:species "bicolor" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Winsor, 1991" ; dwc:authorityName "Winsor" ; dwc:authorityYear "1991" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Caenoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Winsor, 1991" ; dwc:species "variegata" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Winsor, 1997" ; dwc:authorityName "Winsor" ; dwc:authorityYear "1997" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Caenoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Winsor, 1997" ; dwc:species "bicolor" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Alvarez-Presas et al. 2014" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Caenoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Alvarez-Presas et al., 2014" ; dwc:species "bicolor" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "De Waart, 2016" ; dwc:authorityName "De Waart" ; dwc:authorityYear "2016" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Caenoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "De Waart, 2016" ; dwc:species "bicolor" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Sluys, 2016" ; dwc:authorityName "Sluys" ; dwc:authorityYear "2016" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Caenoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Sluys, 2016" ; dwc:species "bicolor" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Vardinoyannis & Alexandrakis, 2019" ; dwc:authorityName "Vardinoyannis & Alexandrakis" ; dwc:authorityYear "2019" ; dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Caenoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Vardinoyannis & Alexandrakis, 2019" ; dwc:species "bicolor" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Caenoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:species "variegata" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Caenoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Rhabditophora" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Rhabditophora" ; dwc:kingdom "Animalia" ; dwc:phylum "Platyhelminthes" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Platyhelminthes" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:species "variegata" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:species "bicolor" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Geoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:species "varigata" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Australopacifica" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:species "bicolor" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Australopacifica" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Australopacifica" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:species "variegata" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Rhabditophora" ; dwc:family "Geoplanidae" ; dwc:genus "Caenoplana" ; dwc:kingdom "Animalia" ; dwc:order "Seriata" ; dwc:phylum "Platyhelminthes" ; dwc:rank "species" ; dwc:species "bicolor" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURE 5. Caenoplana variegata. Photographs of remaining preserved portions of the sectioned specimens, taken 10 August 2018. NHMUK97: dorsal (a) and ventral (b) surfaces, preserved in 1897, now in 80% IMS. Cv7: dorsal (c) and ventral (d) surfaces, killed and preserved 29 January 2015, now in 100% ethanol. 1 mm graph paper background, scale bar 5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 6. Caenoplana variegata. NHMUK97: a, mid-dorsal and, b, mid-ventral TS showing the epithelium and cutaneous muscle, scales 50 µm; c, TS enlargement of dorsal portion of (d), scale 200 µm; d, whole TS, scale 500 µm. Note in (c) lack of sub-cutaneous melanic pigment in regions corresponding with the dark stripes of living animals, cf. Figs 8 & 10; e, LS of the pharynx, * indicates the position of the pharyngeal aperture, scale 2 mm (anterior to the right)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 7. Caenoplana variegata. NHMUK97: a, LS showing several mature testes, scale 200 µm; b, TS showing the ventral nerve cord, ovovitelline duct and testis on one side, scale 100 µm; c, LS showing the immature ovovitelline duct immediately dorsal to the ventral nerve cord, scale 200 µm; d & e, LS of the portion containing the copulatory apparatus: d, the extent of the male atrium (right); e, the extent of the female atrium (left). Both extend beyond the sectioned portion. Scale 2 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 8. Caenoplana bicolor. ZMB3506: mid-dorsal (a) and mid-ventral (b) TS showing the epithelium and cutaneous muscle, scales 50 µm; c, TS, enlargement of dorsal portion of (d), scale 200 µm; d, whole TS, scale 500 µm. The double-ended arrows above c and d indicate regions of sub-cutaneous melanic pigment presumed to correspond with the dark stripes of living animals, cf. Figs 6 & 10; e, LS of the pharynx, * indicates the position of the pharyngeal aperture, scale 2 mm (anterior to the right)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 9. Caenoplana bicolor. ZMB3506: a, LS showing several immature testes, scale 100 µm; b, TS showing ovovitelline duct, ventral nerve cord and immature testis on one side, scale 100 µm; c, longitudinal section showing the immature ovovitelline duct immediately dorsal to the ventral nerve cord, scale 100 µm; d, longitudinal section showing the immature ovovitelline duct and probable immature ovary, scale 100 µm; e, LS showing the maximum extent of the male atrium (right); f, showing the maximum extent of the female atrium (left), scale bar 2 mm; g and h, enlargement of the extremities of the female and male atria, scale bar 250 µm; e, diagram of the copulatory apparatus, male to the right." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 10. Caenoplana variegata. Cv7: mid-dorsal (a) and mid-ventral (b) TS showing the epithelium and cutaneous muscle, scales 50 µm; c, TS enlargement of dorsal portion of (d), scale 200 µm; d, whole TS, scale 500 µm. The double-ended arrows above c and d indicate regions of sub-cutaneous melanic pigment presumed to correspond with the dark stripes of living animals, cf. Figs 6 & 8; e, LS of the pharynx, * indicates the position of the pharyngeal aperture, scale 2 mm (anterior to the right)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 11. Caenoplana variegata. Cv7: a, LS showing several empty testes, scale 100 µm; b, TS showing the sperm duct, ventral nerve cord, ovovitelline duct and empty testis on one side, scale 100 µm; c, LS showing the ovovitelline duct on the dorsal margin of the ventral nerve cord, and the sperm duct on the ventral margin of the ventral nerve cord, scale 100 µm; d, LS of the copulatory apparatus, scale 2 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 12. Caenoplana variegata. Cv7: diagram and selected LS micrographs of the copulatory apparatus: a, overlapping images through the gonopore, female atrium to the left, male to the right; b–e, sections of the female atrium and ducts; f–j, sections through the male atrium and ducts (i is an enlargement of the entry of the combined sperm duct into cavity c1); k, diagrammatic reconstruction. All (except i) to the same scale, 1 mm scale bars shown in a and k (scale bar in i = 200 µm). Vertical lines link the same feature." ; fabio:hasRepresentation ; a fabio:Figure . dwc:collectionCode "NSW, T, NHMUK" ; dwc:verbatimLatitude "-34.66" ; dwc:verbatimLongitude "150.28" ; trt:gbifOccurrenceId "2563048066" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NHMUK" ; dwc:eventDate "2015-01-28" ; dwc:locality "Coventry" ; dwc:recordedBy "J. Tomnie & It & Killed & Cut" ; dwc:verbatimLatitude "52.38545" ; dwc:verbatimLongitude "-1.648287" ; trt:gbifOccurrenceId "2563048067" ; trt:httpUri ; a dwc:MaterialCitation .