@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , ; dc:creator "Manuel, Michaël; Ramahandrison, Andriamirado T." ; dc:title "Laccophilus bergsteni Manuel & Ramahandrison 2020, sp. nov." ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; trt:definesTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "502" ; bibo:issue "4" ; bibo:journal "Zootaxa" ; bibo:pubDate "2020-08-07" ; bibo:startPage "482" ; bibo:volume "4822" ; dc:creator "Manuel, Michaël; Ramahandrison, Andriamirado T." ; dc:date "2020" ; dc:title "Four new species of the diving beetle genus Laccophilus Leach, 1815 from Madagascar (Coleoptera, Dytiscidae, Laccophilini)" ; fabio:hasPart , , , , , ; a fabio:JournalArticle . dwc:authority "Manuel & Ramahandrison, 2020" ; dwc:authorityName "Manuel & Ramahandrison" ; dwc:authorityYear "2020" ; dwc:class "Insecta" ; dwc:family "Dytiscidae" ; dwc:genus "Laccophilus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Manuel & Ramahandrison, 2020" ; dwc:species "bergsteni" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Dytiscidae" ; dwc:genus "Laccophilus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "bergsteni" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dytiscidae" ; dwc:genus "Laccophilus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dytiscidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "FIGURES 1–4. Habitus in dorsal view of: (1) Laccophilus leguyaderi sp. nov. (holotype); (2) Laccophilus bergsteni sp. nov. (holotype); (3) Laccophilus makay sp. nov. (holotype); (4) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 5–8. Ventral surface of: (5) Laccophilus leguyaderi sp. nov. (holotype); (6) Laccophilus bergsteni sp. nov. (holotype); (7) Laccophilus makay sp. nov. (holotype); (8) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 23–30. Left (23, 25, 27, 29) and right (24, 26, 28, 30) parameres of: (23, 24) Laccophilus leguyaderi sp. nov. (holotype); (25, 26) Laccophilus bergsteni sp. nov. (holotype); (27, 28) Laccophilus makay sp. nov. (holotype); (29, 30) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 0.1 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 43–46. Habitats of Laccophilus leguyaderi sp. nov. (43–44) and Laccophilus bergsteni sp. nov. (45–46). (43) Shallow brooklet with peaty bottom in subalpine grassland (Andringitra massif, 2,000 m); (44) Rather deep brook with peaty bottom, slowly flowing among subalpine shrubs (Andringitra massif, 1,970 m; type locality of L. leguyaderi); (45) Slowly flowing stream in natural forest with muddy bottom and lots of tree leaves (Ankaratra massif, Tavolotara, 2,000 m); (46) Slowly flowing stream on flat area in subalpine grassland (Ankaratra massif, 2,400 m; type locality of L. bergsteni)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 53. Map showing the distribution of known localities of the newly described species. Red dot: L. leguyaderi; Blue dots: L. bergsteni; Magenta dot: L. bergsteni and L. rakouthae; yellow dot: L. makay. Names of corresponding protected areas or massifs indicated in italics above dots." ; fabio:hasRepresentation ; a fabio:Figure . dwc:locality "Ankaratra" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "2400" ; dwc:verbatimLatitude "-19.333334" ; dwc:verbatimLongitude "47.25" ; trt:gbifOccurrenceId "3019656346" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:eventDate "2017-12-22" ; dwc:locality "Manjakatompo" ; dwc:recordedBy "Manuel" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "2400" ; dwc:verbatimLatitude "-19.333334" ; dwc:verbatimLongitude "47.25" ; trt:gbifOccurrenceId "3019656315" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH, CMM, DEUA, MNHN, NHRS, NMPC, PBZT, MBC" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "3019656311" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:eventDate "2017-12-22" ; dwc:locality "Manjakatompo" ; dwc:recordedBy "Manuel" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2400" ; dwc:verbatimLatitude "-19.333334" ; dwc:verbatimLongitude "47.25" ; trt:gbifOccurrenceId "3019656340" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2017-12-21" ; dwc:locality "Manjakatompo" ; dwc:recordedBy "Manuel" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2050" ; dwc:verbatimLatitude "-19.35" ; dwc:verbatimLongitude "47.283333" ; trt:gbifOccurrenceId "3019656349" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2017-12-21" ; dwc:locality "Manjakatompo" ; dwc:recordedBy "Ramahandrison & Manuel" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2000" ; dwc:verbatimLatitude "-19.35" ; dwc:verbatimLongitude "47.283333" ; trt:gbifOccurrenceId "3019656305" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "MANS3" ; dwc:collectionCode "MANS" ; dwc:eventDate "2018-07-14" ; dwc:locality "Ankaratra mountain" ; dwc:recordedBy "Ramahandrison" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1800" ; dwc:verbatimLatitude "-19.35" ; dwc:verbatimLongitude "47.3" ; trt:gbifOccurrenceId "3019656312" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "MANS6" ; dwc:collectionCode "MANS" ; dwc:eventDate "2018-07-21" ; dwc:locality "Manjakatompo" ; dwc:recordedBy "Ramahandrison" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2000" ; dwc:verbatimLatitude "-19.35" ; dwc:verbatimLongitude "47.283333" ; trt:gbifOccurrenceId "3019656324" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "MANS5" ; dwc:collectionCode "MANS" ; dwc:eventDate "2019-02-19" ; dwc:locality "Manjakatompo" ; dwc:recordedBy "Ramahandrison" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2000" ; dwc:verbatimLatitude "-19.35" ; dwc:verbatimLongitude "47.283333" ; trt:gbifOccurrenceId "3019656350" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "MANS8" ; dwc:collectionCode "MANS" ; dwc:eventDate "2019-02-20" ; dwc:locality "Manjakatompo" ; dwc:recordedBy "Ramahandrison" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2000" ; dwc:verbatimLatitude "-19.35" ; dwc:verbatimLongitude "47.283333" ; trt:gbifOccurrenceId "3019656303" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "MANS13" ; dwc:collectionCode "MANS" ; dwc:eventDate "2019-02-22" ; dwc:locality "Ankaratra mountain" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1800" ; dwc:verbatimLatitude "-19.366667" ; dwc:verbatimLongitude "47.3" ; trt:gbifOccurrenceId "3019656344" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Manjakatompo" ; dwc:recordedBy "J. H. Randriamihaja" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "500" ; dwc:verbatimLatitude "-19.34485" ; dwc:verbatimLongitude "47.33381" ; trt:gbifOccurrenceId "3019656358" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "river Analamitana" ; dwc:recordedBy "J. H. Randriamihaja" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1750" ; dwc:verbatimLatitude "-19.36458" ; dwc:verbatimLongitude "47.29885" ; trt:gbifOccurrenceId "3019656319" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Manjakatompo" ; dwc:recordedBy "J. H. Randriamihaja" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1757" ; dwc:verbatimLatitude "-19.363972" ; trt:gbifOccurrenceId "3019656302" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Forestiere" ; dwc:recordedBy "J. H. Randriamihaja" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1847" ; dwc:verbatimLatitude "-19.336" ; trt:gbifOccurrenceId "3019656338" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Manjakatompo" ; dwc:recordedBy "J. H. Randriamihaja" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2046" ; dwc:verbatimLatitude "-19.349138" ; trt:gbifOccurrenceId "3019656309" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Manjakatompo" ; dwc:recordedBy "J. H. Randriamihaja" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1808" ; dwc:verbatimLatitude "-19.35425" ; trt:gbifOccurrenceId "3019656310" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2014-11-22" ; dwc:locality "Mad" ; dwc:recordedBy "Leg. J. Bergsten & R. Bukontaite & J. H. Randriamihaja & T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1340" ; dwc:verbatimLatitude "-47.2901" ; trt:gbifOccurrenceId "3019656329" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2014-11-23" ; dwc:locality "Saha forest" ; dwc:recordedBy "Leg. J. Bergsten & R. Bukontaite & J. H. Randriamihaja & T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1240" ; dwc:verbatimLatitude "-47.9443" ; trt:gbifOccurrenceId "3019656307" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2014-11-23" ; dwc:locality "Saha forest" ; dwc:recordedBy "Leg. J. Bergsten & R. Bukontaite & J. H. Randriamihaja & T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1320" ; trt:gbifOccurrenceId "3019656328" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NHRS-" ; dwc:eventDate "2016-02-03" ; dwc:locality "Andalantsoavaly" ; dwc:recordedBy "Leg. T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1599" ; dwc:verbatimLatitude "-19.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656313" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NHRS-" ; dwc:eventDate "2016-02-03" ; dwc:locality "Manjakatompo Reserve" ; dwc:recordedBy "Leg. T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1796" ; dwc:verbatimLatitude "-19.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656333" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2016-02-04" ; dwc:locality "Manjakatompo Reserve" ; dwc:recordedBy "Leg. T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2044" ; dwc:verbatimLatitude "-19.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656337" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2016-02-05" ; dwc:locality "Reserve" ; dwc:recordedBy "Leg. T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1759" ; dwc:verbatimLatitude "-19.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656306" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Ambohitantely Special Reserve" ; dwc:recordedBy "T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1533" ; dwc:verbatimLatitude "-18.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656356" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Ambohitantely Special Reserve" ; dwc:recordedBy "T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1553" ; dwc:verbatimLatitude "-18.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656345" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Foret d'Anjozorobe Angavo" ; dwc:recordedBy "T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1306" ; dwc:verbatimLatitude "-18.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656322" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Foret d'Anjozorobe Angavo" ; dwc:recordedBy "T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1298" ; dwc:verbatimLatitude "-18.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656301" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Foret d'Anjozorobe Angavo" ; dwc:recordedBy "T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1367" ; dwc:verbatimLatitude "-18.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656334" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Ruisseau Ambatovikinina" ; dwc:recordedBy "T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1367" ; dwc:verbatimLatitude "-18.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656364" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Foret d'Anjozorobe Angavo" ; dwc:recordedBy "T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1331" ; dwc:verbatimLatitude "-18.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656336" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Foret d'Anjozorobe Angavo" ; dwc:recordedBy "T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1356" ; dwc:verbatimLatitude "-18.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656335" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Foret d'Anjozorobe Angavo" ; dwc:recordedBy "T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1308" ; dwc:verbatimLatitude "-18.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656342" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Foret d'Anjozorobe Angavo" ; dwc:recordedBy "T. Ranarilalatiana" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1341" ; dwc:verbatimLatitude "-18.0" ; dwc:verbatimLongitude "47.0" ; trt:gbifOccurrenceId "3019656321" ; trt:httpUri ; a dwc:MaterialCitation .