@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , , , , , , , , , ; dc:creator "Lowe, Graeme; Kovařík, František; Stockmann, Mark; Šťáhlavský, František" ; dc:title "Trypanothacus barnesi Lowe & Kovařík & Stockmann & Šťáhlavský 2019, sp. n." ; dwc:basisOfRecord , , , , , , , , ; trt:definesTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "30" ; bibo:journal "Euscorpius" ; bibo:pubDate "2019-12-31" ; bibo:startPage "1" ; bibo:volume "277" ; dc:creator "Lowe, Graeme; Kovařík, František; Stockmann, Mark; Šťáhlavský, František" ; dc:date "2019" ; dc:title "Trypanothacus gen. n., a new genus of burrowing scorpion from the Arabian Peninsula (Scorpiones: Buthidae)" ; fabio:hasPart , , , , , , , , , , , , , , ; a fabio:JournalArticle . dwc:authority "Lowe & Kovařík & Stockmann & Šťáhlavský, 2019" ; dwc:authorityName "Lowe & Kovařík & Stockmann & Šťáhlavský" ; dwc:authorityYear "2019" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Trypanothacus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lowe & Kovařík & Stockmann & Šťáhlavský, 2019" ; dwc:species "barnesi" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Trypanothacus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "barnesi" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Trypanothacus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "Figures 107–110: Morphometric scaling relations of the telson of fossorial ‘Buthus group’ scorpions. Figure 107. Double logarithmic scatterplot of telson vesicle length/ depth (L/ D) ratio vs. aculeus length normalized to vesicle length (aculeus L/ vesicle L), for large pelophiles/ semipsammophiles (4 genera, 53 species) (Buthus, Odontobuthus, Pantobuthus: orange circles; Trypanothacus gen. n.: blue circles) and large psammophiles/ ultrapsammophiles (6 genera, 29 species) (Buthacus: cyan circles; Vachoniolus: dark green circles; Apistobuthus, Buthiscus, Liobuthus, and Plesiobuthus: yellow circles). Dark line: linear least squares regression fit to all data shown (Pearson correlation coefficient, R = 0.57). Upper right inset: diagram of measurements estimated from inverted telson lateral profiles, with dorsal surface of vesicle approximately leveled; SA = ventral profile length of aculeus from base point to tip, LV = length of vesicle from anterior limit or inflection point to aculeus base point; HV = vesicle depth or height (distance between dorsal and ventral tangent planes). Base point was placed at local minimum of inverted ventral profile curve. SA was calculated by cubic spline fit (>10 anchor points). For a minority of broken tips, SA was estimated by intersecting extrapolated dorsal and ventral profile curves (<5%). Figure 108. Scatterplot of fossorial buthids in Figure 107, superposed upon larger scale plot of Order Scorpiones (gray circles). Gray line: linear least squares regression fit to Order Scorpiones data (including data in Figure 107) (Pearson correlation coefficient, R = –0.62).Also plotted are data from small ‘Buthus group’ psammophiles/ ultrapsammophiles (5 genera, 10 species: Compsobuthus arabicus, Lanzatus somalilandus, Neobuthus kutcheri, Picobuthus dundoni, P. wahibaensis, Anomalobuthus krivochatskyi, A. pavlovskyi, A. rickmersi, A. lowei, A. talebii) (dark yellow circles). Figure 109. Double logarithmic scatterplot of carapace length (L) vs. aculeus length normalized to vesicle length (i.e., aculeus L/ vesicle L) for psammophile/ ultrapsammophile members of the ‘Buthus group’, including large species in Figure 107, and small species in Figure 108 (dark yellow circles). Blue line: linear least squares regression fit to all data shown (Pearson correlation coefficient, R = 0.74). Figure 110. Scatterplot of psammophile buthid data in Figure 109 (all in one color, blue circles) superposed upon larger scale plot of Order Scorpiones (gray circles). Gray line: linear least squares regression to Order Scorpiones data (including data in Figure 109) (Pearson correlation coefficient, R = –0.01). Also plotted are data from bothriurid genus Brachistosternus (43 species) (magenta triangles), and corresponding linear least squares regression fit (R = 0.39). All plots in Figures 107-110 include data from both males and females. Data were assembled by mining taxonomic literature or directly measuring specimens. Included are 1854 species or subspecies (1331 ♂, 1201 ♀) representing all known extant genera (1841 species or subspecies) plus 7 fossil genera (13 species)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 1–4: Trypanothacus barnesi gen. et sp. n. Figures 1–2. Male paratype from the type locality in dorsal (1) and ventral (2) views. Figures 3–4: Female paratype from the type locality in dorsal (3) and ventral (4) views. Scale bar: 10 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 15–22: Trypanothacus barnesi gen. et sp. n. Figures 15, 20–22. Male holotype, metasoma V and telson lateral view (15), metasoma and telson lateral (20), ventral (21), and dorsal (22) views. Figures. 16–19. Female paratype from type locality, metasoma V and telson lateral view (16), metasoma and telson lateral (17), ventral (18), and dorsal (19) views. Scale bars: 4 mm (17–19, 20–21)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 29–46: Figures 29–42: Trypanothacus barnesi gen. et sp. n. Figures 29–40. Pedipalp segments. Figures 29–38. Male holotype, chela dorsal (29), external (30), and ventral (31) views. Patella dorsal (32), external (33) and ventral (34) views. Femur and trochanter internal (35), dorsal (36), and ventral (37) views. Movable finger dentition (38). Trichobothrial pattern is indicated in Figures 30–33 and 35–36. Figures 39– 40. Female paratype from the type locality, chela dorsal (39) and external (40) views. Figures 41–42. Telson, male holotype (41) and female paratype from type locality (42), lateral views. Figures 43–44: T. buettikeri comb. n., pedipalp chela dorsal, male holotype (43) and female paratype (44). Figures 45–46: Buthacus stockmanni Kovařík et al., 2016, telson, male holotype (45) and female paratype (46), lateral views." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 71–77: Trypanothacus barnesi gen. et sp. n. Figures 71–73. Paratype male, right chelicera, dorsal (71), ventral (72) and ventral UV fluorescence (73) views. Scale bar: 1 mm. Figures 74–77. Holotype male, right hemispermatophore, capsule region with lobes in anterior (74), convex compressed (75) and posterior (76) views, and whole hemispermatophore in convex view (77). Scale bars: 1 mm (71–73), 500 µm (74–76), 1 mm (77)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 5–10: Trypanothacus barnesi gen. et sp. n. Figures 5, 7, 9. Male holotype, carapace and tergites I–V (5), coxosternal area and sternites (7), right legs I–IV, retrolateral aspect (9.I–IV). Figures 6, 8, 10. Female paratype from the type locality, carapace and tergites I–III (6), coxosternal area and sternites III–IV (8), right legs III–IV, retrolateral aspect (10.III–IV)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 11–12: Trypanothacus barnesi gen. et sp. n. Male paratype from Thumrait (VII.1997), carapace and tergites I–VII (11), coxosternal area and sternites (12). UV fluorescence. Scale bars: 2 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 13–14: Trypanothacus barnesi gen. et sp. n. Female paratype from Yalooni (XII.1988), carapace and tergites I–VII (13), coxosternal area and sternites (14). UV fluorescence. Scale bar: 2 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 47–58: Trypanothacus barnesi gen. et sp. n. Pedipalp segments of male paratype from Thumrait (VII.1997), chela dorsal (47), external (48), ventral (49) and internal (50) views. Patella dorsal (51), external (52), ventral (53) and internal (54) views. Femur and trochanter dorsal (55), external (56), ventral (57) and internal (58) views. UV fluorescence. Scale bar: 2 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 59–70: Trypanothacus barnesi gen. et sp. n. Pedipalp segments of female paratypes, chela dorsal (59), external (60), ventral (61) and internal (62) views. Patella dorsal (63), external (64), ventral (65) and internal (66) views. Femur and trochanter dorsal (67), external (68), ventral (69) and internal (70) views. Dorsal views (59, 63, 67): paratype female from Thumrait (XI.1989) (with normal dorsointernal carina on femur). External (60, 64, 68), ventral (61, 65, 69) and internal (62, 66, 70) views: paratype female from Yalooni (XII.1988) (with malformed dorsointernal carina on femur). UV fluorescence. Scale bars: 2 mm (59, 63, 67), 2 mm (60–62, 64–66, 69–70)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 111–113: Trypanothacus barnesi gen. et sp. n., mitotic metaphase (111), postpachytene (112) and ideogram (113) (y axis - % of the chromosome length of the haploid set) of male holotype. Scale bar = 5 μm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 84–85: Trypanothacus barnesi gen. et sp. n., in vivo habitus, male holotype (84) and female paratype from type locality (85)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 86–89: Trypanothacus barnesi gen. et sp. n.. Figures 86–87. Male (86) and female (87) from Thumrait, photos by J. N. Barnes. Figures 88–89. Type locality, Oman, S of Schelim, 18.041070°N 55.612262°E." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 114–116: Trypanothacus barnesi gen. et sp. n., promenade à deux (courtship dance) in a mating pair of individuals collected from the type locality. Photos taken in laboratory. Male is on the right." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figures 117–118: Trypanothacus barnesi gen. et sp. n., cheliceral massage (117) and spermatophore deposition (118) in a mating pair of individuals collected from the type locality. Photos taken in laboratory. Male is on the right." ; fabio:hasRepresentation ; a fabio:Figure . dwc:collectionCode "FKCP" ; dwc:locality "S of Shalim" ; dwc:typeStatus "holotype" ; dwc:verbatimLatitude "18.04107" ; dwc:verbatimLongitude "55.612263" ; trt:gbifOccurrenceId "3758760305" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ONHM 1186" ; dwc:collectionCode "NHMB" ; dwc:eventDate "1988-12" ; dwc:locality "Jiddat Al Harasis" ; dwc:recordedBy "Yalooni" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "19.95" ; dwc:verbatimLongitude "57.1" ; trt:gbifOccurrenceId "3758760309" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ONHM 1416" ; dwc:collectionCode "NHMB" ; dwc:eventDate "1989-11-10" ; dwc:locality "Thamarit" ; dwc:recordedBy "I. A. J. Brown" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "17.65" ; dwc:verbatimLongitude "54.033333" ; trt:gbifOccurrenceId "3758760310" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NHMB, GLPC, ONHM" ; dwc:eventDate "1997-07" ; dwc:locality "Thumrait" ; dwc:recordedBy "J. N. Barnes" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "17.7" ; dwc:verbatimLongitude "53.983334" ; trt:gbifOccurrenceId "3758760307" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NHMB" ; dwc:eventDate "1998-07-23" ; dwc:locality "Thumrait" ; dwc:recordedBy "J. N. Barnes" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "17.7" ; dwc:verbatimLongitude "53.816666" ; trt:gbifOccurrenceId "3758760303" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "No. 1594" ; dwc:collectionCode "FKCP, GLPC" ; dwc:eventDate "2016-09-06" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "18.04107" ; dwc:verbatimLongitude "55.612263" ; trt:gbifOccurrenceId "3758760312" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FKCP" ; dwc:eventDate "2017-10-27" ; dwc:locality "S of Shalim" ; dwc:recordedBy "M. Stockmann" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "18.04107" ; dwc:verbatimLongitude "55.612263" ; trt:gbifOccurrenceId "3758760308" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "RS8787" ; dwc:collectionCode "MNHN" ; dwc:eventDate "1980-03-22" ; dwc:locality "Dhofar" ; dwc:recordedBy "J. N. Barnes & Barnes" ; dwc:verbatimElevation "450" ; dwc:verbatimLatitude "17.666666" ; dwc:verbatimLongitude "54.033333" ; trt:gbifOccurrenceId "3758760301" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "MDG6020" ; dwc:collectionCode "NHMB" ; dwc:eventDate "1980-11-10" ; dwc:locality "Jiddat Al Harasis" ; dwc:recordedBy "M. D. Gallagher" ; dwc:verbatimLatitude "19.933332" ; dwc:verbatimLongitude "57.1" ; trt:gbifOccurrenceId "3758760304" ; trt:httpUri ; a dwc:MaterialCitation .