@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , , , , , ; dc:creator "López-García, Margarita M.; Deloya, Cuauhtémoc" ; dc:title "Ligyrus (Anagrylius) moroni López-García & Deloya 2022, new combination" ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , , , , , , ; trt:definesTaxonConcept ; trt:deprecates ; trt:publishedIn ; a trt:Treatment . bibo:endPage "119" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:pubDate "2022-11-22" ; bibo:startPage "1" ; bibo:volume "5211" ; dc:creator "López-García, Margarita M.; Deloya, Cuauhtémoc" ; dc:date "2022" ; dc:title "Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" ; fabio:hasPart , , , , , , , , , , ; a fabio:JournalArticle . dwc:authority "(Lopez-Garcia & Deloya, 2019)" ; dwc:authorityName "López-García & Deloya" ; dwc:authorityYear "2022" ; dwc:baseAuthorityName "Lopez-Garcia & Deloya" ; dwc:baseAuthorityYear "2019" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Lopez-Garcia & Deloya, 2019) López-García & Deloya, 2022" ; dwc:species "moroni" ; dwc:subGenus "Anagrylius" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Lopez-Garcia & Deloya, 2019: 131" ; dwc:authorityName "Lopez-Garcia & Deloya" ; dwc:authorityPageNumber "131" ; dwc:authorityYear "2019" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Tomarus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lopez-Garcia & Deloya, 2019" ; dwc:species "moroni" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "moroni" ; dwc:subGenus "Anagrylius" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:subGenus "Anagrylius" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Tomarus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "moroni" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Tomarus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 2. Labium. A) Ligyrus (Anagrylius) moroni, B) Ligyrus (Ligyrus) allonasutus, C) L. (Ligyrus) gibbosus, D) L. (Ligyrus) villosus, E) L. (Ligyrodes) relictus, F) Proculigyrus cicatricosus, G) Euligyrus ebenus, H) E. similis, I) Tomarus gyas, J) Tomarus selanderi, K) Tomarus subtropicus. Abbreviations: LL: Ligular lobes, LP: Labial palpus, M: Mentum, S: Suture." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 3. Maxilla. A) Ligyrus (Anagrylius) moroni, B) L. (Ligyrodes) sallaei, C) Ligyrus (Ligyrus) allonasutus. Numeration of teeth of galea: D) Euligyrus similis, E) L. (Ligyrodes) sallaei, F) Tomarus gyas, G) L. (Ligyrus) fossor, H) L. (Ligyrus) allonasutus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 11. Protibia in frontal view.A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) relictus, E) L. (Ligyrodes) sallaei, F) L. (Ligyrus) allonasutus, G) L. (Ligyrus) villosus, H) Tomarus bituberculatus, I) T. colombianus, J) T. pullus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 12. Mesotibia (A–F). A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei; E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) villosus. Metatibia (G–L). G) Euligyrus similis, H) Proculigyrus cicatricosus, I) Ligyrus (Anagrylius) moroni, J) L. (Ligyrodes) sallaei, K) L. (Ligyrus) allonasutus, L) L. (Ligyrus) villosus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 13. Female genital plates. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) bidentulus, G) L. (Ligyrus) neglectus, H) Tomarus laticaudus, I) T. maternus, J) T. rostratus, K) T. subtropicus. Abbreviations: SCx: Subcoxite, Cx: Coxite." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 17. Parameres in lateral view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus; E) L. (Anagrylius) moroni, F) L. (Ligyrodes) relictus, G) L. (Ligyrodes) sallaei." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 21. Internal sac. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus. Abbreviations:AL:Accessory lamellae, Br: Brush of setae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 25. Habitus of the species of Ligyrus. Subgenus Anagrylius: A) Ligyrus cuniculus, B) L. moroni. Subgenus Ligyrodes: C) L. peruvianus, D) L. relictus, E) L. sallaei. Subgenus Ligyrus: F) L. allonasutus, G) L. bidentulus, H) L. burmeisteri. Scale bar: 5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 32. Distributional records of the subgenus Ligyrus (Anagrylius)." ; fabio:hasRepresentation ; a fabio:Figure . dwc:collectionCode "MZUSP" ; dwc:recordedBy "Original & Male" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3986696666" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MZUSP" ; dwc:eventDate "1967-03-11" ; dwc:locality "Ubatuba" ; dwc:recordedBy "Female" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3986696423" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MZUSP" ; dwc:eventDate "1968-02-15" ; dwc:locality "Ubatuba" ; dwc:recordedBy "Male" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "3986696547" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MZUSP" ; dwc:eventDate "1963-04-02" ; dwc:locality "Baraquecaba" ; dwc:recordedBy "P. Araujo & Female" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "3986696379" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MZUSP" ; dwc:eventDate "1964-03" ; dwc:locality "Rio Janeiro" ; dwc:recordedBy "Female" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "3986696716" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEIOC" ; dwc:eventDate "1957-12" ; dwc:locality "Nossa" ; dwc:recordedBy "J. Lane & Female" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "3986696527" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "CEIOC 32162" ; dwc:collectionCode "CEIOC" ; dwc:locality "Rio" ; dwc:recordedBy "T. Freitas" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "3986696448" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "CEIOC 31535" ; dwc:collectionCode "CEIOC" ; dwc:locality "Ebert" ; dwc:recordedBy "S. Paulo" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "3986697042" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "CEIOC 31546" ; dwc:collectionCode "USNM" ; dwc:locality "Frade" ; dwc:recordedBy "A. Jesus" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "3986696795" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "USNM" ; dwc:eventDate "1957-11" ; dwc:locality "Mongagua" ; dwc:recordedBy "S. Paulo" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "3986696489" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "UNSM" ; dwc:eventDate "1957-11" ; dwc:locality "Mongagua" ; dwc:recordedBy "S. Paulo" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "3986696447" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "UNSM" ; dwc:locality "Barbiellini" ; dwc:recordedBy "Female" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "3986696516" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Barbiellini" ; dwc:recordedBy "Ubatuba" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "3986696398" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696797" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696697" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696345" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696690" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696456" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696359" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696591" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696756" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696874" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696407" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696470" ; trt:httpUri ; a dwc:MaterialCitation . trt:gbifOccurrenceId "3986696562" ; trt:httpUri ; a dwc:MaterialCitation .