@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .
@prefix xlink: <http://www.w3.org/1999/xlink/> .

<http://treatment.plazi.org/id/4E574931E2725135B5A689A4E7DD70E0>
    cito:cites <http://dx.doi.org/10.3897/zookeys.881.38026.figure1>, <http://dx.doi.org/10.3897/zookeys.881.38026.figure2>, <http://dx.doi.org/10.3897/zookeys.881.38026.figure3>, <http://dx.doi.org/10.3897/zookeys.881.38026.figure5>, <http://dx.doi.org/10.3897/zookeys.881.38026.figure6>, <http://dx.doi.org/10.3897/zookeys.881.38026.figure7>, <http://dx.doi.org/10.3897/zookeys.881.38026.figure9> ;
    dc:creator "Kohlmann, Bert; Solis, Angel; Alvarado, Guillermo E." ;
    dc:title "Onthophagus humboldti Kohlmann, Solis & Alvarado, 2019, sp. nov." ;
    trt:publishedIn <http://dx.doi.org/10.3897/zookeys.881.38026> ;
    trt:treatsTaxonName <http://taxon-name.plazi.org/id/Animalia/Onthophagus_humboldti> ;
    a trt:Treatment .

<http://dx.doi.org/10.3897/zookeys.881.38026>
    bibo:endPage "51" ;
    bibo:journal "ZooKeys" ;
    bibo:startPage "23" ;
    bibo:volume "881" ;
    dc:creator "Kohlmann, Bert; Solis, Angel; Alvarado, Guillermo E." ;
    dc:date "2019" ;
    dc:title "Description of Onthophagus humboldti and Uroxys bonplandi, two new scarab beetles (Coleoptera, Scarabaeidae, Scarabaeinae) from Costa Rica, with notes on tropical mountain brachyptery and endemicity" ;
    fabio:hasPart <http://dx.doi.org/10.3897/zookeys.881.38026.figure1>, <http://dx.doi.org/10.3897/zookeys.881.38026.figure2>, <http://dx.doi.org/10.3897/zookeys.881.38026.figure3>, <http://dx.doi.org/10.3897/zookeys.881.38026.figure5>, <http://dx.doi.org/10.3897/zookeys.881.38026.figure6>, <http://dx.doi.org/10.3897/zookeys.881.38026.figure7>, <http://dx.doi.org/10.3897/zookeys.881.38026.figure9> ;
    a fabio:JournalArticle .

<http://taxon-name.plazi.org/id/Animalia/Onthophagus_humboldti>
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Onthophagus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:species "humboldti" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Onthophagus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Onthophagus>
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Onthophagus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Scarabaeidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Scarabaeidae>
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "family" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Coleoptera> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Coleoptera>
    dwc:class "Insecta" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "order" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Insecta> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Insecta>
    dwc:class "Insecta" ;
    dwc:kingdom "Animalia" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "class" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Arthropoda> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Arthropoda>
    dwc:kingdom "Animalia" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "phylum" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia>
    dwc:kingdom "Animalia" ;
    dwc:rank "kingdom" ;
    a dwcFP:TaxonName .

<http://dx.doi.org/10.3897/zookeys.881.38026.figure1>
    dc:description "Figure 1. Dorsal drawing of a male Onthophagus humboldti sp. nov." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/347596> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.881.38026.figure2>
    dc:description "Figure 2. Drawings of the clypeal horn of a O. humboldti sp. nov. and b O. micropterus; pronotum of c O. humboldti sp. nov. and d O. micropterus; head of e O. humboldti sp. nov. and f O. micropterus; and elytral apex of g U. bonplandi sp. nov. and h U. dybasi. Scale bars: 1 mm." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/347597> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.881.38026.figure3>
    dc:description "Figure 3. Aedeagui of a O. humboldti sp. nov. b U. bonplandi sp. nov. c brachypterous wing of O. humboldti sp. nov." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/347598> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.881.38026.figure5>
    dc:description "Figure 5. Known distribution of O. humboldti sp. nov. (orange circle) and U. bonplandi sp. nov. (red triangle). The distribution of the proposed sister species of these new taxa is also depicted, O. micropterus (blue rhombus) and U. dybasi (black square)." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/347600> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.881.38026.figure6>
    dc:description "Figure 6. Known distribution of brachypterous Scarabaeinae in Costa Rica." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/347601> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.881.38026.figure7>
    dc:description "Figure 7. Present day distribution of Onthophagus humboldti sp. nov. (blue dot) and O. micropterus (red dot) and lines indicating proposed localities (rhombi) depressed by 1500 m (14 km in straight line) generated by the last glacial maximum, ~ 25 - 23 ka, in the Cordillera de Talamanca. All mountain systems are 150 m lower than present day height and an estimated sea level descent of 120 m is depicted. Dotted black lines represent present-day sea levels." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/347602> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/zookeys.881.38026.figure9>
    dc:description "Figure 9. A cytochrome c-oxidase I (COI- 5 P) mitochondrial DNA sequence-based BOLD taxon ID tree of the nearest species of O. humboldti sp. nov." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/347604> ;
    a fabio:Figure .