@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , , , , , , , , , , , , ; dc:creator "Newman, Michael J.; Burrow, Carole J.; Den Blaauwen, Jan L.; Davidson, Robert G." ; dc:title "Euthacanthus macnicoli Powrie 1864" ; dwc:basisOfRecord , , , , , ; trt:augmentsTaxonConcept ; trt:citesTaxonName ; trt:deprecates , , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "348" ; bibo:issue "3" ; bibo:journal "Geodiversitas" ; bibo:pubDate "2014-09-26" ; bibo:startPage "321" ; bibo:volume "36" ; dc:creator "Newman, Michael J.; Burrow, Carole J.; Den Blaauwen, Jan L.; Davidson, Robert G." ; dc:date "2014" ; dc:title "The Early Devonian acanthodian Euthacanthus macnicoli Powrie, 1864 from the Midland Valley of Scotland" ; fabio:hasPart , , , , , , , , , , , , , , , , ; a fabio:JournalArticle . dwc:authority "Powrie, 1864" ; dwc:authorityName "Powrie" ; dwc:authorityYear "1864" ; dwc:family "Euthacanthidae" ; dwc:genus "Euthacanthus" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Powrie, 1864" ; dwc:species "macnicoli" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Powrie, 1870: 292" ; dwc:authorityName "Powrie" ; dwc:authorityPageNumber "292" ; dwc:authorityYear "1870" ; dwc:family "Euthacanthidae" ; dwc:genus "Euthacanthus" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Powrie, 1870" ; dwc:species "grandis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Powrie" ; dwc:baseAuthorityYear "1870" ; dwc:class "Acanthodii" ; dwc:family "Climatiidae" ; dwc:genus "Climatius" ; dwc:kingdom "Animalia" ; dwc:order "Climatiiformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Powrie, 1870)" ; dwc:species "grandis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Powrie" ; dwc:baseAuthorityYear "1864" ; dwc:class "Acanthodii" ; dwc:family "Climatiidae" ; dwc:genus "Climatius" ; dwc:kingdom "Animalia" ; dwc:order "Climatiiformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Powrie, 1864)" ; dwc:species "macnicoli" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Powrie" ; dwc:authorityYear "1864" ; dwc:family "Euthacanthidae" ; dwc:genus "Euthacanthus" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Powrie, 1864" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Powrie" ; dwc:baseAuthorityYear "1864" ; dwc:family "Euthacanthidae" ; dwc:genus "Euthacanthus" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Powrie, 1864)" ; dwc:species "macnicolli" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:family "Euthacanthidae" ; dwc:genus "Euthacanthus" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "macnicoli" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Euthacanthidae" ; dwc:genus "Euthacanthus" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Euthacanthidae" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:family "Euthacanthidae" ; dwc:genus "Euthacanthus" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "grandis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Acanthodii" ; dwc:family "Climatiidae" ; dwc:genus "Climatius" ; dwc:kingdom "Animalia" ; dwc:order "Climatiiformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "grandis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Acanthodii" ; dwc:family "Climatiidae" ; dwc:genus "Climatius" ; dwc:kingdom "Animalia" ; dwc:order "Climatiiformes" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Acanthodii" ; dwc:family "Climatiidae" ; dwc:kingdom "Animalia" ; dwc:order "Climatiiformes" ; dwc:phylum "Chordata" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Acanthodii" ; dwc:kingdom "Animalia" ; dwc:order "Climatiiformes" ; dwc:phylum "Chordata" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Acanthodii" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Acanthodii" ; dwc:family "Climatiidae" ; dwc:genus "Climatius" ; dwc:kingdom "Animalia" ; dwc:order "Climatiiformes" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "macnicoli" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Euthacanthidae" ; dwc:genus "Euthacanthus" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "mitchelli" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Euthacanthidae" ; dwc:genus "Euthacanthus" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "macnicolli" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIG. 3.— Euthacanthus macnicoli Powrie,1864:A, NMS G.1891.92.231 from Turin Hill (Tillywhandland),the holotype;B, NMS G.2002.59.97, a moderate sized individual from Duntrune. Scale bars: 10 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 4. — Euthacanthus macnicoli Powrie, 1864, head and branchial region: A, holotype NMS G.1891.92.231 from Turin Hill; B, NMS G.1891.92.236 from Turin Hill;C, NMS G.2011.33.1 from Balruddery Den; D, NMS G.2002.59.97 from Duntrune.Abbreviations: gr, gular rays; lt.br.p, left side branchiostegal plates; lt.ll, left side lateral line; lt.sb, left sclerotic bones; lt.sc, left scapulocoracoid; p.fs, pectoral fin spine; rt.br.p, right side branchiostegal plates; rt.ll, right side lateral line;rt.pp.p, right “prepectoral plate”;rt.sb, right side sclerotic bones; rt.sc, right scapulocoracoid; sgc1, first subsidiary gill cover; sgc2, second subsidiary gill cover; sgc3, third subsidiary gill cover. Scale bars: 5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 5.— Euthacanthus macnicoli Powrie,1864,head and branchial region:A, NMS G.1891.92.238 from Turin Hill;B, NMS G.1891.92.275 from Turin Hill; C, NMS G.2007.24.2 from Tillywhandland; D, NMS G.2002.59.100 from Duntrune. Abbreviations: br.p, branchiostegal plates;gr, gular rays;hy.op, hyoid operculum;io.sl, infraorbital sensory line canal; ll, lateral line;lt.br.p, left branchiostegal plates;lt.oc, left otic capsule; lt.sb, left sclerotic bone; m.op, mandibular operculum; p.fs, pectoral fin spine; pop.c, preopercular sensory line canal; pop.sl, preopercular sensory line; pp.p, “prepectoral plate”; rt.br.p, right branchiostegal plates; rt.sb, right sclerotic bones; sb, scle- rotic bone; sc, scapulocoracoid; so.sl, supraorbital sensory line canal; sgc1-3, first to third subsidiary gill covers. Scale bars: 5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 6. — Euthacanthus macnicoli Powrie, 1864, jaws and branchiostegal plate ornament: A, NMS G.1967.12.7 from Turin Hill (Til- lywhandland); C, D, branchiostegal plates on QMF57176 from Tillywhandland, under normal light (C) and whitened with magnesium oxide (D); E, branchiostegal plate in NHM P.67308,a large regurgitate or coprolite.Abbreviations:br.p, branchiostegal plates;lt.mc, left Meckel’s cartilage; lt.pq, left palatoquadrate cartilage; rt.pq, right palatoquadrate cartilage. Scale bars: A, B, 10 mm; C-E, 1 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 7. — Euthacanthus macnicoli Powrie, 1864, head tesserae: NMS G.1881.5.60 from Turin Hill. Scale bar: 1 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 8. — Euthacanthus macnicoli Powrie, 1864, shoulder girdle elements on NMS G.2010.7.39: A, left prepectoral plate; B, right prepectoral plate, medial surface; C, left scapulocoracoid, lateral surface; D, right scapulocoracoid, lateral surface. Abbreviations: br.p, branchiostegal plates; p.fs, pectoral fin spine. Scale bar: 2 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 9. — Euthacanthus macnicoli Powrie, 1864. Reconstruction of the head in lateral view, much modified after Watson (1937, fig. 3)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 10.— Euthacanthus macnicoli Powrie,1864 fin spines:A, NMS G.1967.12.5 from Turin Hill,posterior and anterior dorsal fin spines; B, NMS G.1885.54.6B, anal fin spine; C, NMS G.2002.59.27p, lower half of an articulated fish showing the fin spines; D-I, ground thin sections of fin spines on NMS G.2010.7.42; D-F, cross-sections of an anal fin spine; D, near the tip of the fin spine with no pulp cavity; E, showing vascular canals of the middle layer,and lamellar bone forming inner layer lining central cavity; F, mid-proximal end of spine, with open central cavity open along leading edge; G-I, thin sections of posterior dorsal fin spine; G, mid-distal end of spine with closed central cavity; H, leading edge ridge with mesodentine forming ridge, osteodentine forming middle layer, and very thin" ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 11. — Euthacanthus macnicoli Powrie, 1864, fin spine his- tological structure: Serial sections of posterior dorsal (pd.fs) and anal (a.fs) fin spines of NMS G.2010.7.42. Scale bar: 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 12. — Euthacanthus macnicoli Powrie, 1864, tail squamation: A, tail of holotype NMS G.1891.92.231; B, squamation pattern in E. macnicoli, based on that for Acanthodes sp. as figured by Miles (1970: fig. 7). Scale bar: 10 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 13. — Euthacanthus macnicoli Powrie, 1864 NMS G.2013.8.1, an articulated moderate sized specimen from Tillywhandland: A, specimen before sectioning;B-E, light microscope views of articulated squamation patches;B, pectoral region;C, midflank,midbody; D, anterior to caudal peduncle between posterior dorsal and anal fin spines;E, tail lobe (zone Z1). Scale bars: A, 10 mm; B-E, 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 14. — Euthacanthus macnicoli Powrie, 1864 NMS G.2013.8.1, thin sections of individual scales in serial sections of an articu- lated, moderate sized specimen from Tillywhandland: A-C, sections in the pectoral region; A, vertical longitudinal section with three crown growth zones; B, median vertical longitudinal section along a groove between crown ridges;C, vertical transverse section with three growth zones; D, E, vertical transverse sections in the anterior midflank region; F-H, sections in the midbody midflank region; F, oblique section; G, H, vertical transverse sections; I-K, sections in the posterior midflank region between the posterior dorsal and anal fin spines; I, vertical transverse section; J, K, vertical longitudinal section; L-N, sections in the caudal peduncle-midtail region; L, M, vertical transverse section; N, vertical longitudinal section. Abbreviations: bcl, bone cell lacunae; igl, incremental growth lines; m, mesodentine; Sf, Sharpey’s fibres; Sg, Stranggewebe; vc, vascular canal. Scale bars: A, L, 0.05 mm; B-K, M, N, 0.1 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 15. — Euthacanthus macnicoli Powrie, 1864 specimen NMS G.2010.7.42, the rear half with the posterior dorsal and anal fin spines of a small articulated specimen from Tillywhandland: A, the specimen before thin sectioning with the anterior to the right; white lines shown the areas thin sectioned; B-M, SEM views of scales; B, C, squamation segment cut midflank below the poste- rior dorsal fin spine; D, E, squamation segment cut at the middle of the caudal peduncle; F-M, scales in residues after acetic acid treatment of remainder of cut portion; F, articulated patch of scales (both sides of fish compressed together); G, H, scale in crown and lateral view; I, J, scale in crown and lateral view; K, scale from one side of the body attached to two scales from the other side; L, M, scale in crown and posterolateral view; N-P, thin sections of scales; N, vertical transverse section of a young fin web scale; O, vertical longitudinal section of a fin web scale; P, vertical transverse sections of two scales abutting a fin spine. Scale bars: A, 10 mm; B, D, 1 mm; C, E-P, 0.1 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 16.— Euthacanthus macnicoli Powrie,1864 specimen NMS G.2010.7.39,SEMs of scales from a large partial articulated specimen from Tillywhandland: A, B, scale in crown and posterolateral view; C-D, scale in crown and anterolateral view; E, scale in laterocrown view; F, G, scale in crown and posterior view; H, scale in laterocrown view; I, J, scale in anterocrown and anterior view; K, L, scale in crown and lateral view; M, N, scale in crown and lateral view; O, scale in laterocrown view; P, scale in anterior view; Q, scale in anterobasal view; R, patch of articulated scales from venter in posterocrown view; S, small scale from head region in posterior view; T, small scale from head region in posterocrown view. Scale bars: 0.1 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 17. — Euthacanthus macnicoli Powrie, 1864 NMS G.2010.7.39, thin sections of scales from a large partial articulated specimen from Tillywhandland:A, a group of scales in various orientations;B-H, sections through individual scales;B, vertical longitudinal section with six crown growth zones;C, vertical transverse section with three crown growth zones;D, crown horizontal section with eight crown growth zones;E, vertical transverse section through posterior part of scale with at least eight growth zones;F, vertical transverse sec- tion through the anterior part of the scale with five crown growth zones; G, vertical longitudinal section with seven crown growth zones;" ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 18. — Euthacanthus macnicoli Powrie, 1864 NMS G.2010.7.43, a coprolitic/regurgitate patch of scales from a large individual from Tillywhandland: A, complete specimen; B-F, separate flank scales in matrix, viewed under a light microscope;G-J, SEM images of specialized scales; G, stellate head tessera in crown view; H, stellate head tessera in lateral view; I, fractured tuberculated head scale in anterior view; J, transitional or umbellate scale in anterocrown view; K-Q, thin sections of scales; K, crown horizontal section of head scale or tessera; L, vertical longitudinal section with three crown growth zones with a vertical transverse section of a small Mesacanthus mitchelli Egerton,1860 scale above anterior crown (to right);M, vertical transverse section of scale with six crown growth" ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIG. 19. — Euthacanthus macnicoli Powrie, 1864 Reconstruction of the whole fish in lateral view, much modified after Watson (1937: fig. 4A)." ; fabio:hasRepresentation ; a fabio:Figure . dwc:locality "National Museums of Scotland. Furthermore" ; dwc:recordedBy "Over & The" ; trt:gbifOccurrenceId "3120747302" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NMS" ; dwc:eventDate "1967-12-05" ; dwc:recordedBy "Apart & Turin Hill" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3120747303" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NHMP, NMS" ; dwc:eventDate "2013-08-01" ; dwc:recordedBy "Tillywhandland" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3120747301" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NMS" ; dwc:recordedBy "Duntrune" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3120747304" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NMS" ; dwc:recordedBy "Balruddery Den" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3120747305" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NMS, NMSG" ; dwc:locality "Of" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3120747306" ; trt:httpUri ; a dwc:MaterialCitation .