@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , , ; dc:creator "Matalin, Andrey V.; Wiesner, Jürgen" ; dc:title "Therates fruhstorferi W. Horn 1902" ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; trt:augmentsTaxonConcept ; trt:deprecates , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "433" ; bibo:issue "5" ; bibo:journal "Zootaxa" ; bibo:pubDate "2023-03-22" ; bibo:startPage "401" ; bibo:volume "5256" ; dc:creator "Matalin, Andrey V.; Wiesner, Jürgen" ; dc:date "2023" ; dc:title "Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)" ; fabio:hasPart , , , , , , , ; a fabio:JournalArticle . dwc:authority "W. Horn, 1902" ; dwc:authorityName "W. Horn" ; dwc:authorityYear "1902" ; dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Therates" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Horn, 1902" ; dwc:species "fruhstorferi" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "W. Horn, 1902" ; dwc:authorityName "W. Horn" ; dwc:authorityYear "1902" ; dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Therates" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Horn, 1902" ; dwc:species "fruhstorferi" ; dwc:subSpecies "fruhstorferi" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "W. Horn, 1913" ; dwc:authorityName "W. Horn" ; dwc:authorityYear "1913" ; dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Therates" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Horn, 1913" ; dwc:species "fruhstorferi" ; dwc:subSpecies "vitalisi" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Wiesner & Bi" ; dwc:baseAuthorityYear "2014" ; dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Therates" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Wiesner & Bi, 2014)" ; dwc:species "vitalisi" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Therates" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "fruhstorferi" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Therates" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Therates" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:species "fruhstorferi" ; dwc:subSpecies "fruhstorferi" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Therates" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:species "fruhstorferi" ; dwc:subSpecies "vitalisi" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Therates" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "vitalisi" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 50–57. Therates fruhstorferi ssp., labrum: 50–55—T. fruhstorferi fruhstorferi (50, 51—Vietnam, Montes Mauson; 52—Vietnam, Na Hang; 53—Vietnam, Tam Dao; 54, 55—Laos, Ban Nape); 56, 57—T. fruhstorferi australis ssp. n. (56—Vietnam, Kon Ka Kinh N.P.; 57—Vietnam, Pak Khe River); 50, 52, 54, 56—males; 51, 53, 55, 57—females; 50, 51—paralectotypes; 56—holotype; 57—paratype." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 136–151. Therates fruhstorferi fruhstorferi, variability of aedeagus, left lateral view: 136–143—Vietnam, Phu Tho Province, ~90 km W of Hanoi, Xuan Son Nat. Park, h = 400–700 m, 21o07’01’’N 104o56’39’’E, 6–15.VI.2014; 144–151— Vietnam, Vinh Phu Province, Tam Dao, different years (144, 145—3–11.VI.1985; 146—27.V–2.VI.1986; 147—2–11.VI.1989; 148—6–10.V.1990; 149—12.VI.1994; 150, 151—9–19.V.1996). Scales in mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 175–178. Morphometric features of Therates spp: 175, 177—relation of the proportions EL/PL and HTbL/HTaL (green rhombus—T. pseudovitalisi sp. n.; blue circles—T. fruhstorferi; open symbols—males; solid symbols—females); 176, 178—relation of the proportions HTbL/HTaL (small quadrates—medians; rectangles—25–75 percentiles; whiskers—min–max; significantly different samples are marked with different lower letters, according to Mann–Whitney U-test, p <0.05 (176—U = 112.0, z = 5.08, p = 0.000; 178—U = 23.5, z = 3.37, p = 0.0007)); 175, 176—all specimens; 177, 178—males." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png." ; fabio:hasRepresentation ; a fabio:Figure . dwc:collectionCode "XX" ; dwc:eventDate "2006-07-24" ; dwc:locality "Hong Mu Da Shan" ; dwc:verbatimElevation "1851" ; trt:gbifOccurrenceId "4063374323" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "ZIN" ; dwc:eventDate "1963-07-05" ; dwc:locality "Ha Giang" ; dwc:recordedBy "O. Kabakov" ; dwc:verbatimElevation "500" ; trt:gbifOccurrenceId "4063374407" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "AA" ; dwc:eventDate "1963-07-05" ; dwc:locality "Yen Bai" ; dwc:recordedBy "O. Kabakov" ; dwc:verbatimElevation "500" ; trt:gbifOccurrenceId "4063374301" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "AA" ; dwc:eventDate "1963-07-05" ; dwc:locality "Ha Giang" ; dwc:recordedBy "O. Kabakov" ; dwc:verbatimElevation "500" ; trt:gbifOccurrenceId "4063374392" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "AN, MSPU" ; dwc:eventDate "1996-10-06" ; dwc:locality "Na Hang" ; dwc:recordedBy "A Napolov & I. Roma" ; dwc:verbatimElevation "175" ; dwc:verbatimLatitude "21.066668" ; dwc:verbatimLongitude "105.36056" ; trt:gbifOccurrenceId "4063374408" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "ZIN" ; dwc:eventDate "1963-06-21" ; dwc:locality "Thai Nguen Prov." ; dwc:recordedBy "O. Kabakov" ; dwc:verbatimElevation "300" ; dwc:verbatimLatitude "21.066668" ; dwc:verbatimLongitude "105.36056" ; trt:gbifOccurrenceId "4063374316" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "ZIN" ; dwc:eventDate "1963-08-30" ; dwc:locality "Tam Dao" ; dwc:recordedBy "O. Kabakov" ; dwc:verbatimElevation "900" ; dwc:verbatimLatitude "21.066668" ; dwc:verbatimLongitude "105.36056" ; trt:gbifOccurrenceId "4063374410" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "JW" ; dwc:eventDate "1985-11-06" ; dwc:locality "Tam Dao" ; dwc:recordedBy "Ploka & Visa" ; dwc:verbatimElevation "175" ; dwc:verbatimLatitude "21.066668" ; dwc:verbatimLongitude "105.36056" ; trt:gbifOccurrenceId "4063374371" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "JW" ; dwc:eventDate "1989-11-06" ; dwc:locality "Vinh Phu Prov." ; dwc:recordedBy "Kuban" ; dwc:verbatimElevation "175" ; dwc:verbatimLatitude "21.066668" ; dwc:verbatimLongitude "105.36056" ; trt:gbifOccurrenceId "4063374367" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "JW" ; dwc:eventDate "1986-06-02" ; dwc:locality "Vinh Phu Prov." ; dwc:recordedBy "Jan Horak" ; dwc:verbatimElevation "175" ; dwc:verbatimLatitude "21.066668" ; dwc:verbatimLongitude "105.36056" ; trt:gbifOccurrenceId "4063374395" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "JW" ; dwc:eventDate "1990-10-05" ; dwc:locality "Vinh Phu Prov." ; dwc:recordedBy "Jan Horak" ; dwc:verbatimElevation "175" ; dwc:verbatimLatitude "21.066668" ; dwc:verbatimLongitude "105.36056" ; trt:gbifOccurrenceId "4063374383" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "JW" ; dwc:eventDate "1989-05-24" ; dwc:locality "Vinh Phu Prov." ; dwc:recordedBy "Jan Strnad" ; dwc:verbatimElevation "175" ; dwc:verbatimLatitude "21.066668" ; dwc:verbatimLongitude "105.36056" ; trt:gbifOccurrenceId "4063374390" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "JW" ; dwc:eventDate "1990-05-06" ; dwc:recordedBy "M. 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Skale" ; dwc:verbatimElevation "1100" ; dwc:verbatimLatitude "22.564722" ; dwc:verbatimLongitude "105.88139" ; trt:gbifOccurrenceId "4063374409" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2014-05-09" ; dwc:locality "Nui Pia Oac Nature Reserve" ; dwc:recordedBy "A. Skale" ; dwc:verbatimElevation "1075" ; dwc:verbatimLatitude "22.631945" ; dwc:verbatimLongitude "105.36056" ; trt:gbifOccurrenceId "4063374317" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V, R" ; dwc:eventDate "2015-05-18" ; dwc:locality "Tay Yen Tu Nature Reserve" ; dwc:recordedBy "Gerstmeier" ; dwc:verbatimElevation "200" ; dwc:verbatimLatitude "21.1805" ; dwc:verbatimLongitude "106.723785" ; trt:gbifOccurrenceId "4063374362" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2012-06-18" ; dwc:locality "Xuan Son National Park" ; dwc:recordedBy "L. Bartolozzi; S. Bambi; F. Fabiano & E. Orbach" ; dwc:verbatimElevation "500" ; trt:gbifOccurrenceId "4063374350" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2013-05-02" ; dwc:locality "Tam Dao National Park" ; dwc:recordedBy "A. Skale & A. Weigel" ; dwc:verbatimElevation "850" ; dwc:verbatimLatitude "21.45" ; dwc:verbatimLongitude "105.63333" ; trt:gbifOccurrenceId "4063374381" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2013-06-01" ; dwc:locality "Tam Dao National Park" ; dwc:recordedBy "L. Bartolozzi; S. Bambi; F. Cianferoni; G. Mazza & E. Orbach" ; dwc:verbatimElevation "950" ; trt:gbifOccurrenceId "4063374354" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2014-06-09" ; dwc:locality "Tam Dao" ; dwc:recordedBy "M. 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