@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , , , , , , , , , , , , , , , , , , , , ; dc:creator "Bellardini, Flavio; Coria, Rodolfo A; Pino, Diego A; Windholz, Guillermo J; Baiano, Mattia A; Martinelli, Augustin G" ; dc:title "Ligabuesaurus leanzai BONAPARTE, GONZALEZ RIGA & APESTEGUIA 2006" ; trt:augmentsTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "1393" ; bibo:issue "4" ; bibo:journal "Zoological Journal of the Linnean Society" ; bibo:pubDate "2022-12-01" ; bibo:startPage "1333" ; bibo:volume "196" ; dc:creator "Bellardini, Flavio; Coria, Rodolfo A; Pino, Diego A; Windholz, Guillermo J; Baiano, Mattia A; Martinelli, Augustin G" ; dc:date "2022" ; dc:title "Osteology and phylogenetic relationships of Ligabuesaurus leanzai (Dinosauria: Sauropoda) from the Early Cretaceous of the Neuquén Basin, Patagonia, Argentina" ; fabio:hasPart , , , , , , , , , , , , , , , , , , , , , , , , , ; a fabio:JournalArticle . dwc:authority "BONAPARTE, GONZALEZ RIGA & APESTEGUIA, 2006" ; dwc:authorityName "BONAPARTE, GONZALEZ RIGA & APESTEGUIA" ; dwc:authorityYear "2006" ; dwc:class "Reptilia" ; dwc:genus "Ligabuesaurus" ; dwc:kingdom "Animalia" ; dwc:order "Saurischia" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Bonaparte, Gonzalez Riga & Apesteguia, 2006" ; dwc:species "leanzai" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Reptilia" ; dwc:genus "Ligabuesaurus" ; dwc:kingdom "Animalia" ; dwc:order "Saurischia" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "leanzai" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:genus "Ligabuesaurus" ; dwc:kingdom "Animalia" ; dwc:order "Saurischia" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:kingdom "Animalia" ; dwc:order "Saurischia" ; dwc:phylum "Chordata" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "Figure 4. The somphospondylan sauropod Ligabuesaurus leanzai from Cerro de los Leones (Neuquén Province, Patagonia, Argentina). A, skeletal reconstruction based on the holotype (MCF-PVPH-233) and referred material (MCF-PVPH-261) from quarry no. 4, plus the newly referred specimens from quarry no. 3 (MCF-PVPH-228, MCF-PVPH-744 and MCF- PVPH-908). B, life restoration of Ligabuesaurus leanzai as a non-titanosaurian somphospondylan by J. L. Blanco." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 1. Location map. A, satellite map of northern Patagonia (Argentina), showing the approximate limits of Neuquén Basin (white dashed line). B, the Cerro de los Leones locality is ~10 km to the south-west of Picún Leufú city, to the south of the Neuquén Province (white crossed pickaxes). C, satellite map of the main quarries opened on the southern flank of the Cerro de los Leones, where outcropping of the fluvial sediments of the Lohan Cura Formation (Albian) occurs." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 5. New autapomorphic feature and revised diagnosis of Ligabuesaurus leanzai. A, Photograph and line drawings of the left coracoid MCF-PVPH-261/05 in lateral view. The quadrangular ventral half of the coracoid in lateral view is here proposedasauniquelyderivedfeatureofLigabuesaurusleanzainotpresentinanyothersauropods.B–K,comparativeoutlines of sauropod coracoids: B, Neuquensaurus MLP-Ly-14; C, Saltasaurus PVL-4017-100; D, Quetecsaurus UNCUYO-LD-300.15; E, Opisthocoelicaudia ZPAL-MgD-I/48; F, Dreadnoughtus MPM-PV-1156; G, Tapuiasaurus MZSP-PV-807; H, Uberabatitan CPP-1109-UrHo; I, Suuwassea ANS 21122; J, Patagotitan MPEF-PV-3400/24; K, Zby ML 368. L–N, line drawings of the posterior cervical vertebra MCF-PVPH-233/02 (L) and anterior dorsal vertebrae MCF-PVPH-233/03 (M) and MCF- PVPH-908 (N), showing the development of the prespinal lamina on the anterior surface of the neural spine (green triangle). B–K modified from González Riga et al. (2019). Not to scale. Scale bar: 10 cm in L–N." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 2. Lithological profile. A schematic log of the the lower section of the Cullin Grande Member (Bajada del Agrio Group, Lohan Cura Formation, Lower Cretaceous, Albian) that outcrops at the Cerro de los Leones locality (modified from Martinelli et al., 2007). Abbreviations: CS, crevasse channel; FF, floodplain fines; FL, fossiliferous level; LA, lateral accretion; LS, laminated sand sheets; LV, levee; SB, sandy bedforms. Architectural element codes follow Miall (1996)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 3. Original picture of quarry no. 4 during the fieldwork of 2004 (A) and a quarry map with tentative arrangement of the holotype (in red) and referred material (in blue) of Ligabuesaurus leanzai (B). Abbreviations: ant, anterior; s/n, without collection number; v, vertebra." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 6. Photographs and line drawings of the teeth of Ligabuesaurus leanzai. A, ten maxillary teeth and fragments of unerupted teeth in sedimentary matrix MCF-PVPH-233/01 in lingual view. B–F, isolated maxillary tooth MCF-PVPH-744 in lingual (B), apical (C), labial (D), mesial (E) and distal (F) views. Abbreviations: awf, apical wear facet; cc, cingular cusp; de, dentine surface; e, enamel surface; mg, mesial groove; mwf, mesial wear facet; rt, replacement tooth fragment; tc, tooth crown; tr, tooth root; t-1/10, tooth no. 1 and no. 10; we, wrinkled enamel surface. Scale bar: 3 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 7. A, detail ofthe microwear pattern on the apicalwear facet of the maxillary tooth MCF-PVPH-744 of Ligabuesaurus leanzai. Most of the pits and coarse grooves are evident with a binocular microscope (α) on the dentine exposed surface, whereas the scanning electron photomicrographs (β, γ, δ and ε) show abundant fine scars in both the enamel and dentine surfaces. Abbreviations: g, coarse groove; p, pit; s, fine scar." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 8. Photographs and line drawings of the middle cervical vertebra MCF-PVPH-261/16 of Ligabuesaurus leanzai in dorsal (A), ventral (B) and left lateral (C) views. Abbreviations: acdl, anterior centrodiapophyseal lamina; apo, accessory pneumatic opening; cpol, centropostzygapophyseal lamina; cprl, centroprezygapophyseal lamina; di, diapophysis; nc, neural canal; pa, parapophysis; pcdl, posterior centrodiapophyseal lamina; pl, pleurocoel; prdl, prezygodiapophyseal lamina; prz, prezygapophysis; pvr, posteroventral ridges; spdl, spinodiapophyseal lamina; sprl, spinoprezygapophyseal lamina. Scale bar: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 9. Photographs and line drawings of the posterior cervical vertebra MCF-PVPH-261/01 of Ligabuesaurus leanzai in anterior (A), posterior (B) and left lateral (C) views. Abbreviations: aas, anterior articular surface; apf, anterior pneumatic fossa; cprl, centroprezygapophyseal lamina; di, diapophysis; pa, parapophysis; pas, posterior articular surface; ppf, posterior pneumatic fossa; prz, prezygapophysis; r?, fragment of rib. Scale bar: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 10. Photographs and line drawings of the posterior cervical vertebra MCF-PVPH-261/02 of Ligabuesaurus leanzai in anterior (A), posterior (B) and right lateral (C) views. Abbreviations: apf, anterior pneumatic fossa; cprl, centroprezygapophyseal lamina; di, diapophysis; nc, neural canal; pa, parapophysis; ppf, posterior pneumatic fossa; prz, prezygapophysis. Scale bar: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 11. Photographs and line drawings of the posterior cervical vertebra MCF-PVPH-228/01 and MCF-PVPH-228/02 of Ligabuesaurus leanzai in anterior (A) view. Abbreviations: aas, anterior articular surface; cpol, centropostzygapophyseal lamina; di, diapophysis; poz, postzygapophysis; prdl, prezygodiapohyseal lamina; prz, prezygapophysis; sprl, spinoprezygapophyseal lamina; tprl, intraprezygapophyseal lamina. Scale bar: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 12. Photographs and line drawings of the posterior cervical vertebra MCF-PVPH-233/02 of Ligabuesaurus leanzai in anterior (A), posterior (B) and right lateral (C) views. Abbreviations: acdl, anterior centrodiapophyseal lamina; cpol, centropostzygapophyseal lamina; cprl, centroprezygapophyseal lamina; di, diapophysis; lat. sprl, lateral spinoprezygapophyseal lamina; med. sprl, medial spinoprezygapophyseal lamina; nc, neural canal; ns, neural spine; pa, parapophysis; pcdl, posterior centrodiapophyseal lamina; pl, pleurocoel; podl, postzygodiapophyseal lamina; posl, postspinal lamina; poz, postzygapophysis; prdl, prezygodiapophyseal lamina; prz, prezygapophysis; r?, fragment of rib; spdl, spinodiapophyseal lamina; spol, spinopostzygapophyseal lamina; sprl, spinoprezygapophyseal lamina. Scale bar: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 13. Photographs and line drawings of the anterior dorsal vertebra MCF-PVPH-233/03 of Ligabuesaurus leanzai in anterior (A), posterior (B) and left lateral (C) views. Abbreviations: acdl, anterior centrodiapophyseal lamina; cpol, centropostzygapophyseal lamina; cprl, centroprezygapophyseal lamina; di, diapophysis; lat. sprl, lateral spinoprezygapophyseal lamina; med. sprl, medial spinoprezygapophyseal lamina; nc, neural canal; pa, parapophysis; pcdl, posterior centrodiapophyseal lamina; pl, pleurocoel; poz, postzygapophysis; prsl, prespinal lamina; prz, prezygapophysis; spdl, spinodiapophyseal lamina; spol, spinopostzygapophyseal lamina; tprl, intraprezygapophyseal lamina. Scale bar: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 14. Photograph and line drawing of the anterior dorsal vertebra MCF-PVPH-908 of Ligabuesaurus leanzai in anterior view.Abbreviations:acdl, anterior centrodiapophyseal lamina; cprl, centroprezygapophyseallamina; di, diapophysis; lat. sprl, lateral spinoprezygapophyseal lamina; med. sprl, medial spinoprezygapophyseal lamina; pa, parapophysis; prsl, prespinal lamina; prz, prezygapophysis; spdl, spinodiapophyseal lamina; sprf, spinoprezygapophyseal fossa; tprl, intraprezygapophyseal lamina. Scale bar: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 15. Photograph and line drawings of the mid-posterior dorsal vertebrae MCF-PVPH-233/04 and MCF- PVPH-233/05 of Ligabuesaurus leanzai in left lateral view. Abbreviations: acdl, anterior centrodiapophyseal lamina; acpl, anterior centroparapophyseal lamina; pa, parapophysis; pcdl, posterior centrodiapophyseal lamina; pcpl, posterior centroparapophyseal lamina; pl, pleurocoel; poz, postzygapophysis; ppdl, parapophysisdiapophyseal lamina; prz, prezygapophysis. Scale bar: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 16. Photographs and line drawings of the mid-posterior dorsal vertebrae MCF-PVPH-228/03 and MCF- PVPH-228/04 of Ligabuesaurus leanzai in left lateral (A), anterior (B) and posterior (C) views. Abbreviations: acdl, anterior centrodiapophyseal lamina; acpl, anterior centroparapophyseal lamina; cpol, centropostzygapophyseal lamina; hypa, hypantrum; pcdl, posterior centrodiapophyseal lamina; pcpl, posterior centroparapophyseal lamina; pl, pleurocoel; poz, postzygapophysis; ppdl, parapophysisdiapophyseal lamina; prz, prezygapophysis; spdl, spinodiapophyseal lamina; spol, spinopostzygapophyseal lamina; sprl, spinoprezygapophyseal lamina. Scale bar: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 17. Photographs and line drawings of the posterior dorsal vertebrae MCF-PVPH-233/06 and MCF-PVPH-233/07 of Ligabuesaurus leanzai in anterior (A) and posterior (B) views. Abbreviations: acc. spdl, accessory spinodiapophyseal lamina; cpol, centropostzygapophyseal lamina; di, diapophysis; hypo, hyposphene; pcdl, posterior centrodiapophyseal lamina; poz, postzygapophysis; prsl, prespinal lamina; prz, prezygapophysis; spdl, spinodiapophyseal lamina; sprl, spinoprezygapophyseal lamina. Scale bar: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 18. Photographs and line drawings of the caudal vertebra and dorsal rib of Ligabuesaurus leanzai. A–E, anterior caudal vertebra MCF-PVPH-261/15 in anterior (A), posterior (B), left lateral (C), dorsal (D) and ventral (E) views. F, G, dorsal rib MCF-PVPH-228/10 in medial view (F), with a detail of the internal pneumatic structure on the process of the capitulum (G). Abbreviations: aas, anterior articular surface; cap, capitulum; nc, neural canal; pas, posterior articular surface; rs, rib shaft; tp, transverse process; tub, tuberculum; vf, vascular foramen. Scale bars: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 19. Pectoral girdle elements of Ligabuesaurus leanzai. A, Right scapula MCF-PVPH-228/11 in lateral view. B, left scapula MCF-PVPH-233/08 in lateral view. C–E, left coracoid MCF-PVPH-261/05 in lateral (C), distal (D) and medial views (E). F–H, right coracoid MCF-PVPH-261/06 in medial (F), distal (G) and lateral (H) views.Abbreviations:af, anterior fossa; ar, acromial ridge; br, attachment surface for muscle biceps brachii; cbb, attachment surface for muscle coracobrachialis brevis; f, coracoid foramen; gl, glenoid; ign, infraglenoid notch; lev sc, attachment surface for levator scapulae; mvp, medioventral process; pf, posterior fossa; scs, scapulocoracoid articular surface; sr, scapular ridge. Scale bars: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 20. Photographs and line drawings of the forelimb elements of Ligabuesaurus leanzai. A–D, left humerus MCF- PVPH-233/10 in anterior (A), proximal (B), distal (C) and posterior (D) views. E, F, distal half of the right humerus MCF- PVPH-233/12 in anterior (E) and distal (F) views. G–I, distal half of the left radius MCF-PVPH-261/07 in posterior (G), distal (H) and lateral (I) views. Abbreviations: ac, accessory condyles; anf, anconeal fossa; das, distal articular surface; dpc, deltopectoral crest; dpf, deltopectoral fossa; hh, humeral head; ltf, lateral triceps fossa; mtf, medial triceps fossa; rc, radial condyle; uas, ulnar articular surface; uc, ulnar condyle. Scale bars: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 21. Photographs and line drawings of the manus of Ligabuesaurus leanzai. A–D, right metacarpal II MCF- PVPH-233/13 in medial (A), proximal (B), distal (C) and lateral (D) views. E, distal epiphysis of the left metacarpal II MCF-PVPH-233/15 in distal view. F–I, right metacarpal III MCF-PVPH-233/14 in lateral (F), proximal (G), distal (H) and medial (I) views. J, distal epiphysis of the left metacarpal IV MCF-PVPH-233/16 in distal view. Abbreviations: am, anterior margin; lm, lateral margin; McI, articular surface for metacarpal I; McII, articular surface for metacarpal II; McIII, articular surface for metacarpal III; McIV, articular surface for metacarpal IV; pm, posterior margin; vlr, ventrolateral ridge. Scale bars: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure22. Photographsand linedrawingsofpelvicgirdleelementsofLigabuesaurus leanzai.A,leftilium MCF-PVPH-261/08 in lateral view. B–D, proximal half of the right pubis MCF-PVPH-261/10 in lateral (B), proximal (C) and medial (D) views. E, F, proximal half of the left pubis MCF-PVPH-261/09 in lateral (E) and proximal (F) views. G, reconstruction of right pubis in lateral view, assembling the proximal and distal halves MCF-PVPH-261/10 and MCF-PVPH-261/11. H–J, distal half of the right pubis MCF-PVPH-261/11 in anterior (H), lateral (I) and distal (J) views. Abbreviations: acet, acetabulum; am, anterior margin; ias, ischiatic articular surface; ip, ischiatic peduncle; lm, lateral margin; mm, medial margin; of, obturator foramen; poap, postacetabular process; pp, pubic peduncle; prap, preacetabular process. Scale bars: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 23. Photographs and line drawings of the femora of Ligabuesaurus leanzai. A–C, right femur MCF-PVPH-233/17 in anterior (A), posterior (B) and distal (C) views. D–G, left femur MCF-PVPH-261/12 in anterior (D), distal (E), posterior (F) and medial (G) views. Abbreviations: ec, epicondyle; fc, fibular condyle; fh, femoral head; ft, fourth trochanter; ftr, femoral trochlea; gt, great trochanter; icf, intracondylar fossa; icg, intracondylar groove; lb, lateral bulge; lic, linea intermuscularis cranialis; tc, tibial condyle. Scale bars: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 24. Photographs and line drawings of the hindlimb elements of Ligabuesaurus leanzai. A–E, right tibia MCF- PVPH-233/18 in anterior (A), lateral (B), posterior (C), proximal (D) and distal (E) views. F–I, proximal epiphysis of the left tibia MCF-PVPH-261/13 in proximal (F), posterior (G), lateral (H) and medial (I) views.J–M, right fibula MCF-PVPH-233/19 in medial (J), anterior (K), proximal (L) and distal (M) views. N, O, proximal epiphysis of the left fibula MCF-PVPH-261/14 in proximal (N) and medial (O) views.Abbreviations:amc, anteromedial crest; amf, anteromedial fossa; ap, anterior process; asap, articular surface for the ascending process; asp, astragal process; at, anterior trochanter; cc, cnemial crest; cf,cnemial fossa; lt, lateral trochanter; mi, insertion surface for muscle iliofibularis; pp, posterior process; tas, tibial articular surface. Scale bars: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 25. Photographs and line drawings of the right astragalus MCF-PVPH-233/20 of Ligabuesaurus leanzai in anterior (A), dorsal (B), lateral (C), posterior (D), ventral (E) and medial (F) views. Abbreviations: apr, ascending process of astragalus; cas, calcaneal articular surface; cpf, crest of posterior fossa; das, distal articular surface; fas, fibular articular surface; ilg, intertarsal ligament groove; pf, posterior fossa; tas, tibial articular surface. Scale bar: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 26. Photographs and line drawings of the pedal elements of Ligabuesaurus leanzai. A–C, right metatarsal I MCF- PVPH-233/21 in posterior (A), proximal (B) and distal (C) views. D–F, right metatarsal II MCF-PVPH-233/22 in posterior (D), proximal (E) and distal (F) views. G–I, right metatarsal III MCF-PVPH-233/23 in posterior (G), proximal (H) and distal (I) views. J, K, right metatarsal IV MCF-PVPH-233/24 in posterior (J) and proximal (K) views. L, M, right metatarsal V MCF- PVPH-233/25 in posterior (L) and proximal (M) views. N, right proximal phalanx I-1 MCF-PVPH-233/26 in posterior view. O, right ungueal phalanx MCF-PVPH-233/27 in medial view. P, right proximal phalanx II-1 MCF-PVPH-233/28 in posterior view. Abbreviations: I, articular surface for metatarsal I; II, articular surface for metatarsal II; III, articular surface for metatarsal III; icg, intercondylar groove; IV, articular surface for metatarsal IV; lc, lateral condyle; mdp, mediodistal process; mh, medial hollow; pas, proximal articular surface; V, articular surface for metatarsal V; vf, vascular foramen. Scale bars: 10 cm." ; fabio:hasRepresentation ; a fabio:Figure .