@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , ; dc:creator "Montoya-Molina, Santiago; Vaz-De, Fernando Z." ; dc:title "Dichotomius (Luederwaldtinia) belus" ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; trt:augmentsTaxonConcept ; trt:deprecates ; trt:publishedIn ; a trt:Treatment . bibo:endPage "64" ; bibo:journal "European Journal of Taxonomy" ; bibo:pubDate "2021-02-11" ; bibo:startPage "1" ; bibo:volume "734" ; dc:creator "Montoya-Molina, Santiago; Vaz-De, Fernando Z." ; dc:date "2021" ; dc:title "Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)" ; fabio:hasPart , ; a fabio:JournalArticle . dwc:authority "(Harold, 1880)" ; dwc:baseAuthorityName "Harold" ; dwc:baseAuthorityYear "1880" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Harold, 1880)" ; dwc:species "belus" ; dwc:subGenus "Luederwaldtinia" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Harold, 1880: 25" ; dwc:authorityName "Harold" ; dwc:authorityPageNumber "25" ; dwc:authorityYear "1880" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Pinotus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Harold, 1880" ; dwc:species "belus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "belus" ; dwc:subGenus "Luederwaldtinia" ; trt:hasParentName , ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:subGenus "Luederwaldtinia" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Pinotus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "belus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Pinotus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "Fig. 5. Dichotomius belus (Harold, 1880). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Anterior view of the shape of the cephalic carina. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Lateral view, showing the pronotum and the cephalic carina flattened antero-posteriorly and curved anteriorly." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." ; fabio:hasRepresentation ; a fabio:Figure . dwc:collectionCode "MNHN" ; dwc:locality "Medellin" ; dwc:recordedBy "F. Z. Vaz-de-Mello" ; dwc:typeStatus "lectotype" ; trt:gbifOccurrenceId "3037418511" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "COLOMBIA" ; dwc:recordedBy "F. Z. Vaz-de-Mello" ; dwc:typeStatus "paralectotype" ; trt:gbifOccurrenceId "3037418378" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "Medellin" ; dwc:recordedBy "F. Z. Vaz-de-Mello" ; trt:gbifOccurrenceId "3037418345" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "Medellin" ; dwc:recordedBy "F. Z. Vaz-de-Mello" ; dwc:typeStatus "paralectotype" ; trt:gbifOccurrenceId "3037418591" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "Medellin" ; dwc:recordedBy "F. Z. Vaz-de-Mello" ; dwc:typeStatus "paralectotype" ; trt:gbifOccurrenceId "3037418497" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "Ocana Landoldt" ; dwc:recordedBy "F. Z. Vaz-de-Mello" ; dwc:typeStatus "paralectotype" ; trt:gbifOccurrenceId "3037418602" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "Ambalema" ; dwc:recordedBy "F. Z. Vaz-de-Mello" ; dwc:typeStatus "paralectotype" ; trt:gbifOccurrenceId "3037418563" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1972-03-01" ; dwc:locality "COLOMBIA" ; dwc:recordedBy "S. and J. Peck" ; trt:gbifOccurrenceId "3037418538" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1964-04-22" ; trt:gbifOccurrenceId "3037418567" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1957-09" ; dwc:recordedBy "Gallego" ; trt:gbifOccurrenceId "3037418321" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1964-04-21" ; dwc:locality "Antioquia" ; trt:gbifOccurrenceId "3037418380" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1964-04-21" ; dwc:locality "Antioquia" ; dwc:recordedBy "Medellin" ; trt:gbifOccurrenceId "3037418342" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:recordedBy "Medellin" ; trt:gbifOccurrenceId "3037418617" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1962-12-15" ; dwc:locality "Medellin" ; trt:gbifOccurrenceId "3037418467" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1947-04" ; dwc:locality "Encorredo" ; dwc:recordedBy "Gallego" ; trt:gbifOccurrenceId "3037418351" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "2015-03" ; dwc:locality "Puerto Berrio" ; dwc:verbatimElevation "300" ; dwc:verbatimLatitude "6.4919996" ; dwc:verbatimLongitude "-74.577736" ; trt:gbifOccurrenceId "3037418441" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "2012-10" ; dwc:locality "Vda. La Carlota" ; dwc:recordedBy "J. Leon" ; dwc:verbatimElevation "590" ; dwc:verbatimLatitude "6.518" ; dwc:verbatimLongitude "-74.582" ; trt:gbifOccurrenceId "3037418426" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "2012-10" ; dwc:locality "Vda. La Carlota" ; dwc:recordedBy "J. Leon" ; dwc:verbatimElevation "590" ; dwc:verbatimLatitude "6.518" ; dwc:verbatimLongitude "-74.582" ; trt:gbifOccurrenceId "3037418566" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "1996-07" ; dwc:locality "Rio Cane" ; dwc:recordedBy "F. Escobar" ; dwc:verbatimElevation "2200" ; trt:gbifOccurrenceId "3037418377" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "1996-09" ; dwc:locality "Villa de Leyva" ; dwc:recordedBy "M. Alvares" ; dwc:verbatimElevation "2200" ; trt:gbifOccurrenceId "3037418394" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "1998-08" ; dwc:locality "Claustro San Agustin" ; dwc:recordedBy "K. Schutze" ; dwc:verbatimElevation "2200" ; trt:gbifOccurrenceId "3037418492" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "1998-08" ; dwc:locality "Claustro San Agustin" ; dwc:recordedBy "K. Schutze" ; dwc:verbatimElevation "2200" ; dwc:verbatimLatitude "2.0" ; dwc:verbatimLongitude "-76.95" ; trt:gbifOccurrenceId "3037418526" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "ECC" ; dwc:eventDate "2010-01" ; dwc:locality "Area de reserva Lerma" ; dwc:recordedBy "W. Guzman" ; dwc:verbatimElevation "1700" ; dwc:verbatimLatitude "2.0" ; dwc:verbatimLongitude "-76.95" ; trt:gbifOccurrenceId "3037418520" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "ECC" ; dwc:eventDate "1995-01" ; dwc:locality "Fusagasuga" ; dwc:recordedBy "A. Amezquita & A. Lopera" ; dwc:verbatimElevation "1900" ; trt:gbifOccurrenceId "3037418423" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1962-09-06" ; dwc:locality "Fusagasuga" ; trt:gbifOccurrenceId "3037418415" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1991-04-14" ; dwc:locality "alrededores colegio Fatima" ; dwc:recordedBy "A. Mera & R. Lozano" ; dwc:verbatimElevation "1500" ; trt:gbifOccurrenceId "3037418468" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "1997-10" ; dwc:locality "Juan de Rio Seco" ; dwc:recordedBy "H. Pulido-Riveros" ; dwc:verbatimElevation "1300" ; trt:gbifOccurrenceId "3037418516" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:locality "Bogota" ; trt:gbifOccurrenceId "3037418483" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "2002-09" ; dwc:locality "Bogota" ; dwc:recordedBy "M. Ospina" ; dwc:verbatimElevation "1000" ; dwc:verbatimLatitude "2.2958333" ; dwc:verbatimLongitude "-75.59361" ; trt:gbifOccurrenceId "3037418392" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:locality "La Guajira" ; dwc:recordedBy "S. Montoya-M." ; dwc:verbatimLatitude "10.8248" ; dwc:verbatimLongitude "-72.8246" ; trt:gbifOccurrenceId "3037418524" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1972-03-02" ; dwc:locality "La Guajira" ; dwc:recordedBy "S. Montoya-M." ; dwc:verbatimLatitude "10.8248" ; dwc:verbatimLongitude "-72.8246" ; trt:gbifOccurrenceId "3037418438" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1972-03-02" ; dwc:locality "La Guajira" ; dwc:recordedBy "S. Montoya-M." ; dwc:verbatimLatitude "10.8248" ; dwc:verbatimLongitude "-72.8246" ; trt:gbifOccurrenceId "3037418408" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "2008-08-22" ; dwc:locality "Potrero Ganadero" ; dwc:recordedBy "A. N. L. Hidalgo" ; dwc:samplingProtocol "pitfall" ; dwc:verbatimElevation "2500" ; dwc:verbatimLatitude "1.3772222" ; dwc:verbatimLongitude "-77.21917" ; trt:gbifOccurrenceId "3037418445" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "1994-04" ; dwc:locality "La Planada." ; dwc:recordedBy "F. Escobar" ; dwc:verbatimElevation "1800" ; trt:gbifOccurrenceId "3037418571" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "2013-10" ; dwc:locality "Vda. Algodonal" ; dwc:recordedBy "J. Salazar & J. Castillo" ; dwc:verbatimElevation "628" ; dwc:verbatimLatitude "1.6683334" ; dwc:verbatimLongitude "-77.2875" ; trt:gbifOccurrenceId "3037418532" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "2013-10" ; dwc:locality "Vda. Algodonal" ; dwc:recordedBy "J. Salazar & J. Castillo" ; dwc:verbatimElevation "628" ; dwc:verbatimLatitude "1.6683334" ; dwc:verbatimLongitude "-77.2875" ; trt:gbifOccurrenceId "3037418431" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "2013-10" ; dwc:locality "Vda. Algodonal" ; dwc:recordedBy "J. Salazar & J. 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Sarmiento" ; dwc:verbatimElevation "2280" ; dwc:verbatimLatitude "6.1077776" ; dwc:verbatimLongitude "-73.05212" ; trt:gbifOccurrenceId "3037418547" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "1998-02-02" ; dwc:locality "SFF Guanenta Alto rio Fonce" ; dwc:recordedBy "J. Martinez" ; dwc:verbatimElevation "2450" ; trt:gbifOccurrenceId "3037418365" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "BMNH 1918-14" ; dwc:collectionCode "IAvH" ; dwc:eventDate "1998-02-02" ; dwc:locality "SFF Guanenta Alto rio Fonce" ; dwc:recordedBy "J. Martinez" ; dwc:verbatimElevation "2450" ; trt:gbifOccurrenceId "3037418372" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "1995-11" ; dwc:locality "Loma Santo Tomas" ; dwc:recordedBy "F. Escobar" ; dwc:samplingProtocol "excr. humano" ; dwc:verbatimElevation "300" ; dwc:verbatimLatitude "5.0919447" ; dwc:verbatimLongitude "-49.866665" ; trt:gbifOccurrenceId "3037418607" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IAvH" ; dwc:eventDate "1995-11" ; dwc:locality "Loma Santo Tomas" ; dwc:recordedBy "F. Escobar" ; dwc:samplingProtocol "excr. humano" ; dwc:verbatimElevation "300" ; dwc:verbatimLatitude "5.0919447" ; dwc:verbatimLongitude "-49.866665" ; trt:gbifOccurrenceId "3037418487" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "ECC" ; dwc:eventDate "2003-09-28" ; dwc:locality "Bosque Municipal" ; dwc:recordedBy "V. Fuentes" ; dwc:verbatimElevation "650" ; trt:gbifOccurrenceId "3037418496" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "ECC" ; dwc:eventDate "2015-04" ; dwc:locality "Valle del Cauca" ; dwc:recordedBy "A. 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