@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , , , , , , , , , ; dc:creator "Rahayu, Dwi Listyo; Ng, Peter K. L." ; dc:title "Lyphira heterograna" ; dwc:basisOfRecord , , , , , , , , ; trt:augmentsTaxonConcept ; trt:citesTaxonName ; trt:deprecates , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "392" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:pubDate "2024-07-04" ; bibo:startPage "358" ; bibo:volume "5476" ; dc:creator "Rahayu, Dwi Listyo; Ng, Peter K. L." ; dc:date "2024" ; dc:title "On the genus Lyphira Galil, 2009 (Crustacea: Brachyura: Leucosiidae), with descriptions of four new species from the West Pacific" ; fabio:hasPart , , , , , , , , , , , , , , ; a fabio:JournalArticle . dwc:authority "(Ortmann, 1892)" ; dwc:baseAuthorityName "Ortmann" ; dwc:baseAuthorityYear "1892" ; dwc:class "Malacostraca" ; dwc:family "Leucosiidae" ; dwc:genus "Lyphira" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Ortmann, 1892)" ; dwc:species "heterograna" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ortmann, 1892: 582" ; dwc:authorityName "Ortmann" ; dwc:authorityPageNumber "582" ; dwc:authorityYear "1892" ; dwc:class "Malacostraca" ; dwc:family "Leucosiidae" ; dwc:genus "Philyra" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ortmann, 1892" ; dwc:species "heterograna" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "De Haan" ; dwc:authorityYear "1841" ; dwc:class "Malacostraca" ; dwc:family "Leucosiidae" ; dwc:genus "Philyra" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "De Haan, 1841" ; dwc:species "pisum" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Shen, 1932: 18" ; dwc:authorityName "Shen" ; dwc:authorityPageNumber "18" ; dwc:authorityYear "1932" ; dwc:class "Malacostraca" ; dwc:family "Leucosiidae" ; dwc:genus "Philyra" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Shen, 1932" ; dwc:species "peitahoensis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Malacostraca" ; dwc:family "Leucosiidae" ; dwc:genus "Lyphira" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "heterograna" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Malacostraca" ; dwc:family "Leucosiidae" ; dwc:genus "Lyphira" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Malacostraca" ; dwc:family "Leucosiidae" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Malacostraca" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Malacostraca" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Malacostraca" ; dwc:family "Leucosiidae" ; dwc:genus "Philyra" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "heterograna" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Malacostraca" ; dwc:family "Leucosiidae" ; dwc:genus "Philyra" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Malacostraca" ; dwc:family "Leucosiidae" ; dwc:genus "Philyra" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "pisum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Malacostraca" ; dwc:family "Leucosiidae" ; dwc:genus "Philyra" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "peitahoensis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Malacostraca" ; dwc:family "Leucosiidae" ; dwc:genus "Philyra" ; dwc:kingdom "Animalia" ; dwc:order "Decapoda" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "anatum" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 2. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191) (specimen number Ortmann 658), Japan.A, overall dorsal habitus; B, overall dorsal habitus (carapace slightly tilted posteriorly); C, overall ventral habitus; D, anterior thoracic sternum, buccal cavity and third maxillipeds." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 12. Third maxillipeds. A, Lyphira chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; B, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; C, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore; D, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.97), Malaysia; E, L. linda n. sp., female (17.0 × 16.9 mm) (ZRC 1995.0547), Singapore; F, L. linda n. sp., male (12.1 × 12.4 mm) (ZRC 1995.0547), Singapore." ; fabio:hasRepresentation ; a fabio:Figure . dwc:collectionCode "MZS" ; dwc:locality "Tokyo Bay" ; dwc:recordedBy "L. Doderlein" ; dwc:typeStatus "lectotype" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "KPM" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "KPM, T" ; dwc:eventDate "1958-11-20" ; dwc:locality "Mikawa-Isshiki" ; dwc:recordedBy "Odawara" ; trt:httpUri ; a dwc:MaterialCitation . trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "KPM, T" ; dwc:locality "Isshiki" ; dwc:recordedBy "Sakai" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NHM" ; dwc:locality "Peichihli Bay" ; dwc:recordedBy "S. S. King" ; dwc:typeStatus "paratype" ; trt:httpUri ; a dwc:MaterialCitation . trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "NHM" ; dwc:locality "Chusan" ; dwc:recordedBy "P. Basset-Smith" ; dwc:verbatimElevation "22" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Bay" ; dwc:typeStatus "holotype" ; trt:httpUri ; a dwc:MaterialCitation .