@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .
@prefix xlink: <http://www.w3.org/1999/xlink/> .

<http://treatment.plazi.org/id/A209493BFD1DFFC2AFA61F71FCE7B870>
    cito:cites <http://dx.doi.org/10.5281/zenodo.7345011>, <http://dx.doi.org/10.5281/zenodo.7345041>, <http://dx.doi.org/10.5281/zenodo.7345051>, <http://dx.doi.org/10.5281/zenodo.7345059>, <http://dx.doi.org/10.5281/zenodo.7345065>, <http://dx.doi.org/10.5281/zenodo.7345083>, <http://dx.doi.org/10.5281/zenodo.7345087>, <http://dx.doi.org/10.5281/zenodo.7345107>, <http://dx.doi.org/10.5281/zenodo.7345180> ;
    dc:creator "López-García, Margarita M.; Deloya, Cuauhtémoc" ;
    dc:title "Tomarus pullus" ;
    dwc:basisOfRecord <http://tb.plazi.org/GgServer/dwcaRecords/A209493BFD1DFFC2AFA61F71FCE7B870.mc.9AC8F270FD1CFFC2AD3A1A31FB95BF8C> ;
    trt:augmentsTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Tomarus_pullus_Prell_1937> ;
    trt:deprecates <http://taxon-concept.plazi.org/id/Animalia/Ligyrus_pullus_Prell_1937> ;
    trt:publishedIn <http://dx.doi.org/10.11646/zootaxa.5211.1.1> ;
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<http://dx.doi.org/10.11646/zootaxa.5211.1.1>
    bibo:endPage "119" ;
    bibo:issue "1" ;
    bibo:journal "Zootaxa" ;
    bibo:pubDate "2022-11-22" ;
    bibo:startPage "1" ;
    bibo:volume "5211" ;
    dc:creator "López-García, Margarita M.; Deloya, Cuauhtémoc" ;
    dc:date "2022" ;
    dc:title "Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" ;
    fabio:hasPart <http://dx.doi.org/10.5281/zenodo.7345011>, <http://dx.doi.org/10.5281/zenodo.7345041>, <http://dx.doi.org/10.5281/zenodo.7345051>, <http://dx.doi.org/10.5281/zenodo.7345059>, <http://dx.doi.org/10.5281/zenodo.7345065>, <http://dx.doi.org/10.5281/zenodo.7345083>, <http://dx.doi.org/10.5281/zenodo.7345087>, <http://dx.doi.org/10.5281/zenodo.7345107>, <http://dx.doi.org/10.5281/zenodo.7345180> ;
    a fabio:JournalArticle .

<http://taxon-concept.plazi.org/id/Animalia/Tomarus_pullus_Prell_1937>
    dwc:authority "(Prell, 1937)" ;
    dwc:baseAuthorityName "Prell" ;
    dwc:baseAuthorityYear "1937" ;
    dwc:class "Insecta" ;
    dwc:family "Dynastidae" ;
    dwc:genus "Tomarus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Prell, 1937)" ;
    dwc:species "pullus" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Tomarus_pullus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Ligyrus_pullus_Prell_1937>
    dwc:authority "Prell 1937: 89" ;
    dwc:authorityName "Prell" ;
    dwc:authorityPageNumber "89" ;
    dwc:authorityYear "1937" ;
    dwc:class "Insecta" ;
    dwc:family "Dynastidae" ;
    dwc:genus "Ligyrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Prell, 1937" ;
    dwc:species "pullus" ;
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<http://taxon-name.plazi.org/id/Animalia/Tomarus_pullus>
    dwc:class "Insecta" ;
    dwc:family "Dynastidae" ;
    dwc:genus "Tomarus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:species "pullus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Tomarus> ;
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<http://taxon-name.plazi.org/id/Animalia/Tomarus>
    dwc:class "Insecta" ;
    dwc:family "Dynastidae" ;
    dwc:genus "Tomarus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Dynastidae> ;
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<http://taxon-name.plazi.org/id/Animalia/Dynastidae>
    dwc:class "Insecta" ;
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    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
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    dwc:rank "family" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Coleoptera> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Coleoptera>
    dwc:class "Insecta" ;
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<http://taxon-name.plazi.org/id/Animalia/Insecta>
    dwc:class "Insecta" ;
    dwc:kingdom "Animalia" ;
    dwc:phylum "Arthropoda" ;
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<http://taxon-name.plazi.org/id/Animalia/Arthropoda>
    dwc:kingdom "Animalia" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "phylum" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia>
    dwc:kingdom "Animalia" ;
    dwc:rank "kingdom" ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Ligyrus_pullus>
    dwc:class "Insecta" ;
    dwc:family "Dynastidae" ;
    dwc:genus "Ligyrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:species "pullus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Ligyrus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Ligyrus>
    dwc:class "Insecta" ;
    dwc:family "Dynastidae" ;
    dwc:genus "Ligyrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Dynastidae> ;
    a dwcFP:TaxonName .

<http://dx.doi.org/10.5281/zenodo.7345011>
    dc:description "FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." ;
    fabio:hasRepresentation <https://zenodo.org/record/7345011/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7345041>
    dc:description "FIGURE 9. Pygidium of male. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) cuniculus, D) L. (Ligyrus) fossor, E) L. (Ligyrus) paranaensis, F) Tomarus bituberculatus, G) T. colombianus, H) T. maracaiboensis, I) T. pilcopataensis, J) T. pullus, K) T. roigjunenti, L) T. rostratus." ;
    fabio:hasRepresentation <https://zenodo.org/record/7345041/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7345051>
    dc:description "FIGURE 11. Protibia in frontal view.A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) relictus, E) L. (Ligyrodes) sallaei, F) L. (Ligyrus) allonasutus, G) L. (Ligyrus) villosus, H) Tomarus bituberculatus, I) T. colombianus, J) T. pullus." ;
    fabio:hasRepresentation <https://zenodo.org/record/7345051/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7345059>
    dc:description "FIGURE 14. Spiculum gastrale. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) L. (Ligyrodes) relictus, D) L. (Ligyrus) villosus, E) Tomarus pullus, F) T. rostratus, G) T. selanderi, H) T. subtropicus." ;
    fabio:hasRepresentation <https://zenodo.org/record/7345059/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7345065>
    dc:description "FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus." ;
    fabio:hasRepresentation <https://zenodo.org/record/7345065/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7345083>
    dc:description "FIGURE 20. Parameres in lateral view. A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus." ;
    fabio:hasRepresentation <https://zenodo.org/record/7345083/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7345087>
    dc:description "FIGURE 22. Internal sac. A) Tomarus pullus, B) T. rostratus, C) T. maternus. Abbreviations: AL: Accessory lamellae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula, SD: Sclerotized diverticulum." ;
    fabio:hasRepresentation <https://zenodo.org/record/7345087/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7345107>
    dc:description "FIGURE 28. Habitus of the species of Tomarus: A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus. Scale bar: 5 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/7345107/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7345180>
    dc:description "FIGURE 48. Distributional records of Tomarus pullus." ;
    fabio:hasRepresentation <https://zenodo.org/record/7345180/files/figure.png> ;
    a fabio:Figure .

<http://tb.plazi.org/GgServer/dwcaRecords/A209493BFD1DFFC2AFA61F71FCE7B870.mc.9AC8F270FD1CFFC2AD3A1A31FB95BF8C>
    dwc:collectionCode "MPUJ, UPN, USNM, ZMHB" ;
    dwc:locality "Rio Orinoco" ;
    dwc:recordedBy "La Urbana" ;
    trt:gbifOccurrenceId "3986696826" ;
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