@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . cito:cites , , , , , , , , , , , , , , , ; dc:creator "López-García, Margarita M.; Deloya, Cuauhtémoc" ; dc:title "Ligyrus (Ligyrodes) relictus" ; dwc:basisOfRecord ; trt:augmentsTaxonConcept ; trt:deprecates , , , , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "119" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:pubDate "2022-11-22" ; bibo:startPage "1" ; bibo:volume "5211" ; dc:creator "López-García, Margarita M.; Deloya, Cuauhtémoc" ; dc:date "2022" ; dc:title "Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" ; fabio:hasPart , , , , , , , , , , , ; a fabio:JournalArticle . dwc:authority "(Say, 1825)" ; dwc:baseAuthorityName "Say" ; dwc:baseAuthorityYear "1825" ; dwc:class "Insecta" ; dwc:family "Dynastidae", "Scarabaeidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Say, 1825)" ; dwc:species "relictus" ; dwc:subGenus "Ligyrodes" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Say, 1825: 194" ; dwc:authorityName "Say" ; dwc:authorityPageNumber "194" ; dwc:authorityYear "1825" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Scarabaeus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Say, 1825" ; dwc:species "relictus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Casey, 1915: 181" ; dwc:authorityName "Casey" ; dwc:authorityPageNumber "181" ; dwc:authorityYear "1915" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrodes" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Casey, 1915" ; dwc:species "clypealis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Casey, 1915: 181" ; dwc:authorityName "Casey" ; dwc:authorityPageNumber "181" ; dwc:authorityYear "1915" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrodes" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Casey, 1915" ; dwc:species "parviceps" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Casey, 1915: 182" ; dwc:authorityName "Casey" ; dwc:authorityPageNumber "182" ; dwc:authorityYear "1915" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrodes" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Casey, 1915" ; dwc:species "quadripennis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Casey, 1915: 183" ; dwc:authorityName "Casey" ; dwc:authorityPageNumber "183" ; dwc:authorityYear "1915" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrodes" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Casey, 1915" ; dwc:species "relictus" ; dwc:subSpecies "vernicicollis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Casey, 1924: 334" ; dwc:authorityName "Casey" ; dwc:authorityPageNumber "334" ; dwc:authorityYear "1924" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrodes" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Casey, 1924" ; dwc:species "dawsoni" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Bates" ; dwc:authorityYear "1888" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Bates, 1888" ; dwc:species "sallaei" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Linnaeus" ; dwc:authorityYear "1758" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Scarabaeus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Linnaeus, 1758" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "sensu Moron & Grossi" ; dwc:baseAuthorityYear "2015" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "(sensu Moron & Grossi, 2015)" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Dejean" ; dwc:authorityYear "1833" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Heteronychus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Dejean, 1833" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Dynastidae", "Scarabaeidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "relictus" ; dwc:subGenus "Ligyrodes" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:rank "subGenus" ; dwc:subGenus "Ligyrodes" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Scarabaeidae" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:order "Coleoptera" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Scarabaeus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "relictus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Scarabaeus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrodes" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "clypealis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:genus "Ligyrodes" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Dynastidae" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrodes" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "parviceps" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrodes" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "quadripennis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrodes" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:species "relictus" ; dwc:subSpecies "vernicicollis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:rank "species" ; dwc:species "relictus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Ligyrodes" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "dawsoni" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURE 2. Labium. A) Ligyrus (Anagrylius) moroni, B) Ligyrus (Ligyrus) allonasutus, C) L. (Ligyrus) gibbosus, D) L. (Ligyrus) villosus, E) L. (Ligyrodes) relictus, F) Proculigyrus cicatricosus, G) Euligyrus ebenus, H) E. similis, I) Tomarus gyas, J) Tomarus selanderi, K) Tomarus subtropicus. Abbreviations: LL: Ligular lobes, LP: Labial palpus, M: Mentum, S: Suture." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 4. Maxilla. A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) relictus, D) L. (Ligyrus) burmeisteri, E) L. (Ligyrus) villosus, F) Tomarus gyas, G) Tomarus maternus, H) T. selanderi, I) T. subtropicus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 5. Mandible. A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) relictus, D) L. (Ligyrus) burmeisteri, E) L. (Ligyrus) villosus, F) Tomarus rostratus, G) T. gyas, H) T. maternus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 6. Labrum. A) Ligyrus (Ligyrus) burmeisteri, B) L. (Ligyrus) villosus, C) Ligyrus (Ligyrodes) relictus, D) Tomarus gyas, E) T. maternus, F) T. selanderi, G) T. subtropicus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 8. Stridulatory apparatus. A, D) Proculigyrus cicatricosus, B, E) Ligyrus (Ligyrus) burmeisteri, C, F) L. (Ligyrodes) relictus, G, K) Tomarus amazonicus, H, L) T. maimon, I, M) T. maternus, J, N) T. gyas." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 11. Protibia in frontal view.A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) relictus, E) L. (Ligyrodes) sallaei, F) L. (Ligyrus) allonasutus, G) L. (Ligyrus) villosus, H) Tomarus bituberculatus, I) T. colombianus, J) T. pullus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 14. Spiculum gastrale. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) L. (Ligyrodes) relictus, D) L. (Ligyrus) villosus, E) Tomarus pullus, F) T. rostratus, G) T. selanderi, H) T. subtropicus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 17. Parameres in lateral view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus; E) L. (Anagrylius) moroni, F) L. (Ligyrodes) relictus, G) L. (Ligyrodes) sallaei." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 25. Habitus of the species of Ligyrus. Subgenus Anagrylius: A) Ligyrus cuniculus, B) L. moroni. Subgenus Ligyrodes: C) L. peruvianus, D) L. relictus, E) L. sallaei. Subgenus Ligyrus: F) L. allonasutus, G) L. bidentulus, H) L. burmeisteri. Scale bar: 5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 33. Distributional records of the subgenus Ligyrus (Ligyrodes)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 10. Protarsus and inner claw. A–B) Euligyrus similis, C–E) Ligyrus (Ligyrodes) sallaei." ; fabio:hasRepresentation ; a fabio:Figure . dwc:collectionCode "CEIOC, FSCA, IEXA, MZUSP, UNSM, USNM" ; dwc:locality "Chalk River" ; dwc:recordedBy "La Porte & Van Buren" ; trt:gbifOccurrenceId "3986696396" ; trt:httpUri ; a dwc:MaterialCitation .