@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .
@prefix xlink: <http://www.w3.org/1999/xlink/> .

<http://treatment.plazi.org/id/F4AB9D33A3825C038F5C4566985CAF35>
    cito:cites <http://dx.doi.org/10.3897/vz.71.e60921.figure3>, <http://dx.doi.org/10.3897/vz.71.e60921.figure4>, <http://dx.doi.org/10.3897/vz.71.e60921.figure7>, <http://dx.doi.org/10.3897/vz.71.e60921.figure8>, <http://dx.doi.org/10.3897/vz.71.e60921.figure9>, <http://dx.doi.org/10.3897/vz.71.e60921.figure10> ;
    dc:creator "Quah, Evan S. H.; Grismer, L. Lee; Wood Jr., Perry L.; Lim, Kelvin K. P.; Imbun, Paul Y.; Anuar, M. S. Shahrul" ;
    dc:title "Abavorana luctuosa" ;
    trt:augmentsTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Abavorana_luctuosa_Peters_1871> ;
    trt:deprecates <http://taxon-concept.plazi.org/id/Animalia/Limnodytes_luctuosus_Peters_1871>, <http://taxon-concept.plazi.org/id/Animalia/Rana_luctuosa_Peters_1871>, <http://taxon-concept.plazi.org/id/Animalia/Hylorana_luctuosa_Peters_1871>, <http://taxon-concept.plazi.org/id/Animalia/Pulchrana_luctuosa_Peters_1871>, <http://taxon-concept.plazi.org/id/Animalia/Hylarana_luctuosa_Peters_1871> ;
    trt:publishedIn <http://dx.doi.org/10.3897/vz.71.e60921> ;
    a trt:Treatment .

<http://dx.doi.org/10.3897/vz.71.e60921>
    bibo:endPage "99" ;
    bibo:journal "Vertebrate Zoology" ;
    bibo:pubDate "2021-03-11" ;
    bibo:startPage "75" ;
    bibo:volume "71" ;
    dc:creator "Quah, Evan S. H.; Grismer, L. Lee; Wood Jr., Perry L.; Lim, Kelvin K. P.; Imbun, Paul Y.; Anuar, M. S. Shahrul" ;
    dc:date "2021" ;
    dc:title "An investigation into the taxonomy of Abavorana luctuosa (Peters, 1871) (Anura, Ranidae) and the resurrection of Rana decorata Mocquard, 1890 from Borneo" ;
    fabio:hasPart <http://dx.doi.org/10.3897/vz.71.e60921.figure3>, <http://dx.doi.org/10.3897/vz.71.e60921.figure4>, <http://dx.doi.org/10.3897/vz.71.e60921.figure7>, <http://dx.doi.org/10.3897/vz.71.e60921.figure8>, <http://dx.doi.org/10.3897/vz.71.e60921.figure9>, <http://dx.doi.org/10.3897/vz.71.e60921.figure10> ;
    a fabio:JournalArticle .

<http://taxon-concept.plazi.org/id/Animalia/Abavorana_luctuosa_Peters_1871>
    dwc:LSID "F4AB9D33-A382-5C03-8F5C-4566985CAF35" ;
    dwc:authority "(Peters, 1871)" ;
    dwc:baseAuthorityName "Peters" ;
    dwc:baseAuthorityYear "1871" ;
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Abavorana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Peters, 1871)" ;
    dwc:species "luctuosa" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Abavorana_luctuosa> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Limnodytes_luctuosus_Peters_1871>
    dwc:authorityName "Peters" ;
    dwc:authorityYear "1871" ;
    dwc:family "Ranidae" ;
    dwc:genus "Limnodytes" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Peters, 1871" ;
    dwc:species "luctuosus" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Limnodytes_luctuosus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Rana_luctuosa_Peters_1871>
    dwc:baseAuthorityName "Peters" ;
    dwc:baseAuthorityYear "1871" ;
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Rana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Peters, 1871)" ;
    dwc:species "luctuosa" ;
    dwc:subGenus "Limnodytes", "Hylorana", "Hylarana", "Pulchrana" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Rana_luctuosa> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Hylorana_luctuosa_Peters_1871>
    dwc:baseAuthorityName "Peters" ;
    dwc:baseAuthorityYear "1871" ;
    dwc:family "Ranidae" ;
    dwc:genus "Hylorana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Peters, 1871)" ;
    dwc:species "luctuosa" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Hylorana_luctuosa> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pulchrana_luctuosa_Peters_1871>
    dwc:baseAuthorityName "Peters" ;
    dwc:baseAuthorityYear "1871" ;
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Pulchrana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Peters, 1871)" ;
    dwc:species "luctuosa" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pulchrana_luctuosa> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Hylarana_luctuosa_Peters_1871>
    dwc:baseAuthorityName "Peters" ;
    dwc:baseAuthorityYear "1871" ;
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Hylarana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Peters, 1871)" ;
    dwc:species "luctuosa" ;
    dwc:subGenus "Abavorana" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Hylarana_luctuosa> ;
    a dwcFP:TaxonConcept .

<http://taxon-name.plazi.org/id/Animalia/Abavorana_luctuosa>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Abavorana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:species "luctuosa" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Abavorana> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Abavorana>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Abavorana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Ranidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Ranidae>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "family" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Anura> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Anura>
    dwc:class "Amphibia" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "order" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Amphibia>, <http://taxon-name.plazi.org/id/Animalia> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Amphibia>
    dwc:class "Amphibia" ;
    dwc:kingdom "Animalia" ;
    dwc:phylum "Chordata" ;
    dwc:rank "class" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Chordata> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Chordata>
    dwc:kingdom "Animalia" ;
    dwc:phylum "Chordata" ;
    dwc:rank "phylum" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia>
    dwc:kingdom "Animalia" ;
    dwc:rank "kingdom" ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Limnodytes_luctuosus>
    dwc:family "Ranidae" ;
    dwc:genus "Limnodytes" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:rank "species" ;
    dwc:species "luctuosus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Limnodytes> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Limnodytes>
    dwc:family "Ranidae" ;
    dwc:genus "Limnodytes" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Ranidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Rana_luctuosa>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Rana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:species "luctuosa" ;
    dwc:subGenus "Limnodytes", "Hylorana", "Hylarana", "Pulchrana" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Rana>, <http://taxon-name.plazi.org/id/Animalia/Rana_Limnodytes>, <http://taxon-name.plazi.org/id/Animalia/Rana_Hylorana>, <http://taxon-name.plazi.org/id/Animalia/Rana_Hylarana>, <http://taxon-name.plazi.org/id/Animalia/Rana_Pulchrana> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Rana>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Rana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Ranidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Rana_Limnodytes>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Rana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Limnodytes" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Rana> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Rana_Hylorana>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Rana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Hylorana" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Rana> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Rana_Hylarana>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Rana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Hylarana" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Rana> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Hylorana_luctuosa>
    dwc:family "Ranidae" ;
    dwc:genus "Hylorana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:rank "species" ;
    dwc:species "luctuosa" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Hylorana> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Hylorana>
    dwc:family "Ranidae" ;
    dwc:genus "Hylorana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Ranidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Rana_Pulchrana>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Rana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Pulchrana" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Rana> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Pulchrana_luctuosa>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Pulchrana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:species "luctuosa" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Pulchrana> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Pulchrana>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Pulchrana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Ranidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Hylarana_luctuosa>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Hylarana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:species "luctuosa" ;
    dwc:subGenus "Abavorana" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Hylarana>, <http://taxon-name.plazi.org/id/Animalia/Hylarana_Abavorana> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Hylarana>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Hylarana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Ranidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Hylarana_Abavorana>
    dwc:class "Amphibia" ;
    dwc:family "Ranidae" ;
    dwc:genus "Hylarana" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Anura" ;
    dwc:phylum "Chordata" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Abavorana" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Hylarana> ;
    a dwcFP:TaxonName .

<http://dx.doi.org/10.3897/vz.71.e60921.figure3>
    dc:description "Figure 3. Holotype of Abavorana luctuosa; A: Original illustration of the holotype of Abavorana luctuosa from Peters (1872). B: Dorsum of the holotype of A. luctuosa (MSNG 29344). C: Venter of the holotype of A. luctuosa (MSNG 29344). Photographs by Giuliano Doria." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/518976> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/vz.71.e60921.figure4>
    dc:description "Figure 4. Bornean Abavorana luctuosa; Dorsal (A & C) and ventral (B & D) pattern of Abavorana luctuosa from Borneo. A & B: Top row (from left to right); FMNH 234972, FMNH 234973, FMNH 234975 and FMNH 234977 (all specimens from Sipitang District, Sabah). Bottom row (from left to right); FMNH 234980 (Sipitang District, Sabah), FMNH 269142, FMNH 273216 and FMNH 273217 (specimens from Bintulu Division, Sarawak). C & D (from left to right): ZRC 1.11938, ZRC 1.11943 (both from Kubah, Sarawak) and ZRC 1.2270 (unspecified location in Borneo)." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/518977> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/vz.71.e60921.figure7>
    dc:description "Figure 7. Living specimens of Abavorana decorata and Bornean A. luctuosa; A: Abavorana decorata from Tawau Hills, Sabah, East Malaysia. B: A. decorata from Maliau Basin, Sabah, East Malaysia. C: A. decorata from Kubah National Park, Sarawak, East Malaysia. D: A. decorata from Melawi Regency, West Kalimantan Province, Indonesia. E & F: A. luctuosa from Kubah National Park, Sarawak, East Malaysia. G: A. luctuosa (FMNH 238720) from Sipitang, Sabah, East Malaysia. H: A. luctuosa (FMNH 242883) from Sipitang, Sabah, East Malaysia. (Photographs by Alexander Haas [A, C & E], Paul Bertner [B], Muhammad Aini [D], Chien C. Lee [F], and Robert F. Inger [G & H], Figs G & H © Field Museum of Natural History. Created by Field Museum of Natural History, Amphibian and Reptile Collection and licensed under CC-BY-SA 4.0)." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/518980> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/vz.71.e60921.figure8>
    dc:description "Figure 8. Peninsular Malaysian, Thai and Sumatran Abavorana luctuosa, and A. nazgul; A: A. luctuosa from Sungai Durian Forest Reserve, Kelantan, Peninsular Malaysia. B: A. luctuosa from Janda Baik, Pahang, Peninsular Malaysia. C: A. luctuosa from Semenyih, Selangor, Peninsular Malaysia. D: A. luctuosa from Fraser's Hill, Pahang, Peninsular Malaysia. E: A. luctuosa from Hala-Bala Wildlife Sanctuary, Thailand. F: A. luctuosa from Sumatra, Indonesia. G: A. nazgul (USMHC 2572) from Gunung Jerai, Kedah, Peninsular Malaysia. H. A. cf. nazgul from Thale Ban National Park, Satun Province, Thailand. (Photographs by Zaharil Dzulkafly [A], Steven Wong [B], Kurt H. P. Guek [C], Evan Quah [D & G], Ian Dugdale [E], Hellen Kurniati [F] & Bunyarit Dechochai [H])." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/518981> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/vz.71.e60921.figure9>
    dc:description "Figure 9. Colour pattern comparison among species of the genus Abavorana; Dorsum (A, C & E) and venter (B, D & F) in life of A. luctuosa (USMHC 2576) (A & B) from Penang Hill, Penang Island, Malaysia, A. decorata from Maliau Basin, Sabah, East Malaysia, Borneo (C & D) and A. nazgul (USMHC 2233) (E & F) from Gunung Jerai, Kedah, West Malaysia. (Photographs by Evan Quah [A, B, E & F] and Stefan T. Hertwig [C & D])." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/518982> ;
    a fabio:Figure .

<http://dx.doi.org/10.3897/vz.71.e60921.figure10>
    dc:description "Figure 10. Distribution map of Abavorana decorata and A. luctuosa in Borneo; Known distribution of Abavorana decorata (red) and A. luctuosa (blue) in Borneo. Star = type locality." ;
    fabio:hasRepresentation <https://binary.pensoft.net/fig/518983> ;
    a fabio:Figure .