@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .

<http://treatment.plazi.org/id/03AED64E3E550A66FF37305F24C5FBC5>
    cito:cites <http://taxon-concept.plazi.org/id/Animalia/Hoplia_Illiger_1803>, <http://taxon-concept.plazi.org/id/Animalia/Hopliini_Latreille_1829>, <http://taxon-concept.plazi.org/id/Animalia/Melolontha_crassipes_Fabricius_1775>, <http://taxon-concept.plazi.org/id/Animalia/Melolontha_tibialis_Olivier_1789>, <http://taxon-concept.plazi.org/id/Animalia/Pachycnema_Cupello_2019>, <http://taxon-concept.plazi.org/id/Animalia/Pachycnema_Saint_1828>, <http://taxon-concept.plazi.org/id/Animalia/Pachycnema_abdominalis_Burmeister_1844>, <http://taxon-concept.plazi.org/id/Animalia/Pachycnema_kochi_Schein_1959>, <http://taxon-concept.plazi.org/id/Animalia/Pachycnema_moerens_Peringuey_1902>, <http://taxon-concept.plazi.org/id/Animalia/Pachycnema_signatipennis_Burmeister_1844>, <http://taxon-concept.plazi.org/id/Animalia/Pachycnemida_multiguttata_Thunberg_1818>, <http://taxon-concept.plazi.org/id/Animalia/Pachypodini_Erichson_1840>, <http://taxon-concept.plazi.org/id/Animalia/Pachypus_Dejean_1821> ;
    dc:creator "Cupello, Mario; Ribeiro-Costa, Cibele Stramare" ;
    dc:title "Pachycnema (Macacoplia) Cupello & Ribeiro-Costa 2019, new subgenus" ;
    trt:citesTaxonName <http://taxon-name.plazi.org/id/Animalia/Melolonthinae> ;
    trt:definesTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Pachycnema_Macacoplia_Cupello_2019> ;
    trt:deprecates <http://taxon-concept.plazi.org/id/Animalia/Pachycnema_Pachycnemula_Schein_1959>, <http://taxon-concept.plazi.org/id/Animalia/Pachycnemida_Peringuey_1902>, <http://taxon-concept.plazi.org/id/Animalia/Pachypus_Billberg_1820>, <http://taxon-concept.plazi.org/id/Animalia/Physocnema_Burmeister_1844>, <http://taxon-concept.plazi.org/id/Animalia/Stenocnema_Burmeister_1844> ;
    trt:publishedIn <http://dx.doi.org/10.11646/zootaxa.4543.1.9> ;
    a trt:Treatment .

<http://dx.doi.org/10.11646/zootaxa.4543.1.9>
    bibo:endPage "144" ;
    bibo:issue "1" ;
    bibo:journal "Zootaxa" ;
    bibo:pubDate "2019-01-04" ;
    bibo:startPage "137" ;
    bibo:volume "4543" ;
    dc:creator "Cupello, Mario; Ribeiro-Costa, Cibele Stramare" ;
    dc:date "2019" ;
    dc:title "Nomenclatural clarification on some Southern African monkey beetle genera (Scarabaeidae: Hopliini: Pachycnemina)" ;
    a fabio:JournalArticle .

<http://taxon-concept.plazi.org/id/Animalia/Pachycnema_Macacoplia_Cupello_2019>
    dwc:authority "Cupello & Ribeiro-Costa", "Cupello & Ribeiro-Costa, 2019" ;
    dwc:authorityName "Cupello & Ribeiro-Costa" ;
    dwc:authorityYear "2019" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachycnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "subGenus" ;
    dwc:scientificNameAuthorship "Cupello & Ribeiro-Costa, 2019" ;
    dwc:subGenus "Macacoplia" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/8F2T> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachycnema_Macacoplia> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Hopliini_Latreille_1829>
    dwc:authorityName "Latreille" ;
    dwc:authorityYear "1829" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "tribe" ;
    dwc:scientificNameAuthorship "Latreille, 1829" ;
    dwc:tribe "Hopliini" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Hopliini> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Hoplia_Illiger_1803>
    dwc:authority "Illiger, 1803" ;
    dwc:authorityName "Illiger" ;
    dwc:authorityYear "1803" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Hoplia" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    dwc:scientificNameAuthorship "Illiger, 1803" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Hoplia> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pachycnema_moerens_Peringuey_1902>
    dwc:authority "Peringuey, 1902" ;
    dwc:authorityName "Peringuey" ;
    dwc:authorityYear "1902" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachycnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Peringuey, 1902" ;
    dwc:species "moerens" ;
    dwc:subGenus "Macacoplia" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/4BQR7> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachycnema_moerens> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pachycnema_Cupello_2019>
    dwc:authority "Cupello & Ribeiro-Costa, 2019" ;
    dwc:authorityName "Cupello & Ribeiro-Costa" ;
    dwc:authorityYear "2019" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachycnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    dwc:scientificNameAuthorship "Cupello & Ribeiro-Costa, 2019" ;
    dwc:sensu "stricto" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/6BR8> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachycnema> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pachycnema_abdominalis_Burmeister_1844>
    dwc:authority "(Burmeister, 1844)" ;
    dwc:authorityName "Burmeister" ;
    dwc:authorityYear "1844" ;
    dwc:baseAuthorityName "Burmeister" ;
    dwc:baseAuthorityYear "1844" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachycnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Burmeister, 1844)" ;
    dwc:species "abdominalis" ;
    dwc:subGenus "Macacoplia" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachycnema_abdominalis> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pachycnema_kochi_Schein_1959>
    dwc:authority "Schein, 1959" ;
    dwc:authorityName "Schein" ;
    dwc:authorityYear "1959" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachycnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Schein, 1959" ;
    dwc:species "kochi" ;
    dwc:subGenus "Macacoplia" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachycnema_kochi> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Melolontha_tibialis_Olivier_1789>
    dwc:authority "(Olivier, 1789)", "Olivier, 1789" ;
    dwc:authorityName "Olivier" ;
    dwc:authorityYear "1789" ;
    dwc:baseAuthorityName "Olivier" ;
    dwc:baseAuthorityYear "1789" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Melolontha" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Olivier, 1789)" ;
    dwc:species "tibialis" ;
    dwc:subGenus "Macacoplia" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Melolontha_tibialis> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pachycnema_Pachycnemula_Schein_1959>
    dwc:authority "Schein, 1959" ;
    dwc:authorityName "Schein" ;
    dwc:authorityYear "1959" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachycnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "subGenus" ;
    dwc:scientificNameAuthorship "Schein, 1959" ;
    dwc:subGenus "Pachycnemula" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/7Q88X> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachycnema_Pachycnemula> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pachycnemida_Peringuey_1902>
    dwc:authority ", Peringuey (1902: 689)", "Peringuey, 1902" ;
    dwc:authorityName ", Peringuey" ;
    dwc:authorityPageNumber "689" ;
    dwc:authorityYear "1902" ;
    dwc:class "Insecta" ;
    dwc:genus "Pachycnemida" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    dwc:scientificNameAuthorship "Peringuey, 1902" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/63JJP> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachycnemida> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pachycnema_signatipennis_Burmeister_1844>
    dwc:authority "(Burmeister, 1844)" ;
    dwc:authorityName "Burmeister" ;
    dwc:authorityYear "1844" ;
    dwc:baseAuthorityName "Burmeister" ;
    dwc:baseAuthorityYear "1844" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachycnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Burmeister, 1844)" ;
    dwc:species "signatipennis" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachycnema_signatipennis> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pachycnemida_multiguttata_Thunberg_1818>
    dwc:authority "(Thunberg, 1818)" ;
    dwc:authorityName "Thunberg" ;
    dwc:authorityYear "1818" ;
    dwc:baseAuthorityName "Thunberg" ;
    dwc:baseAuthorityYear "1818" ;
    dwc:class "Insecta" ;
    dwc:genus "Pachycnemida" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Thunberg, 1818)" ;
    dwc:species "multiguttata" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachycnemida_multiguttata> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Physocnema_Burmeister_1844>
    dwc:authority "Burmeister, 1844" ;
    dwc:authorityName "Burmeister" ;
    dwc:authorityYear "1844" ;
    dwc:class "Insecta" ;
    dwc:genus "Physocnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    dwc:scientificNameAuthorship "Burmeister, 1844" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Physocnema> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pachycnema_Saint_1828>
    dwc:authority "Le Peletier de Saint-Fargeau & Audinet-Serville, 1828" ;
    dwc:authorityName "Le Peletier de Saint-Fargeau & Audinet-Serville" ;
    dwc:authorityYear "1828" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachycnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    dwc:scientificNameAuthorship "Le Peletier de Saint-Fargeau & Audinet-Serville, 1828" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/6BR8> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachycnema> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pachypus_Billberg_1820>
    dwc:authority "Billberg", "Billberg, 1820" ;
    dwc:authorityName "Billberg" ;
    dwc:authorityYear "1820" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachypus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    dwc:scientificNameAuthorship "Billberg, 1820" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/63JKJ> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachypus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Melolontha_crassipes_Fabricius_1775>
    dwc:authority "Fabricius, 1775" ;
    dwc:authorityName "Fabricius" ;
    dwc:authorityYear "1775" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Melolontha" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Fabricius, 1775" ;
    dwc:species "crassipes" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Melolontha_crassipes> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pachypus_Dejean_1821>
    dwc:authority "Dejean, 1821" ;
    dwc:authorityName "Dejean" ;
    dwc:authorityYear "1821" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachypus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    dwc:scientificNameAuthorship "Dejean, 1821" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachypus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pachypodini_Erichson_1840>
    dwc:authorityName "Erichson" ;
    dwc:authorityYear "1840" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "tribe" ;
    dwc:scientificNameAuthorship "Erichson, 1840" ;
    dwc:tribe "Pachypodini" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pachypodini> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Stenocnema_Burmeister_1844>
    dwc:authority "Burmeister, 1844" ;
    dwc:authorityName "Burmeister" ;
    dwc:authorityYear "1844" ;
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Stenocnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    dwc:scientificNameAuthorship "Burmeister, 1844" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Stenocnema> ;
    a dwcFP:TaxonConcept .

<http://taxon-name.plazi.org/id/Animalia/Pachycnema_Macacoplia>
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachycnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Macacoplia" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Pachycnema> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Pachycnema>
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachycnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Scarabaeidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Scarabaeidae>
    dwc:family "Scarabaeidae" ;
    dwc:rank "family" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Coleoptera> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Coleoptera>
    dwc:order "Coleoptera" ;
    dwc:rank "order" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Insecta> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Insecta>
    dwc:class "Insecta" ;
    dwc:rank "class" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Arthropoda> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Arthropoda>
    dwc:phylum "Arthropoda" ;
    dwc:rank "phylum" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia>
    dwc:kingdom "Animalia" ;
    dwc:rank "kingdom" ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Pachypus>
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachypus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Scarabaeidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Pachycnema_Pachycnemula>
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Pachycnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Pachycnemula" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Pachycnema> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Stenocnema>
    dwc:class "Insecta" ;
    dwc:family "Scarabaeidae" ;
    dwc:genus "Stenocnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Scarabaeidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Physocnema>
    dwc:class "Insecta" ;
    dwc:genus "Physocnema" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Coleoptera> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Pachycnemida>
    dwc:class "Insecta" ;
    dwc:genus "Pachycnemida" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Coleoptera> ;
    a dwcFP:TaxonName .