Cyrtodactylus muluensis Davis & Bauer & Jackman & Nashriq & Das 2019, sp. nov.
cites: zenodo.3242438
cites: zenodo.3242440
cites: zenodo.3242442
cites: zenodo.3242444
cites: Cyrtodactylus agusanensis Taylor 1915
cites: Cyrtodactylus annulatus Taylor 1915
cites: Cyrtodactylus aurensis Grismer 2005
cites: Cyrtodactylus baluensis Mocquard 1890
cites: Cyrtodactylus batucolus Grismer 2008
cites: Cyrtodactylus condorensis Smith 1921
cites: Cyrtodactylus consobrinus Peters 1871
cites: Cyrtodactylus darmandvillei Weber 1890
cites: Cyrtodactylus grismeri Tri 2008
cites: Cyrtodactylus gunungsenyumensis Grismer 2016
cites: Cyrtodactylus ingeri Hikida 1990
cites: Cyrtodactylus irregularis Smith 1921
cites: Cyrtodactylus jellesmae Boulenger 1897
cites: Cyrtodactylus macrotuberculatus Grismer 2008
cites: Cyrtodactylus malayanus Rooij
cites: Cyrtodactylus matsuii Hikida 1990
cites: Cyrtodactylus pantiensis Grismer 2008
cites: Cyrtodactylus papuensis Brongersma 1934
cites: Cyrtodactylus paradoxus Darevsky 1997
cites: Cyrtodactylus philippinicus Steindachner 1867
cites: Cyrtodactylus psarops Harvey 2015
cites: Cyrtodactylus pubisulcus Inger 1958
cites: Cyrtodactylus rosichonariefi Riyanto 2015
cites: Cyrtodactylus semenanjungensis Grismer 2005
cites: Cyrtodactylus seribuatensis Youmans 2006
cites: Cyrtodactylus sworderi Smith 1925
cites: Cyrtodactylus tautbatorum Welton 2009
cites: Cyrtodactylus tiomanensis Das 2000
cites: Cyrtodactylus yoshii Hikida 1990
creator: Davis, Hayden R.; Bauer, Aaron M.; Jackman, Todd R.; Nashriq, Izneil; Das, Indraneil
cites taxon name: Cyrtodactylus
cites taxon name: Cyrtodactylus cavernicolus
cites taxon name: Cyrtodactylus jambangan
cites taxon name: Cyrtodactylus jarujini
cites taxon name: Cyrtodactylus malayanus
cites taxon name: Cyrtodactylus metropolis
cites taxon name: Cyrtodactylus quadrivirgatus
cites taxon name: Cyrtodactylus redimiculus
cites taxon name: Cyrtodactylus semicinctus
defines taxon concept: Cyrtodactylus muluensis Davis 2019
published in: Uncovering karst endemism within Borneo: two new Cyrtodactylus species from Sarawak, Malaysia
type: Treatment

Uncovering karst endemism within Borneo: two new Cyrtodactylus species from Sarawak, Malaysia
end page: 352
issue: 2
journal: Zootaxa
pub date: 2019-06-10
start page: 331
volume: 4614
creator: Davis, Hayden R.; Bauer, Aaron M.; Jackman, Todd R.; Nashriq, Izneil; Das, Indraneil
date: 2019
has part: zenodo.3242438
has part: zenodo.3242440
has part: zenodo.3242442
has part: zenodo.3242444
type: JournalArticle

Cyrtodactylus muluensis Davis 2019
authority: Davis & Bauer & Jackman & Nashriq & Das, 2019
authority name: Davis & Bauer & Jackman & Nashriq & Das
authority year: 2019
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Davis, Bauer, Jackman, Nashriq & Das, 2019
species: muluensis
has taxon name: Cyrtodactylus muluensis
type: TaxonConcept

Cyrtodactylus agusanensis Taylor 1915
base authority name: Taylor
base authority year: 1915
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (Taylor, 1915)
species: agusanensis
has taxon name: Cyrtodactylus agusanensis
type: TaxonConcept

Cyrtodactylus annulatus Taylor 1915
base authority name: Taylor
base authority year: 1915
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (Taylor, 1915)
species: annulatus
has taxon name: Cyrtodactylus annulatus
type: TaxonConcept

Cyrtodactylus aurensis Grismer 2005
authority name: Grismer
authority year: 2005
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Grismer, 2005
species: aurensis
has taxon name: Cyrtodactylus aurensis
type: TaxonConcept

Cyrtodactylus baluensis Mocquard 1890
authority: (Mocquard)
authority name: Mocquard
base authority name: Mocquard
base authority year: 1890
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (Mocquard, 1890)
species: baluensis
has taxon name: Cyrtodactylus baluensis
type: TaxonConcept

Cyrtodactylus batucolus Grismer 2008
authority name: Grismer, Chan, Grismer, Wood & Belabut
authority year: 2008
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Grismer, Chan, Grismer, Wood & Belabut, 2008
species: batucolus
has taxon name: Cyrtodactylus batucolus
type: TaxonConcept

Cyrtodactylus condorensis Smith 1921
base authority name: Smith
base authority year: 1921
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (Smith, 1921)
species: condorensis
has taxon name: Cyrtodactylus condorensis
type: TaxonConcept

Cyrtodactylus consobrinus Peters 1871
authority: (Peters)
authority name: Peters
base authority name: Peters
base authority year: 1871
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (Peters, 1871)
species: consobrinus
has taxon name: Cyrtodactylus consobrinus
type: TaxonConcept

Cyrtodactylus darmandvillei Weber 1890
base authority name: Weber
base authority year: 1890
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (Weber, 1890)
species: darmandvillei
has taxon name: Cyrtodactylus darmandvillei
type: TaxonConcept

Cyrtodactylus grismeri Tri 2008
authority name: Ngo Van Tri
authority year: 2008
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Ngo Van Tri, 2008
species: grismeri
has taxon name: Cyrtodactylus grismeri
type: TaxonConcept

Cyrtodactylus gunungsenyumensis Grismer 2016
authority name: Grismer, Wood, Anuar, Davis, Cobos & Murdoch
authority year: 2016
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Grismer, Wood, Anuar, Davis, Cobos & Murdoch, 2016
species: gunungsenyumensis
has taxon name: Cyrtodactylus gunungsenyumensis
type: TaxonConcept

Cyrtodactylus irregularis Smith 1921
base authority name: Smith
base authority year: 1921
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (Smith, 1921)
species: irregularis
has taxon name: Cyrtodactylus irregularis
type: TaxonConcept

Cyrtodactylus jellesmae Boulenger 1897
base authority name: Boulenger
base authority year: 1897
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (Boulenger, 1897)
species: jellesmae
has taxon name: Cyrtodactylus jellesmae
type: TaxonConcept

Cyrtodactylus macrotuberculatus Grismer 2008
authority name: Grismer & Ahmad
authority year: 2008
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Grismer & Ahmad, 2008
species: macrotuberculatus
has taxon name: Cyrtodactylus macrotuberculatus
type: TaxonConcept

Cyrtodactylus pantiensis Grismer 2008
authority name: Grismer, Chan, Grismer, Wood & Belabut
authority year: 2008
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Grismer, Chan, Grismer, Wood & Belabut, 2008
species: pantiensis
has taxon name: Cyrtodactylus pantiensis
type: TaxonConcept

Cyrtodactylus papuensis Brongersma 1934
base authority name: Brongersma
base authority year: 1934
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (Brongersma, 1934)
species: papuensis
has taxon name: Cyrtodactylus papuensis
type: TaxonConcept

Cyrtodactylus paradoxus Darevsky 1997
base authority name: Darevsky & Szczerbak
base authority year: 1997
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (Darevsky & Szczerbak, 1997)
species: paradoxus
has taxon name: Cyrtodactylus paradoxus
type: TaxonConcept

Cyrtodactylus philippinicus Steindachner 1867
base authority name: Steindachner
base authority year: 1867
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (Steindachner, 1867)
species: philippinicus
has taxon name: Cyrtodactylus philippinicus
type: TaxonConcept

Cyrtodactylus psarops Harvey 2015
authority name: Harvey, O'connell, Barraza, Riyanto, Kurniawan & Smith
authority year: 2015
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Harvey, Oconnell, Barraza, Riyanto, Kurniawan & Smith, 2015
species: psarops
has taxon name: Cyrtodactylus psarops
type: TaxonConcept

Cyrtodactylus pubisulcus Inger 1958
authority: Inger
authority name: Inger
authority year: 1958
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Inger, 1958
species: pubisulcus
has taxon name: Cyrtodactylus pubisulcus
type: TaxonConcept

Cyrtodactylus rosichonariefi Riyanto 2015
authority name: Riyanto, Grismer & Wood
authority year: 2015
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Riyanto, Grismer & Wood, 2015
species: rosichonariefi
has taxon name: Cyrtodactylus rosichonariefi
type: TaxonConcept

Cyrtodactylus semenanjungensis Grismer 2005
authority name: Grismer & Leong
authority year: 2005
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Grismer & Leong, 2005
species: semenanjungensis
has taxon name: Cyrtodactylus semenanjungensis
type: TaxonConcept

Cyrtodactylus seribuatensis Youmans 2006
authority name: Youmans & Grismer
authority year: 2006
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Youmans & Grismer, 2006
species: seribuatensis
has taxon name: Cyrtodactylus seribuatensis
type: TaxonConcept

Cyrtodactylus sworderi Smith 1925
base authority name: Smith
base authority year: 1925
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (Smith, 1925)
species: sworderi
has taxon name: Cyrtodactylus sworderi
type: TaxonConcept

Cyrtodactylus tautbatorum Welton 2009
authority name: Welton, Siler, Diesmos & Brown
authority year: 2009
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Welton, Siler, Diesmos & Brown, 2009
species: tautbatorum
has taxon name: Cyrtodactylus tautbatorum
type: TaxonConcept

Cyrtodactylus tiomanensis Das 2000
authority name: Das & Lim
authority year: 2000
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Das & Lim, 2000
species: tiomanensis
has taxon name: Cyrtodactylus tiomanensis
type: TaxonConcept

Cyrtodactylus yoshii Hikida 1990
authority: Hikida
authority name: Hikida
authority year: 1990
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Hikida, 1990
species: yoshii
has taxon name: Cyrtodactylus yoshii
type: TaxonConcept

Cyrtodactylus ingeri Hikida 1990
authority: Hikida
authority name: Hikida
authority year: 1990
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Hikida, 1990
species: ingeri
has taxon name: Cyrtodactylus ingeri
type: TaxonConcept

Cyrtodactylus malayanus Rooij
authority: (de Rooij)
authority name: de Rooij
base authority name: de Rooij
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: (de Rooij)
species: malayanus
has taxon name: Cyrtodactylus malayanus
type: TaxonConcept

Cyrtodactylus matsuii Hikida 1990
authority: Hikida
authority name: Hikida
authority year: 1990
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
scientific name authorship: Hikida, 1990
species: matsuii
has taxon name: Cyrtodactylus matsuii
type: TaxonConcept

Cyrtodactylus muluensis
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: species
species: muluensis
has parent name: Cyrtodactylus
type: TaxonName

Cyrtodactylus
class: Reptilia
family: Gekkonidae
genus: Cyrtodactylus
kingdom: Animalia
order: Squamata
phylum: Chordata
rank: genus
has parent name: Gekkonidae
type: TaxonName

Gekkonidae
family: Gekkonidae
rank: family
has parent name: Squamata
type: TaxonName

Squamata
order: Squamata
rank: order
has parent name: Reptilia
type: TaxonName

Reptilia
class: Reptilia
rank: class
has parent name: Chordata
type: TaxonName

Chordata
phylum: Chordata
rank: phylum
has parent name: Animalia
type: TaxonName

Animalia
kingdom: Animalia
rank: kingdom
type: TaxonName

zenodo.3242438
description: FIGURE 1. Preserved type series of Cyrtodactylus muluensis sp. nov. A) type series displaying dorsal side of body (from left to right: CAS 262995; CAS 262986; CAS 262983; CAS 262997; CAS 262996; and CAS 262984; CAS 262985 not shown); and B) type series displaying ventral side of body (specimens arranged in the same order as figure 1A).
type: Figure

zenodo.3242440
description: FIGURE 2. Photographs of live individuals of Cyrtodactylus muluensis sp. nov. A) adult holotype (CAS 262983); B) juvenile paratype (CAS 262995).
type: Figure

zenodo.3242442
description: FIGURE 3. Map of Eastern Malaysia with circles indicating the type locality of the three Bornean karst-dwelling Cyrtodactylus species. Map is an altered version of 2018 Google Earth, US Dept of State Geographer map.
type: Figure

zenodo.3242444
description: FIGURE 4. A) Mulu karst formation demonstrating Cyrtodactylus muluensis sp. nov. microhabitat; B) type locality of Cyrtodactylus limajalur sp. nov. The type locality of C. muluensis sp. nov. is primarily old growth with low-lying plants near the karst walls; the type locality of C. limajalur sp. nov. is on the edge of a local palm oil plantation with some old growth remaining at the base of the karst wall and on the portions of the formation at higher elevation.
type: Figure

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collection code: CAS, WGS
event date: 2017-07-21
locality: Adult
recorded by: Izneil Nashriq & Hayden Davis
type status: holotype
verbatim elevation: 155
verbatim latitude: 4.026206
verbatim longitude: 114.824104
gbif occurrence id: 2251813388
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gbif occurrence id: 2397873677
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locality: Bunker Trail
gbif occurrence id: 2397873653
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gbif occurrence id: 2397873669
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collection code: MZB
locality: Kerinci
gbif occurrence id: 2397873666
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collection code: MZB
locality: Kerinci
gbif occurrence id: 2397873663
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collection code: LSUHC
locality: Pulau Mentigi
gbif occurrence id: 2397873655
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collection code: LSUHC
locality: Pulau Mentigi
gbif occurrence id: 2397873661
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locality: Peta
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gbif occurrence id: 2397873657
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collection code: KU
locality: Municipality of Brooke's Point
gbif occurrence id: 2397873652
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collection code: LSUHC
locality: Pulau Tioman
gbif occurrence id: 2397873671
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catalog number: ZRC2.4851
collection code: ZRC
locality: Sungai Kipungit
gbif occurrence id: 2397873656
http uri: https://treatment.plazi.org/id/03A3CA13FFC5FFCFFF369260FBC2E5FC#3B627158FFC1FFC3FB829345F9B3E17F
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catalog number: RMBR00847
collection code: RMBR
locality: Bukit Baka Bukit Raya
gbif occurrence id: 2397873654
http uri: https://treatment.plazi.org/id/03A3CA13FFC5FFCFFF369260FBC2E5FC#3B627158FFC1FFC3FB82931DF98EE147
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catalog number: CAS 262984, CAS 262986
collection code: CAS
event date: 2017-07-21
locality: Adult
recorded by: Izneil Nashriq & Hayden Davis
type status: paratype
verbatim elevation: 155
verbatim latitude: 4.026206
verbatim longitude: 114.824104
gbif occurrence id: 2251813323
http uri: https://treatment.plazi.org/id/03A3CA13FFC5FFCFFF369260FBC2E5FC#3B627158FFCFFFCDFEF2917FFF53E30A
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catalog number: CAS 262995-262997, WGS 1984
collection code: CAS
locality: Clearwater Cave
type status: paratype
verbatim elevation: 205
verbatim latitude: 4.063069
verbatim longitude: 114.83072
gbif occurrence id: 2251813310
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