@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . cito:cites , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; dc:creator "Rossini, Michele; Vaz-De, Fernando Z.; Zunino, Mario" ; dc:title "Onthophagus hircus Billberg 1815" ; trt:augmentsTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "21" ; bibo:journal "European Journal of Taxonomy" ; bibo:pubDate "2018-05-04" ; bibo:startPage "1" ; bibo:volume "432" ; dc:creator "Rossini, Michele; Vaz-De, Fernando Z.; Zunino, Mario" ; dc:date "2018" ; dc:title "Toward a comprehensive taxonomic revision of the ā€œ hirculus ā€ group of American Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae)" ; fabio:hasPart ; a fabio:JournalArticle . dwc:authority "Billberg, 1815" ; dwc:authorityName "Billberg" ; dwc:authorityYear "1815" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Billberg, 1815" ; dwc:species "hircus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Latreille, 1809" ; dwc:authorityName "Latreille" ; dwc:authorityYear "1809" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Latreille, 1809" ; dwc:species "curvicornis" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Say, 1835" ; dwc:authorityName "Say" ; dwc:authorityYear "1835" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Say, 1835" ; dwc:species "incensus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Bates, 1886" ; dwc:authorityName "Bates" ; dwc:authorityYear "1886" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Bates, 1886" ; dwc:species "nitidior" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Howden & Young, 1981" ; dwc:authorityName "Howden &Young" ; dwc:authorityYear "1981" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Howden &Young, 1981" ; dwc:species "stockwelli" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Howden & Cartwright, 1963" ; dwc:authorityName "Howden & Cartwright" ; dwc:authorityYear "1963" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Howden & Cartwright, 1963" ; dwc:species "batesi" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Harold, 1875" ; dwc:authorityName "Harold" ; dwc:authorityYear "1875" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Harold, 1875" ; dwc:species "janthinus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Harold, 1880" ; dwc:authorityName "Harold" ; dwc:authorityYear "1880" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Harold, 1880" ; dwc:species "acuminatus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Harold, 1873" ; dwc:authorityName "Harold" ; dwc:authorityYear "1873" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Harold, 1873" ; dwc:species "tristis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . # Warning: Removing "(Palisot de Beauvois" from authority ; dwc:authority "striatulus (Palisot de Beauvois, 1809)" ; dwc:authorityName "striatulus (Palisot de Beauvois" ; dwc:authorityYear "1809" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "striatulus, 1809" ; dwc:species "striatulus" ; dwc:subSpecies "striatulus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Blatchley, 1928" ; dwc:authorityName "Blatchley" ; dwc:authorityYear "1928" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Blatchley, 1928" ; dwc:species "striatulus" ; dwc:subSpecies "floridanus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Erichson, 1847" ; dwc:authorityName "Erichson" ; dwc:authorityYear "1847" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Erichson, 1847" ; dwc:species "ptox" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Blanchard, 1846" ; dwc:authorityName "Blanchard" ; dwc:authorityYear "1846" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Blanchard, 1846" ; dwc:species "aeneus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Drapiez, 1819" ; dwc:authorityName "Drapiez" ; dwc:authorityYear "1819" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Drapiez, 1819" ; dwc:species "bidentatus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Harold, 1880" ; dwc:authorityName "Harold" ; dwc:authorityYear "1880" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Harold, 1880" ; dwc:species "marginicollis" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Arrow, 1903" ; dwc:authorityName "Arrow" ; dwc:authorityYear "1903" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Arrow, 1903" ; dwc:species "antillarum" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Mannerheim, 1829" ; dwc:authorityName "Mannerheim" ; dwc:authorityYear "1829" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Mannerheim, 1829" ; dwc:species "buculus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Arrow, 1913" ; dwc:authorityName "Arrow" ; dwc:authorityYear "1913" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Arrow, 1913" ; dwc:species "ranunculus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ", Erichson, 1847" ; dwc:authorityName ", Erichson" ; dwc:authorityYear "1847" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Erichson, 1847" ; dwc:species "ophion" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Guerin-Meneville, 1855" ; dwc:authorityName "Guerin-Meneville" ; dwc:authorityYear "1855" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Guerin-Meneville, 1855" ; dwc:species "osculatii" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Bates, 1886" ; dwc:authorityName "Bates" ; dwc:authorityYear "1886" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Bates, 1886" ; dwc:species "nyctopus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Harold, 1880" ; dwc:authorityName "Harold" ; dwc:authorityYear "1880" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Harold, 1880" ; dwc:species "steinheili" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Howden & Young, 1981" ; dwc:authorityName "Howden & Young" ; dwc:authorityYear "1981" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Howden & Young, 1981" ; dwc:species "transisthmius" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "sensu Boucomont, 1932" ; dwc:authorityName "Boucomont" ; dwc:authorityYear "1932" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Boucomont, 1932" ; dwc:species "ophion" ; dwc:subSpecies "confusus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Boheman, 1858" ; dwc:authorityName "Boheman" ; dwc:authorityYear "1858" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Boheman, 1858" ; dwc:species "insularis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Rossini, Vaz-de-Mello & Zunino, 2018" ; dwc:authorityName "Rossini" ; dwc:authorityYear "2018" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Rossini, 2018" ; dwc:species "basicarinatus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Rossini, Vaz-de-Mello & Zunino, 2018" ; dwc:authorityName "Rossini" ; dwc:authorityYear "2018" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Rossini, 2018" ; dwc:species "chacoensis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Blanchard, 1846" ; dwc:authorityName "Blanchard" ; dwc:authorityYear "1846" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Blanchard, 1846" ; dwc:species "rubrescens" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Harold, 1875" ; dwc:authorityName "Harold" ; dwc:authorityYear "1875" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Harold, 1875" ; dwc:species "haematopus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Zunino & Halffter, 1997" ; dwc:authorityName "Zunino & Halffter" ; dwc:authorityYear "1997" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Zunino & Halffter, 1997" ; dwc:species "onorei" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Harold, 1869" ; dwc:authorityName "Harold" ; dwc:authorityYear "1869" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Harold, 1869" ; dwc:species "crinitus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Bates, 1886" ; dwc:authorityName "Bates" ; dwc:authorityYear "1886" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Bates, 1886" ; dwc:species "crinitus" ; dwc:subSpecies "panamensis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Paulian, 1936" ; dwc:authorityName "Paulian" ; dwc:authorityYear "1936" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Paulian, 1936" ; dwc:species "embrikianus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Balthasar, 1939" ; dwc:authorityName "Balthasar" ; dwc:authorityYear "1939" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Balthasar, 1939" ; dwc:species "nabeleki" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Paulian, 1936" ; dwc:authorityName "Paulian" ; dwc:authorityYear "1936" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Paulian, 1936" ; dwc:species "schunckei" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Paulian, 1936" ; dwc:authorityName "Paulian" ; dwc:authorityYear "1936" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Paulian, 1936" ; dwc:species "catharinensis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "hircus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:genus "Onthophagus" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Scarabaeidae" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:order "Coleoptera" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "Fig. 3. A. Protibial apical tooth acuminate. B. Protibial apical tooth obtuse. C. Details of the elytral interstriae of Onthophagus marginicollis Harold, 1880 (hircus complex). Dā€“E. Detail of the elytral paler spots. F. Pronotal sulcus of O. haematopus Harold, 1875 (rubrescens complex). G. Detail of the basal carina of the cephalic horns. H. Shape of the clypeus and lateral margin of pronotum of a species of the ophion complex. I. Right antenna of species of the ophion complex. J. Right antenna of species of the osculatii complex. K. Detail of the metasternal punctuation of a species of the osculatii complex. Scale bars: 1 mm." ; fabio:hasRepresentation ; a fabio:Figure .