@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .
@prefix xlink: <http://www.w3.org/1999/xlink/> .

<http://treatment.plazi.org/id/112B87BCFFD9FFF8FF29FE71FBFAF9F4>
    cito:cites <http://dx.doi.org/10.5281/zenodo.10012477>, <http://dx.doi.org/10.5281/zenodo.10012480>, <http://dx.doi.org/10.5281/zenodo.10012482>, <http://dx.doi.org/10.5281/zenodo.10012522>, <http://dx.doi.org/10.5281/zenodo.10012475>, <http://dx.doi.org/10.5281/zenodo.10012526>, <http://dx.doi.org/10.5281/zenodo.10012469>, <http://dx.doi.org/10.5281/zenodo.10012471>, <http://dx.doi.org/10.5281/zenodo.10012473> ;
    dc:creator "Tan, Desmond Kai Teck; Fong, Jonathan J.; Zhang, Chao" ;
    dc:title "Heterobiantes geniculatus" ;
    dwc:basisOfRecord <http://tb.plazi.org/GgServer/dwcaRecords/112B87BCFFD9FFF8FF29FE71FBFAF9F4.mc.29EA3CF7FFD9FFF6FF41FC7AFF26FC08>, <http://tb.plazi.org/GgServer/dwcaRecords/112B87BCFFD9FFF8FF29FE71FBFAF9F4.mc.29EA3CF7FFD9FFF6FEBFFC26FEFCFBEC>, <http://tb.plazi.org/GgServer/dwcaRecords/112B87BCFFD9FFF8FF29FE71FBFAF9F4.mc.29EA3CF7FFD9FFF6FE8CFBC2FE88FBB0>, <http://tb.plazi.org/GgServer/dwcaRecords/112B87BCFFD9FFF8FF29FE71FBFAF9F4.mc.29EA3CF7FFD9FFF6FED3FBEEFC53FB78>, <http://tb.plazi.org/GgServer/dwcaRecords/112B87BCFFD9FFF8FF29FE71FBFAF9F4.mc.29EA3CF7FFD9FFF6FC30FBB6FD1DFB5C>, <http://tb.plazi.org/GgServer/dwcaRecords/112B87BCFFD9FFF8FF29FE71FBFAF9F4.mc.29EA3CF7FFD9FFF6FD6BFB52FB3DFB5C>, <http://tb.plazi.org/GgServer/dwcaRecords/112B87BCFFD9FFF8FF29FE71FBFAF9F4.mc.29EA3CF7FFD9FFF6FB4BFB52FBB0FB20>, <http://tb.plazi.org/GgServer/dwcaRecords/112B87BCFFD9FFF8FF29FE71FBFAF9F4.mc.29EA3CF7FFD9FFF6FBCCFB7EFE88FB04>, <http://tb.plazi.org/GgServer/dwcaRecords/112B87BCFFD9FFF8FF29FE71FBFAF9F4.mc.29EA3CF7FFD9FFF6FED9FB1AFCA6FB04>, <http://tb.plazi.org/GgServer/dwcaRecords/112B87BCFFD9FFF8FF29FE71FBFAF9F4.mc.29EA3CF7FFD9FFF6FCC6FB1AFD8DFAE8> ;
    trt:augmentsTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Heterobiantes_geniculatus_Pocock_1903> ;
    trt:deprecates <http://taxon-concept.plazi.org/id/Animalia/Epedanus_geniculatus_1903_1903>, <http://taxon-concept.plazi.org/id/Animalia/Pseudobiantes_geniculatus_Hirst_1911>, <http://taxon-concept.plazi.org/id/Animalia/Epedanestus_geniculatus_Roewer_1938>, <http://taxon-concept.plazi.org/id/Animalia/Heterobiantes_geniculatus_Roewer_1912> ;
    trt:publishedIn <http://dx.doi.org/10.11646/zootaxa.5353.6.2> ;
    a trt:Treatment .

<http://dx.doi.org/10.11646/zootaxa.5353.6.2>
    bibo:endPage "550" ;
    bibo:issue "6" ;
    bibo:journal "Zootaxa" ;
    bibo:pubDate "2023-10-13" ;
    bibo:startPage "533" ;
    bibo:volume "5353" ;
    dc:creator "Tan, Desmond Kai Teck; Fong, Jonathan J.; Zhang, Chao" ;
    dc:date "2023" ;
    dc:title "Re-description of the genus Heterobiantes and its type species H. geniculatus (Opiliones: Laniatores: Epedanidae) with additional genital and external morphology data" ;
    fabio:hasPart <http://dx.doi.org/10.5281/zenodo.10012477>, <http://dx.doi.org/10.5281/zenodo.10012480>, <http://dx.doi.org/10.5281/zenodo.10012482>, <http://dx.doi.org/10.5281/zenodo.10012522>, <http://dx.doi.org/10.5281/zenodo.10012475>, <http://dx.doi.org/10.5281/zenodo.10012526>, <http://dx.doi.org/10.5281/zenodo.10012469>, <http://dx.doi.org/10.5281/zenodo.10012471>, <http://dx.doi.org/10.5281/zenodo.10012473> ;
    a fabio:JournalArticle .

<http://taxon-concept.plazi.org/id/Animalia/Heterobiantes_geniculatus_Pocock_1903>
    dwc:authority "(Pocock, 1903)" ;
    dwc:baseAuthorityName "Pocock" ;
    dwc:baseAuthorityYear "1903" ;
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Heterobiantes" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Pocock, 1903)" ;
    dwc:species "geniculatus" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/7WLKB> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Heterobiantes_geniculatus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Epedanus_geniculatus_1903_1903>
    # Warning: Removing "(Pocock" from authority ;
    dwc:authority "Pocock, 1903: 446" ;
    dwc:authorityName ", (Pocock" ;
    dwc:authorityPageNumber "446" ;
    dwc:authorityYear "1903" ;
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Epedanus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "1903" ;
    dwc:species "geniculatus" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/7WLKB> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Epedanus_geniculatus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Pseudobiantes_geniculatus_Hirst_1911>
    dwc:authority "Hirst, 1911: 633" ;
    dwc:authorityName "Hirst" ;
    dwc:authorityPageNumber "633" ;
    dwc:authorityYear "1911" ;
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Pseudobiantes" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Hirst, 1911" ;
    dwc:species "geniculatus" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/7WLKB> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Pseudobiantes_geniculatus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Epedanestus_geniculatus_Roewer_1938>
    dwc:authority "Roewer, 1938: 109" ;
    dwc:authorityName "Roewer" ;
    dwc:authorityPageNumber "109" ;
    dwc:authorityYear "1938" ;
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Epedanestus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Roewer, 1938" ;
    dwc:species "geniculatus" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/7WLKB> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Epedanestus_geniculatus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Heterobiantes_geniculatus_Roewer_1912>
    dwc:authority "Roewer, 1912: 219" ;
    dwc:authorityName "Roewer" ;
    dwc:authorityPageNumber "219" ;
    dwc:authorityYear "1912" ;
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Heterobiantes" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Roewer, 1912" ;
    dwc:species "geniculatus" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/7WLKB> ;
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<http://taxon-name.plazi.org/id/Animalia/Heterobiantes_geniculatus>
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Heterobiantes" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:species "geniculatus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Heterobiantes> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Heterobiantes>
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Heterobiantes" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Epedanidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Epedanidae>
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "family" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Opiliones> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Opiliones>
    dwc:class "Arachnida" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "order" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Arachnida> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Arachnida>
    dwc:class "Arachnida" ;
    dwc:kingdom "Animalia" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "class" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Arthropoda> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Arthropoda>
    dwc:kingdom "Animalia" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "phylum" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia>
    dwc:kingdom "Animalia" ;
    dwc:rank "kingdom" ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Epedanus_geniculatus>
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Epedanus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:species "geniculatus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Epedanus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Epedanus>
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Epedanus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Epedanidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Pseudobiantes_geniculatus>
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Pseudobiantes" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:species "geniculatus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Pseudobiantes> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Pseudobiantes>
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Pseudobiantes" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Epedanidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Epedanestus_geniculatus>
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Epedanestus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:species "geniculatus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Epedanestus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Epedanestus>
    dwc:class "Arachnida" ;
    dwc:family "Epedanidae" ;
    dwc:genus "Epedanestus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Opiliones" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Epedanidae> ;
    a dwcFP:TaxonName .

<http://dx.doi.org/10.5281/zenodo.10012477>
    dc:description "FIGURE 7. Comparisons of apical structures of male genitals. A – C Penis comparisons of ‘spoon’ and ‘cup’ form between H. geniculatus and other Epedanidae species; A Pseudobiantes japonicus Hirst, 1911, taken from Kury et al. 2019; B Heterobiantes geniculatus (Pocock, 1903); C Parabeloniscus corneus Zhang & Zhang, 2012; D – F Examples and comparisons of ‘prickly funnels’ in everted Epedanidae genitals, all genitals beside H. geniculatus are taken from Zhang & Martens 2020; D Toccolus globitarsis Suzuki, 1969; E Heterobiantes geniculatus (Pocock, 1903); F Euepedanus dashdamirovi Zhang & Martens, 2020; G – N Comparison of lateral genitals of male Epedanidae; G & N Heterobiantes geniculatus (Pocock, 1903); H Toccolus globitarsis Suzuki, 1969, taken from Zhang & Martens, 2020; I Epedanus pictus Thorell, 1876, taken from Kury et al. 2019; J Euepedanus dashdamirovi Zhang & Martens, 2020, taken from Zhang & Martens 2020; K Pseudobiantes japonicus Hirst, 1911, taken from Kury & Villarreal 2015; L Kilungius bimaculatus Roewer, 1915, taken from Suzuki 1973; M Kilungius insulanus (Hirst, 1911), taken from Suzuki 1973." ;
    fabio:hasRepresentation <https://zenodo.org/record/10012477/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.10012480>
    dc:description "FIGURE 8. Comparison of dorsal and ventral genitals of male Epedanidae. A & H Kilungius bimaculatus Roewer, 1915, taken from Suzuki 1973; B, E & I Kilungius insulanus (Hirst, 1911), taken from Suzuki 1973; C & J Plistobunus columnaris Lian et al., 2011; D Pseudobiantes japonicus Hirst, 1911, taken from Kury & Villarreal 2015; F & M Epedanus pictus Thorell, 1876, taken from Kury et al. 2019; G & N Euepedanus dashdamirovi Zhang & Martens, 2020, taken from Zhang & Martens 2020; K Toccolus globitarsis Suzuki, 1969, taken from Zhang & Martens 2020; L Heterobiantes geniculatus (Pocock, 1903)." ;
    fabio:hasRepresentation <https://zenodo.org/record/10012480/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.10012482>
    dc:description "FIGURE 9. Type specimen tags found with the specimens studied by Roewer, photo taken by the British Museum of Natural History (BMNH)." ;
    fabio:hasRepresentation <https://zenodo.org/record/10012482/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.10012522>
    dc:description "FIGURE 1. Heterobiantes geniculatus (Pocock, 1903), males, body. A lateral view (LINGU-INV02); B dorsal view (LINGU- INV06); C ventral view (LINGU-INV06). Scale bars: 1 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/10012522/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.10012475>
    dc:description "FIGURE 6. Photograph comparing chelicerae and the dorsal body of Heterobiantes geniculatus (Pocock, 1903) on sexual dimorphism.A Chelicerae of male (LINGU-INV06); B Dorsal body of male (LINGU-INV06); C Chelicerae of female (LINGU- INV07); D Dorsal body of female (LINGU-INV07). Scale bar: 1 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/10012475/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.10012526>
    dc:description "FIGURE 2. Heterobiantes geniculatus (Pocock, 1903), male (A–D, LINGU-INV06), male (E–F, LINGU-INV05), chelicera.A Distal segment of left chelicera, frontal view; B Distal segment of right chelicera, frontal view; C Left chelicera, lateral view; D Right chelicera, lateral view; E Full right chelicera, lateral view; F Full left chelicera, lateral view. Scale bars: 1mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/10012526/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.10012469>
    dc:description "FIGURE 3. Heterobiantes geniculatus (Pocock, 1903), male (LINGU-INV06). A Close-up of pedipalpal tibia-tarsus-claw, lateral view; B Right pedipalp, lateral view; C Left pedipalp, lateral view; D Leg I, lateral view; E Leg II, lateral view; F Leg III, lateral view; G Leg IV, lateral view. Scale bars: 1 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/10012469/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.10012471>
    dc:description "FIGURE 4. Heterobiantes geniculatus (Pocock, 1903). A photograph of the distitarsus I (LINGU-INV01); B Close-up of the distitarsus I (LINGU-INV06); C Scanning electron microscopic leg IV tarsus (LINGU-INV01), a part of the distitarsus is missing, but the lack of scopulae is still clearly shown; D Scanning electron microscopic full dorso-lateral view of male genitalia (LINGU-INV10); E Scanning electron microscopic lateral view of expanded male genitalia (LINGU-INV08), arrows indicate how the lobes unfold and expand; F Scanning electron microscopic dorsal view of unexpanded male genitalia (LINGU-INV10), arrows indicate how the lobes unfurl and the direction of penial expansion. Scale bar: 1 mm (A, B); 500 µm (C); 100 µm (D, E, F)." ;
    fabio:hasRepresentation <https://zenodo.org/record/10012471/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.10012473>
    dc:description "FIGURE 5. Heterobiantes geniculatus (Pocock, 1903), penis. A apical structure (LINGU-INV10), ventral view; B apical structure (LINGU-INV17), lateral view; C full penis (LINGU-INV04), lateral view. Scale bars: 200 µm." ;
    fabio:hasRepresentation <https://zenodo.org/record/10012473/files/figure.png> ;
    a fabio:Figure .

<http://tb.plazi.org/GgServer/dwcaRecords/112B87BCFFD9FFF8FF29FE71FBFAF9F4.mc.29EA3CF7FFD9FFF6FF41FC7AFF26FC08>
    dwc:catalogNumber "LINGU-INV01" ;
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    dwc:verbatimLongitude "114.01322" ;
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    dwc:catalogNumber "LINGU-INV05" ;
    dwc:collectionCode "LINGU-INV" ;
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    dwc:catalogNumber "LINGU-INV06" ;
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    dwc:catalogNumber "LINGU-INV07, LINGU-INV02, LINGU-INV03, LINGU-INV04, LINGU-INV08, LINGU-INV09, LINGU-INV10" ;
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    dwc:eventDate "2022-05-24" ;
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