@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . # Warning: Not adding 'trt:citesTaxonName ' due to issues with rank ; cito:cites , , , , , , , , , , , , , , , , ; dc:creator "Humara-Gil, Karla J.; Granja-Fernández, Rebeca; Bautista-Guerrero, Eric; Rodríguez-Troncoso, Alma P." ; dc:title "Ophioderma occultum Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso 2022, sp. nov." ; dwc:basisOfRecord , , , , , , , , ; trt:citesTaxonName , , , ; trt:definesTaxonConcept ; trt:deprecates , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "384" ; bibo:issue "5 - 8" ; bibo:journal "Journal of Natural History" ; bibo:pubDate "2022-06-09" ; bibo:series "J. Nat. Hist." ; bibo:startPage "365" ; bibo:volume "56" ; dc:creator "Humara-Gil, Karla J.; Granja-Fernández, Rebeca; Bautista-Guerrero, Eric; Rodríguez-Troncoso, Alma P." ; dc:date "2022" ; dc:title "Overlooked for over a century: Ophioderma occultum sp. nov. (Echinodermata), a new species of brittle star from the Eastern Pacific" ; fabio:hasPart , , , , ; a fabio:JournalArticle . dwc:authority "Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" ; dwc:authorityName "Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" ; dwc:authorityYear "2022" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Humara-Gil, Granja-Fernández, Bautista-Guerrero & Rodríguez-Troncoso, 2022" ; dwc:species "occultum" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Lütken" ; dwc:baseAuthorityYear "1859" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiuridae" ; dwc:genus "Ophiura" ; dwc:kingdom "Animalia" ; dwc:order "Ophiurida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Lütken, 1859)" ; dwc:species "panamensis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Lutken" ; dwc:authorityYear "1859" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lutken, 1859" ; dwc:species "panamense" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Lyman" ; dwc:authorityYear "1860" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiuridae" ; dwc:genus "Ophiura" ; dwc:kingdom "Animalia" ; dwc:order "Ophiurida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lyman, 1860" ; dwc:species "teres" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "H. L. Clark" ; dwc:authorityYear "1917" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Clark, 1917" ; dwc:species "pentacanthum" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Caso" ; dwc:authorityYear "1986" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Caso, 1986" ; dwc:species "sodipallaresi" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Hendler" ; dwc:authorityYear "1996" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Hendler, 1996" ; dwc:species "vansyoci" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Lutken" ; dwc:authorityYear "1856" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lutken, 1856" ; dwc:species "variegatum" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Muller and Troschel" ; dwc:authorityYear "1840" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Muller & Troschel, 1840" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Lutken, 1856" ; dwc:authorityName "Lutken" ; dwc:authorityYear "1856" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lutken, 1856" ; dwc:species "brevicaudum" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Granja-Fernandez et al. 2020" ; dwc:authorityName "Granja-Fernandez" ; dwc:authorityYear "2020" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Granja-Fernandez, 2020" ; dwc:species "hendleri" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ziesenhenne 1955" ; dwc:authorityName "Ziesenhenne" ; dwc:authorityYear "1955" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ziesenhenne, 1955" ; dwc:species "panamense" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Lutken, 1856" ; dwc:authorityName "Lutken" ; dwc:authorityYear "1856" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lutken, 1856" ; dwc:species "rubicundum" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Lyman" ; dwc:baseAuthorityYear "1860" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:isUncertain "true" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Lyman, 1860)" ; dwc:species "teres" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "H. L. Clark" ; dwc:authorityYear "1940" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Clark, 1940" ; dwc:species "teres" ; dwc:subSpecies "unicolor" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "(COI)" ; dwc:baseAuthorityName "COI" ; dwc:class "Ophiuroidea" ; dwc:family "Ophiopezidae" ; dwc:genus "Ophiopeza" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Coi)" ; dwc:species "fallax" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:species "occultum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Ophiodermatidae" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:order "Ophiacanthida" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Ophiuroidea" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:phylum "Echinodermata" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Ophiuroidea" ; dwc:family "Ophiuridae" ; dwc:genus "Ophiura" ; dwc:kingdom "Animalia" ; dwc:order "Ophiurida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:species "panamensis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:genus "Ophiura" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Ophiuridae" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:order "Ophiurida" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Ophiuroidea" ; dwc:family "Ophiodermatidae" ; dwc:genus "Ophioderma" ; dwc:kingdom "Animalia" ; dwc:order "Ophiacanthida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:species "panamense" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Ophiuroidea" ; dwc:family "Ophiuridae" ; dwc:genus "Ophiura" ; dwc:kingdom "Animalia" ; dwc:order "Ophiurida" ; dwc:phylum "Echinodermata" ; dwc:rank "species" ; dwc:species "teres" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 3. Morphological variations recorded in Ophioderma occultum sp. nov. (a) Disc with partially uncoveredradial shields (arrowheads;CE-UAM 390,DD = 15.4mm);(b) white-spottedoral shields (paratype: ICML-EMU 12868, DD = 19.9 mm); (c) white oral shields (CE-UAM 214, DD = 7.3 mm); (d) granules on the proximalmost DAPs (ICML-UNAM 11593, DD = 26.7 mm); (e) arm segment with three tentacle scales (arrowhead; paratype: ICML-UNAM 18430, DD = 17.2 mm); (f) abnormal arm spines (arrowheads; CE-UAM 679,DD = 16.3 mm); (g) brown-coloured specimen (CE-UAM 673, DD = 23.5 mm); (h–i) in vivo colouration (Photographs: M.A. Medina-López)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 4. Some morphological characters of Eastern Pacific Ophioderma that differ from those of Ophioderma occultum sp. nov. Ophioderma hendleri (holotype: ICML-UNAM 18315, DD = 14.8 mm): (a) close-up of a jaw; (b) dorsal arm. Ophioderma panamense (holotype: NHMD-107679, DD = 21.9 mm): (c) portion of dorsal disc and arm. Ophioderma pentacanthum (paratype: MCZ OPH-4519," ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 5. Maximum likelihood tree of the species analysed in this study. (a) COI; (b) 16S. Values at branches indicate bootstrap values (maximum likelihood/neighbour-joining); asterisks indicate specimens of Ophioderma occultum sp. nov. previously identified as Ophioderma panamense. Sequences obtained in this study are indicated in bold. See the Supplemental material for more information." ; fabio:hasRepresentation ; a fabio:Figure . dwc:catalogNumber "ICML-EMU 12867" ; dwc:collectionCode "COI" ; dwc:eventDate "2019-11-07" ; dwc:locality "Punta Pichilingue" ; dwc:recordedBy "K. J. Humara-Gil & R. Granja-Fernandez & B. Maya-Alvarado & A. Hernandez-Morales & M. A. Medina-Lopez" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "2" ; dwc:verbatimLatitude "24.285" ; dwc:verbatimLongitude "-110.334" ; trt:gbifOccurrenceId "3819603308" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ICML-EMU 12867" ; dwc:collectionCode "COI" ; dwc:eventDate "2019-11-07" ; dwc:locality "Punta Pichilingue" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2" ; dwc:verbatimLatitude "24.285" ; dwc:verbatimLongitude "-110.334" ; trt:gbifOccurrenceId "3819603305" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ICML-EMU 12867" ; dwc:collectionCode "R, COI" ; dwc:eventDate "2019-11-07" ; dwc:locality "Punta Pichilingue" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "2" ; dwc:verbatimLatitude "24.285" ; dwc:verbatimLongitude "-110.334" ; trt:gbifOccurrenceId "3819603302" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ICML-EMU 12867" ; dwc:collectionCode "R, COI" ; dwc:eventDate "2019-11-07" ; dwc:locality "Punta Pichilingue" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "2" ; dwc:verbatimLatitude "24.285" ; dwc:verbatimLongitude "-110.334" ; trt:gbifOccurrenceId "3819603301" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ICML-EMU 12868" ; dwc:collectionCode "R, COI" ; dwc:eventDate "2019-11-07" ; dwc:locality "Punta Pichilingue" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "2" ; dwc:verbatimLatitude "24.285" ; dwc:verbatimLongitude "-110.334" ; trt:gbifOccurrenceId "3819603309" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ICML-UNAM 3.24" ; dwc:collectionCode "ICML-UNAM" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3819603303" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "OPH-112" ; dwc:collectionCode "MCZ" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3819603304" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "OPH-114" ; dwc:collectionCode "MCZ" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3819603307" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "LACM 90, 391.1" ; dwc:collectionCode "LACM" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3819603306" ; trt:httpUri ; a dwc:MaterialCitation .