@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .

<http://treatment.plazi.org/id/03A1C039E21CFFCAFF08472EFCDAFEF7>
    cito:cites <http://dx.doi.org/10.5281/zenodo.7863394>, <http://dx.doi.org/10.5281/zenodo.7863398>, <http://dx.doi.org/10.5281/zenodo.7863400>, <http://dx.doi.org/10.5281/zenodo.7863388>, <http://dx.doi.org/10.5281/zenodo.7863392>, <http://dx.doi.org/10.5281/zenodo.7863404>, <http://dx.doi.org/10.5281/zenodo.7863406>, <http://taxon-concept.plazi.org/id/Animalia/Otus_murivorus_Milne_1873>, <http://taxon-concept.plazi.org/id/Animalia/Papasula_Olson_1988>, <http://taxon-concept.plazi.org/id/Animalia/Papasula_abbotti_Cheke_2008>, <http://taxon-concept.plazi.org/id/Animalia/Sula_sula_Linnaeus_1766> ;
    dc:creator "Hume, Julian P." ;
    dc:title "Papasula abbotti subsp. nelsoni Hume 2023, ssp. nov." ;
    dwc:basisOfRecord <http://tb.plazi.org/GgServer/dwcaRecords/03A1C039E21CFFCAFF08472EFCDAFEF7.mc.3B607B72E21BFFC8FF084108FD3FFE83> ;
    trt:citesTaxonName <http://taxon-name.plazi.org/id/Animalia/Cylindraspis>, <http://taxon-name.plazi.org/id/Animalia/Papasula_undetermined> ;
    trt:definesTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Papasula_abbotti_nelsoni_Hume_2023> ;
    trt:deprecates <http://taxon-concept.plazi.org/id/Animalia/Sula_abbotti_Cheke_2001>, <http://taxon-concept.plazi.org/id/Animalia/Sula_abbotti_Nelson_1974>, <http://taxon-concept.plazi.org/id/Animalia/Sula_capensis_Milne_1875>, <http://taxon-concept.plazi.org/id/Animalia/Sula_piscator_Milne_1873> ;
    trt:publishedIn <http://dx.doi.org/10.11646/zootaxa.5270.3.5> ;
    a trt:Treatment .

<http://dx.doi.org/10.11646/zootaxa.5270.3.5>
    bibo:endPage "536" ;
    bibo:issue "3" ;
    bibo:journal "Zootaxa" ;
    bibo:pubDate "2023-04-21" ;
    bibo:startPage "507" ;
    bibo:volume "5270" ;
    dc:creator "Hume, Julian P." ;
    dc:date "2023" ;
    dc:title "A new fossil subspecies of booby (Aves, Sulidae: Papasula) from Mauritius and Rodrigues, Mascarene Islands, with notes on P. abbotti from Assumption Island" ;
    fabio:hasPart <http://dx.doi.org/10.5281/zenodo.7863394>, <http://dx.doi.org/10.5281/zenodo.7863398>, <http://dx.doi.org/10.5281/zenodo.7863400>, <http://dx.doi.org/10.5281/zenodo.7863388>, <http://dx.doi.org/10.5281/zenodo.7863392>, <http://dx.doi.org/10.5281/zenodo.7863404>, <http://dx.doi.org/10.5281/zenodo.7863406> ;
    a fabio:JournalArticle .

<http://taxon-concept.plazi.org/id/Animalia/Papasula_abbotti_nelsoni_Hume_2023>
    dwc:authority "Hume, 2023" ;
    dwc:authorityName "Hume" ;
    dwc:authorityYear "2023" ;
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Papasula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Suliformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "subSpecies" ;
    dwc:scientificNameAuthorship "Hume, 2023" ;
    dwc:species "abbotti" ;
    dwc:subSpecies "nelsoni" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Papasula_abbotti_nelsoni> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Sula_piscator_Milne_1873>
    dwc:authority "Milne-Edwards, 1873, p. 22" ;
    dwc:authorityName "Milne-Edwards" ;
    dwc:authorityPageNumber "22" ;
    dwc:authorityYear "1873" ;
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Sula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Pelecaniformes", "Suliformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Milne-Edwards, 1873" ;
    dwc:species "piscator" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Sula_piscator> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Sula_capensis_Milne_1875>
    dwc:authority "Milne-Edwards, 1875, p. 8" ;
    dwc:authorityName "Milne-Edwards" ;
    dwc:authorityPageNumber "8" ;
    dwc:authorityYear "1875" ;
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Sula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Suliformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Milne-Edwards, 1875" ;
    dwc:species "capensis" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Sula_capensis> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Sula_abbotti_Nelson_1974>
    dwc:authority "Nelson, 1974" ;
    dwc:authorityName "Nelson" ;
    dwc:authorityYear "1974" ;
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Sula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Pelecaniformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Nelson, 1974" ;
    dwc:species "abbotti" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Sula_abbotti> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Sula_abbotti_Cheke_2001>
    dwc:authority "Cheke, 2001, p. 73" ;
    dwc:authorityName "Cheke" ;
    dwc:authorityPageNumber "73" ;
    dwc:authorityYear "2001" ;
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Sula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Suliformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Cheke, 2001" ;
    dwc:species "abbotti" ;
    dwc:subGenus "Papasula" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/53FBB> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Sula_abbotti> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Papasula_abbotti_Cheke_2008>
    dwc:authority ", Cheke & Hume, 2008, p. 47" ;
    dwc:authorityName ", Cheke & Hume" ;
    dwc:authorityPageNumber "47" ;
    dwc:authorityYear "2008" ;
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Papasula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Suliformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Cheke & Hume, 2008" ;
    dwc:species "abbotti" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/4CJ75> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Papasula_abbotti> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Papasula_Olson_1988>
    dwc:authorityName "Olson and Warheit" ;
    dwc:authorityYear "1988" ;
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Papasula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Suliformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "genus" ;
    dwc:scientificNameAuthorship "Olson & Warheit, 1988" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Papasula> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Otus_murivorus_Milne_1873>
    dwc:authority "(Milne-Edwards, 1873)" ;
    dwc:baseAuthorityName "Milne-Edwards" ;
    dwc:baseAuthorityYear "1873" ;
    dwc:class "Aves" ;
    dwc:family "Strigidae" ;
    dwc:genus "Otus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Ciconiiformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Milne-Edwards, 1873)" ;
    dwc:species "murivorus" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Otus_murivorus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Sula_sula_Linnaeus_1766>
    dwc:baseAuthorityName "Linnaeus" ;
    dwc:baseAuthorityYear "1766" ;
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Sula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Pelecaniformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Linnaeus, 1766)" ;
    dwc:species "sula" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Sula_sula> ;
    a dwcFP:TaxonConcept .

<http://taxon-name.plazi.org/id/Animalia/Papasula_abbotti_nelsoni>
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Papasula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Suliformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "subSpecies" ;
    dwc:species "abbotti" ;
    dwc:subSpecies "nelsoni" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Papasula_abbotti> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Papasula_abbotti>
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Papasula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Suliformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:species "abbotti" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Papasula> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Papasula>
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Papasula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Suliformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Sulidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Sulidae>
    dwc:family "Sulidae" ;
    dwc:rank "family" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Pelecaniformes>, <http://taxon-name.plazi.org/id/Animalia/Suliformes> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Suliformes>
    dwc:order "Suliformes" ;
    dwc:rank "order" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Aves> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Aves>
    dwc:class "Aves" ;
    dwc:rank "class" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Chordata> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Chordata>
    dwc:phylum "Chordata" ;
    dwc:rank "phylum" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia>
    dwc:kingdom "Animalia" ;
    dwc:rank "kingdom" ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Sula_piscator>
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Sula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Pelecaniformes", "Suliformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:species "piscator" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Sula> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Sula>
    dwc:genus "Sula" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Sulidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Sula_capensis>
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Sula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Suliformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:species "capensis" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Sula> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Sula_abbotti>
    dwc:class "Aves" ;
    dwc:family "Sulidae" ;
    dwc:genus "Sula" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Pelecaniformes", "Suliformes" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:species "abbotti" ;
    dwc:subGenus "Papasula" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Sula>, <http://taxon-name.plazi.org/id/Animalia/Sula_Papasula> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Pelecaniformes>
    dwc:order "Pelecaniformes" ;
    dwc:rank "order" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Aves> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Sula_Papasula>
    dwc:rank "subGenus" ;
    dwc:subGenus "Papasula" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Sula> ;
    a dwcFP:TaxonName .

<http://dx.doi.org/10.5281/zenodo.7863394>
    dc:description "FIGURE 4. Comparison of humeri of Papasula a. nelsoni ssp. nov. with Papasula a. abbotti, caudal surface, left side. A. NHMUK PVA9049 P. a. nelsoni (Plaine Corail, Rodrigues); B. UMZC 262.AA holotype P. a. nelsoni (Mare aux Songes, Mauritius); C. USNM 560863 P. a. abbotti u/s (Christmas Island); USNM 560862 P. a. abbotti immature u/s (Christmas Island). Abbreviations: td, tuberculum dorsale; tv, tuberculum ventrale; ch, caput humeri; ic, incisura capitis; mc, margo caudalis; cvf, crus ventrale fossa; cb, crista bicipitalis; fp, fossa pneumotricipitalis; sh, sulcus humerotricipitalis; ss, sulcus scapulotricipitalis; pf, processus flexorius. Scale bar = 20 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/7863394/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7863398>
    dc:description "FIGURE 5. Comparison of proximal humeri of Papasula a. nelsoni ssp. nov. with Papasula a. abbotti, cranial surface, left side. A. NHMUK PVA9049 P. a. nelsoni (Plaine Corail, Rodrigues); B. UMZC 262.AA holotype P. a. nelsoni (Mare aux Songes, Mauritius); C. ANWC 40544 P. a. abbotti F (Christmas Island). Abbreviations: st, sulcus lig. transversus; cb, crista bicipitalis. Scale bar = 10 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/7863398/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7863400>
    dc:description "FIGURE 6. Comparison of ulnae of Papasula a. nelsoni ssp. nov. with Papasula a. abbotti, left side, A, B, C, ventral aspect, D, E, F, dorsal aspect.A & D. USNM 560862 P. a. abbotti immature u/s (Christmas Island); B & E. USNM 560863 P. a. abbotti u/s (Christmas Island); C & F UMZC 262.AAparatype P. a. nelsoni (Mare aux Songes, Mauritius).Abbreviations: ib, impressio brachialis; prv, papillae remigales ventralis; si, sulcus intercondylaris; dr, depressio radialis; cv, cotyla ventralis; tc, tuberculum carpale; cvu, condylus ventralis ulnaris. Scale bar = 20 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/7863400/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7863388>
    dc:description "FIGURE 2. Comparison of coracoid of Papasula a. nelsoni ssp. nov. (partially reconstructed) with Papasula a. abbotti, ventral aspect, left side. A. USNM 560863 P. a. abbotti u/s (Christmas Island); B. NHMUK PV A9076 P. a. nelsoni (Plaine Corail, Rodrigues). Abbreviations: le, labrum externum; ia, impressio lig. acrocoracohumeralis. Scale bar = 10 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/7863388/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7863392>
    dc:description "FIGURE 3. Sternum fragment of Papasula a. nelsoni ssp. nov. NHMUK PV A9076. A. right lateral aspect; B. dorsal aspect; C. cranial aspect. Abbreviations: li, labrum interna; tle, tuberculum labri externa; sac, sulcus articularis coracoideus. Scale bar = 10 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/7863392/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7863404>
    dc:description "FIGURE 7. Comparison of tarsometatarsi of Papasula a. nelsoni ssp. nov. with Papasula a. abbotti, right side, A, B, dorsal aspect, C, D, plantar aspect. A & D. USNM 560863 P. a. abbotti u/s (Christmas Island); B & C UMZC 995 P. a. nelsoni (Plaine Corail, Rodrigues). Abbreviations: fvp, foramina vascularia proximalia; ttc, tuberositas m. tibialis cranialis; se, sulcus extensorius; iil. incisura intertrochlearis lateralis; iim, incisura intertrochlearis medialis; cpl, crista plantares lateralis; cpm, crista plantares medialis; fvd, foramen vasculare distale. Scale bar = 10 mm." ;
    fabio:hasRepresentation <https://zenodo.org/record/7863404/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.7863406>
    dc:description "FIGURE 8. The original illustration of UMZC 995 (Fig. 6B, 6C) from Milne-Edwards (1873: pl. 11, figs. 5, 5a, 2d, 2e), in which he tentatively referred it to Sula piscator (= S. sula). Since its execution, damage has occurred, in particular the crista medialis hypotarsi (shown in Fig. 2e) is now missing." ;
    fabio:hasRepresentation <https://zenodo.org/record/7863406/files/figure.png> ;
    a fabio:Figure .

<http://tb.plazi.org/GgServer/dwcaRecords/03A1C039E21CFFCAFF08472EFCDAFEF7.mc.3B607B72E21BFFC8FF084108FD3FFE83>
    dwc:catalogNumber "UMZC 262.AA" ;
    dwc:collectionCode "UMZC" ;
    dwc:typeStatus "holotype" ;
    trt:gbifOccurrenceId "4089740301" ;
    trt:httpUri <https://treatment.plazi.org/id/03A1C039E21CFFCAFF08472EFCDAFEF7#3B607B72E21BFFC8FF084108FD3FFE83> ;
    a dwc:MaterialCitation .