@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . cito:cites , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; dc:creator "Nunes, Rafael V.; Vaz-de-Mello, Fernando Z." ; dc:title "Dichotomius (Cephagonus) Luederwaldt 1929" ; trt:augmentsTaxonConcept ; trt:citesTaxonName , , , , , , , , ; trt:deprecates ; trt:publishedIn ; a trt:Treatment . bibo:endPage "2351" ; bibo:issue "37" ; bibo:journal "Journal of Natural History" ; bibo:pubDate "2020-01-13" ; bibo:startPage "2231" ; bibo:volume "53" ; dc:creator "Nunes, Rafael V.; Vaz-de-Mello, Fernando Z." ; dc:date "2019" ; dc:title "Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" ; fabio:hasPart , , , , , , , , , , , , , , , , ; a fabio:JournalArticle . dwc:authority "", "Luederwaldt 1929" ; dwc:authorityName "Luederwaldt" ; dwc:authorityYear "1929" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:scientificNameAuthorship "Luederwaldt, 1929" ; dwc:subGenus "Cephagonus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Erichson" ; dwc:authorityYear "1847" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Pinotus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Erichson, 1847" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "" ; dwc:authorityName "Burmeister" ; dwc:authorityYear "1846" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:scientificNameAuthorship "Burmeister, 1846" ; dwc:subGenus "Selenocopris" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Nunes & Vaz-de-Mello" ; dwc:authorityYear "2019" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Nunes & Vaz-de-Mello, 2019" ; dwc:species "barbarae" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Nunes & Vaz-de-Mello" ; dwc:authorityYear "2019" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Nunes & Vaz-de-Mello, 2019" ; dwc:species "frizzasae" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Luederwaldti" ; dwc:baseAuthorityYear "1935" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Luederwaldti, 1935)" ; dwc:species "ingens" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Germar 1824" ; dwc:authorityName "Germar" ; dwc:authorityYear "1824" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Copris" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Germar, 1824" ; dwc:species "bicuspis" ; dwc:subGenus "Selenocopris" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Haroldi 1867" ; dwc:authorityName "Haroldi" ; dwc:authorityYear "1867" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Pinotus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Haroldi, 1867" ; dwc:species "fissus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Hope" ; dwc:authorityYear "1838" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Hope, 1838" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Latreille" ; dwc:authorityYear "1802" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; dwc:scientificNameAuthorship "Latreille, 1802" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Linnaeus" ; dwc:authorityYear "1758" ; dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; dwc:scientificNameAuthorship "Linnaeus, 1758" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "(Harold)" ; dwc:baseAuthorityName "Harold" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Pinotus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Harold)" ; dwc:species "fissus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Luederwaldti" ; dwc:authorityYear "1929" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Pinotus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Luederwaldti, 1929" ; dwc:species "socius" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Luederwaldt" ; dwc:baseAuthorityYear "1928" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Luederwaldt, 1928)" ; dwc:species "spadiceus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Nunes & Vaz-de-Mello" ; dwc:authorityYear "2019" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Nunes & Vaz-de-Mello, 2019" ; dwc:species "lucianomourai" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Nunes & Vaz-de-Mello" ; dwc:authorityYear "2019" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Nunes & Vaz-de-Mello, 2019" ; dwc:species "vargasae" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Nunes & Vaz-de-Mello" ; dwc:authorityYear "2019" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Nunes & Vaz-de-Mello, 2019" ; dwc:species "rondoniaensis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "(Luederwaldt)" ; dwc:baseAuthorityName "Luederwaldt" ; dwc:baseAuthorityYear "1930" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Luederwaldt, 1930)" ; dwc:species "punctulatipennis" ; dwc:subGenus "Selenocopris" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Nunes & Vaz-de-Mello" ; dwc:authorityYear "2019" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Nunes & Vaz-de-Mello, 2019" ; dwc:species "baiano" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Haroldi" ; dwc:authorityYear "1869" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Pinotus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Haroldi, 1869" ; dwc:species "mundus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:subGenus "Cephagonus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Dichotomius" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Pinotus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "Figure 7. (a) Mandible of D. fissus (Harold), inc – incisory lobe (membranous), mo – molar lobe. (b) Maxilla and maxillary palp of same species, mxp – maxillary palp, ga – galea, las – lacinia. (c) Epipharynx of same species. (d) Labrum and labial palpomere of same species. (e) Ventral clypeal process of D. fissceps indicated by white arrow.(f) Antena of same species.esc – escapo, lam – lamella. (g) Macropterous hind wig of D. fissiceps. (h) Brachypterous hindwing of D. ingens." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 8. (a) Part of D. ascanius ventral portion evidencing metasternum puncture, cox – coxae ely – elytra, mes – mesepisternum, met – metasternum. (b) Pygidium of D. reichei (c) abdominal ventrites of a male of D. ascaniusı (d) abdominal ventrites of a female of D. ascaniusı (e) abdominal ventrites of a female of D. sociusı (f) detail of the sixth ventrite of a female of D. Socius." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 11. Geographical records of Dichotomius (Cephagonus)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 4. (a) Intern sac sclerites of D. fissiceps. (b) Axial sclerites of D. reichei. (c) Axial sclerites of D. barbarae. (d) Lamella copulatrix of D. reichei. (e) Lamella copulatrix of D. frizzasae.(f) Lamella copulatrix of D. fissiceps. (g) Lamella copulatrix of D. barbarae." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 1. Dichotomius Hope morphological structures." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 2. Dichotomius Aedeagous: (a) lateral view; (b) dorsal view of paramera; (c) ventral view of paramera (D. quadraticeps); (d) aedeagous with everted inter sac (D. ingens)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 5. (a) Ventral view of the open pygidium of a female of D. ascanius: sixth – sixth ventrite, int – intestine;vgn – approximately location of vagina aperture;spt – spermatecha indicated by white arrow.(b) General dorsal aspect of Cephagonus female genitalia, the elytra and abdominal superior ventrites were removed: pyg – pygidium; spt – spermatheca; spdc – spermatheca duct; spgl – spermatheca gland; bcpl – bursa copulatrix. (c) D. ascanius spermatheca. (d) D. ingens spermatheca. (e) D. fissiceps spermatheca." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 13. Types of elytral disc surfaces and striae: (a) smooth and shiny elytral interstriae with deeply impressed striae and well defined puncture of D. reichei (female); (b) Eltral striae and interstriae of D. fissus (female); (c) elytral striae and interstriae of D. Baiano (female); (d) detail of the microsculpture in C (35x); (e) detail of C (50x); (f) elytral striae and interstriae of D. Mundus (male); (g) detail of chagrination in E (35x)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 14. Types of punctures and head/pronotal disc surfaces: (a) smooth surface with fine puncture; (b) ocellate punctures type I; (c) elliptical punctures; (d) ocellate punctures type II; (e) fine punctures (pronotum) and coarse punctures (head, indicated by white arrow); (f) clypeus and gena surfacewrinkled, wrinkled." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 33. D. arnaudi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view (grey arrow indicates the position of longitudinal excavation on lateral view). D. asenjoi: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. barbarae (i) male dorsal habitus; (j) paramera dorsal view (black arrow indicating slight medial constriction); (k) paramera ventral view; (l) paramera lateral view (black arrow indicates the surface of longitudinal excavation, which is smooth with fine punctures (see D. paschoali) and the differentiation of both species)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 35. D. blancoi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view; (e) detail of the carena found on the middle of paramera longitudinal excavation. D. candangoi: (f) male dorsal habitus; (g) paramera dorsal view; (h) paramera ventral view; (i) paramera lateral view." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 39. D. emas: (a) male dorsal habitus; (b) paramera dorsal view (black arrow indicating strong medial constriction on paramera side); (c) paramera ventral view (black arrow indicating strong medial constriction on paramera side); (d) paramera lateral view. D. marcoscarvalhoi: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. paschoali (i) male dorsal habitus; (j) paramera dorsal view; (k) paramera ventral view; (l) paramera lateral view (black arrow indicating basal excavation)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 19. D. fissus: (a) male dorsal habitus; (b) female dorsal habitus; (c) female ventral habitus; (d) paramera dorsal view; (e) paramera ventral view; (f) paramera lateral view; (g) type labels." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 23. D. fernandosilvai: (a) male dorsal habitus (Holotype); (b) female dorsal habitus; (c) female ventral habitus; (d) paramera dorsal view; (e) paramera ventral view; (f) paramera lateral view." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 32. D. clypeatus: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view; (e) type labels. D. aielloae: (f) Holotype dorsal habitus; (g) detail of male cephalic carina in frontal view; (h) type labels." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 37. D. edmondsi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view. D. frizzasae: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. gardneri (i) male dorsal habitus; (j) paramera dorsal view; (k) paramera ventral view; (l) paramera lateral view." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Figure 24. D. filgueirasi: (a) male dorsal habitus; (b) female dorsal habitus; (c) female ventral habitus; (d) paramera dorsal view; (e) paramera ventral view; (f) paramera lateral view." ; fabio:hasRepresentation ; a fabio:Figure .