@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , ; dc:creator "Bagaturov, Mikhail F.; Hillert, Oliver" ; dc:title "Lethrus (Paralethrus) bituberculatus subsp. impressifrons Ballion 1870" ; dwc:basisOfRecord , ; trt:augmentsTaxonConcept ; trt:citesTaxonName ; trt:deprecates , , , , , , , , , , , , , , , , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "316" ; bibo:issue "3" ; bibo:journal "Zootaxa" ; bibo:pubDate "2023-03-29" ; bibo:startPage "301" ; bibo:volume "5258" ; dc:creator "Bagaturov, Mikhail F.; Hillert, Oliver" ; dc:date "2023" ; dc:title "Sinolethrus, a new subgenus of the genus Lethrus Scopoli, 1777 from China (Coleoptera: Geotrupidae: Lethrinae) and new synonymy of the subgenus Paralethrus Nikolajev, 2003" ; fabio:hasPart , , , , , ; a fabio:JournalArticle . dwc:authority "Ballion, 1870" ; dwc:authorityName "Ballion" ; dwc:authorityYear "1870" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Ballion, 1870" ; dwc:species "bituberculatus" ; dwc:subGenus "Paralethrus" ; dwc:subSpecies "impressifrons" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Ballion, 1870: 336" ; dwc:authorityName "Ballion" ; dwc:authorityPageNumber "336" ; dwc:authorityYear "1870" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ballion, 1870" ; dwc:species "impressifrons" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Reitter 1890: 292" ; dwc:authorityName "Reitter" ; dwc:authorityPageNumber "292" ; dwc:authorityYear "1890" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "variety" ; dwc:scientificNameAuthorship "Reitter, 1890" ; dwc:species "laevigatus" ; dwc:variety "impressifrons" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Semenov 1892: 237" ; dwc:authorityName "Ballion" ; dwc:authorityPageNumber "237" ; dwc:authorityYear "1870" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ballion, 1870" ; dwc:species "laevigatus" ; dwc:subGenus "Autolethrus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Boucomont 1912: 37" ; dwc:authorityName "Boucomont" ; dwc:authorityPageNumber "37" ; dwc:authorityYear "1912" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Boucomont, 1912" ; dwc:species "hauseri" ; dwc:subGenus "Autolethrus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Semenov 1934: 1396" ; dwc:authorityName "Semenov" ; dwc:authorityPageNumber "1396" ; dwc:authorityYear "1934" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Semenov, 1934" ; dwc:species "hauseri" ; dwc:subGenus "Autolethrus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Nikolajev 1974: 81" ; dwc:authorityName "Nikolajev" ; dwc:authorityPageNumber "81" ; dwc:authorityYear "1974" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Nikolajev, 1974" ; dwc:species "hauseri" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Nikolajev 1969: 529" ; dwc:authorityName "Nikolajev" ; dwc:authorityPageNumber "529" ; dwc:authorityYear "1969" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Nikolajev, 1969" ; dwc:species "hauseri" ; dwc:subGenus "Autolethrus" ; dwc:subSpecies "hauseri" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Semenov 1894: 484" ; dwc:authorityName "Semenov" ; dwc:authorityPageNumber "484" ; dwc:authorityYear "1894" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Semenov, 1894" ; dwc:species "eous" ; dwc:subGenus "Autolethrus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Boucomont 1912: 37" ; dwc:authorityName "Boucomont" ; dwc:authorityPageNumber "37" ; dwc:authorityYear "1912" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Boucomont, 1912" ; dwc:species "eous" ; dwc:subGenus "Autolethrus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Semenov & Medvedev 1935: 285" ; dwc:authorityName "Semenov & Medvedev" ; dwc:authorityPageNumber "285" ; dwc:authorityYear "1935" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Semenov & Medvedev, 1935" ; dwc:species "jacobsoni" ; dwc:subGenus "Autolethrus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Semenov & Medvedev 1936: 87" ; dwc:authorityName "Semenov & Medvedev" ; dwc:authorityPageNumber "87" ; dwc:authorityYear "1936" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Semenov & Medvedev, 1936" ; dwc:species "jacobsoni" ; dwc:subGenus "Autolethrus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Protzenko 1972: 7" ; dwc:authorityName "Protzenko" ; dwc:authorityPageNumber "7" ; dwc:authorityYear "1972" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Protzenko, 1972" ; dwc:species "jacobsoni" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Skopin 1955: 150" ; dwc:authorityName "Skopin" ; dwc:authorityPageNumber "150" ; dwc:authorityYear "1955" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Skopin, 1955" ; dwc:species "nigroaeneus" ; dwc:subGenus "Autolethrus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Nikolajev 1969: 529" ; dwc:authorityName "Nikolajev" ; dwc:authorityPageNumber "529" ; dwc:authorityYear "1969" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Nikolajev, 1969" ; dwc:species "nigroaeneus" ; dwc:subGenus "Autolethrus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Nikolajev 1974: 81" ; dwc:authorityName "Nikolajev" ; dwc:authorityPageNumber "81" ; dwc:authorityYear "1974" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Nikolajev, 1974" ; dwc:species "bituberculatus" ; dwc:subGenus "Autolethrus" ; dwc:subSpecies "impressifrons" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Nikolajev 1987: 45" ; dwc:authorityName "Nikolajev" ; dwc:authorityPageNumber "45" ; dwc:authorityYear "1987" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Nikolajev, 1987" ; dwc:species "bituberculatus" ; dwc:subGenus "Lethrus" ; dwc:subSpecies "impressifrons" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Nikolajev 2003: 199" ; dwc:authorityName "Nikolajev" ; dwc:authorityPageNumber "199" ; dwc:authorityYear "2003" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Nikolajev, 2003" ; dwc:species "bituberculatus" ; dwc:subGenus "Paralethrus" ; dwc:subSpecies "impressifrons" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Fairmaire, 1892" ; dwc:authorityName "Fairmaire" ; dwc:authorityYear "1892" ; dwc:baseAuthorityName "Scopoli" ; dwc:baseAuthorityYear "1777" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "(Scopoli, 1777) Fairmaire, 1892" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ": Boucomont 1912: 37" ; dwc:authorityName ": Boucomont" ; dwc:authorityPageNumber "37" ; dwc:authorityYear "1912" ; dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Boucomont, 1912" ; dwc:species "mediocris" ; dwc:subGenus "Autolethrus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:species "bituberculatus" ; dwc:subGenus "Paralethrus", "Autolethrus", "Lethrus" ; dwc:subSpecies "impressifrons" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "bituberculatus" ; dwc:subGenus "Paralethrus", "Autolethrus", "Lethrus" ; trt:hasParentName , , ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:subGenus "Paralethrus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "impressifrons" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "variety" ; dwc:species "laevigatus" ; dwc:variety "impressifrons" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "laevigatus" ; dwc:subGenus "Autolethrus" ; trt:hasParentName , ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:subGenus "Autolethrus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "hauseri" ; dwc:subGenus "Autolethrus" ; trt:hasParentName , ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:species "hauseri" ; dwc:subGenus "Autolethrus" ; dwc:subSpecies "hauseri" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "eous" ; dwc:subGenus "Autolethrus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "jacobsoni" ; dwc:subGenus "Autolethrus" ; trt:hasParentName , ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "nigroaeneus" ; dwc:subGenus "Autolethrus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:subGenus "Lethrus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Geotrupidae" ; dwc:genus "Lethrus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "mediocris" ; dwc:subGenus "Autolethrus" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURES 36. Metafemur of Lethrus (Teratolethrus) baiocchii Hillert & Sechi, 2014 after Hillert & Sechi 2014)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 18–23. Head of Lethrus species with mandibular appendix, keels and hornlike apophyses of mandibles (males), schematically. 18–20 L. (Furcilethrus) banghaasi Reitter, 1893 (18 frontal view, 19 lateral view, 20 dorsal view); 21–23 L. (Furcilethrus) tenuidens tenuidens Reitter, 1890 (21 frontal view, 22 lateral view, 23 dorsal view)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 37–42. Head of Lethrus species with mandibular appendix, keels and hornlike apophyses of mandibles (males), schematically.37–39 L. (Lethrus) fallax Nikolajev, 1975 (37 view frontal, 38 lateral view, 39 dorsal view); 40–42 L.(Paralethrus) crassus Hillert, 2014 (40 frontal view, 41 lateral view, 42 dorsal view, after Hillert 2004)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 30–35. Head of Lethrus species with mandibular appendix, keels and hornlike apophyses of mandibles (males), schematically. 30–32 L. (Autolethrus) tshatkalensis Protzenko, 1965 (30 frontal view, 31 lateral view, 32 dorsal view); 33–35 L. (Teratolethrus) baiocchii Hillert & Sechi, 2014 (33 frontal view, 34 lateral view)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 1–4. Aedeagus, schematically. 1–2 L. (Ceratodirus) dostojewskii Semenov, 1899 (1 lateral view, 2 dorsal view); 3–4 L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935 (3 lateral view, 4 dorsal view)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 24–29. Head of Lethrus species with mandibular appendix, keels and hornlike apophyses of mandibles (males), schematically. 24–26 L. (Hetroplistodus) bulbocerus bulbocerus Fischer von Waldheim, 1845 (24 frontal view, 25 lateral view, 26 dorsal view); 27–29 L.(Scelolethrus) sulcatus Kraatz, 1883 (27 fronta view l, 28 lateral view, 29 dorsal view)." ; fabio:hasRepresentation ; a fabio:Figure . dwc:locality "Karschan Tau Mts." ; trt:gbifOccurrenceId "4067802304" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:recordedBy "L. Fairmaire & Det. R. Pittino" ; dwc:typeStatus "lectotype" ; trt:gbifOccurrenceId "4067802303" ; trt:httpUri ; a dwc:MaterialCitation .